Pison ocellare Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 312-315

Pison ocellare Pulawski , species nova

Figures 740 -748.

NAME DERIVATION.– Ocellare , a Neolatin neuter adjective, is derived from ocellus; with reference to the large ocellocular distance combined with the small distance between the hindocelli.

RECOGNITION.– Pison ocellare is an all black species with three submarginal cells. In the female, the setae are erect on the side of tergum I, but in the male there are only a few erect setae or all the setae are appressed. Furthermore, the mandibular apex of P. ocellare is simple, the frontal punctures are small, no more than 0.1-0.2 × midocellar diameter, the gena is punctate and setose on both sides of the oral fossa, the scutal punctures are less than one diameter apart, the mesopleural punctures are less than one diameter apart at the center, the basodorsal hindcoxal tooth is inconspicuous, the apical depression of tergum I is inconspicuous, almost in the same plane as the adjacent more anterior part of tergum, the sterna are evenly, densely punctate (punctures well defined), and the scutal setae are erect and the genal setae are sinuous. Important recognition feature are: the ocellocular distance about twice the the distance between hindocelli ( Figs. 743, 744 ) and sternum II mesally with punctures averaging 2-3 diameters apart. As in P. tibiale and P. dispar , most tergal setae are golden, forming well-defined fasciae on the apical depressions. Unlike the female of P. tibiale (in which the clypeal lamella is divided by an ill-defined, arcuate sulcus into dorsal and ventral portions), the clypeal lamella is simple in the female of P. ocellare , and unlike the male of that species, sternum VIII of P. ocellare has no setose median sulcus. Unlike P. dispar , the inclined part of tergum I is uniformly, finely punctate and the male gaster is black (in P. dispar , the inclined part of tergum I is covered with fine, dense punctures and also with somewhat larger, much sparser punctures that are several to many diameters apart and the male gaster is ferruginous).

from gastral socket area toward spiracle, with conspicuous, short transverse ridges emerging on its admedian side; dorsum irregularly obliquely ridged, punctate between ridges; side punctate, with interspaces merging into irregular ridges; posterior surface ridged. Posteroventral forefemoral surface with small but well-defined punctures that average 2-3 diameters apart. Punctures of tergum I averaging less than one diameter apart on horizontal part. Sterna punctate throughout, punctures minute on apical depression of sternum II.

Setae silvery, erect on upper frons, lower gena, and scutum (here about as long as midocellar diameter), oriented dorsally above dorsal end of midfrontal carina, oriented transversely immediately below midocellus ( Fig. 742 ); not completely concealing integument on clypeus; setae of lower gena sinuous, about twice as long as midocellar diameter. Setae erect on side of tergum I in female, but in male only a few erect setae are present or all setae are appressed. Terga II-V in female, II-VI in male, with golden setae that form conspicuous fasciae on apical depressions.

Body all black except apical depressions of terga II-V in female (II-VI in male) brown.

♀.– Upper interocular distance equal to 0.74-0.76 × lower interocular distance; ocellocular distance equal to 1.3-1.6 × hindocellar diameter, distance between hindocelli equal to 0.7-0.9 × hindocellar diameter ( Fig. 743 ); eye height equal to 0.92-0.94 × distance between eye notches. Free margin of clypeal lamella roundly triangular ( Fig. 740 ). Dorsal length of flagellomere I 2.5-2.7 × apical width, of flagellomere IX 1.4 × apical width. Mandible: trimmal carina with small incision shortly beyond midlength. Length 9.4-10.1 mm; head width 3.0- 3.2 mm.

♂.– Upper interocular distance equal to 0.88-0.90 × lower interocular distance; ocellocular distance equal to 1.8-2.1 × hindocellar diameter, distance between hindocelli equal to 1.0-1.1 × hindocellar diameter; eye height equal to 0.92 × distance between eye notches. Free margin of clypeal lamella acutely angulate ( Fig. 741 ).

Dorsal length of flagellomere I 2.2-2.3 × apical width, of flagellomere X 1.2-1.3 × apical width. Sternum VIII shallowly, narrowly emarginate; apicolateral arm nonprominent, broadly rounded ( Fig. 745 ). Genitalia: Figs. 746, 747 .

Length 7.5-8.7 mm; head width 2.5-2.9 mm.

GEOGRAPHIC DISTRIBUTION (Fig. 748). –

South Australia, Queensland.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA: South

Australia: Wilpena in Flinders Ranges National

Park at 31°31.7ʹS 138°36.2ʹE, 26 Jan 2011,

V. Ahrens and W.J. Pulawski ( SAM).

PARATYPES: AUSTRALIA: Queensland: ca 5 km FIGURE 748. Collecting localities of Pison ocellare N Biloela at 24°13.7ʹS 150°34.7ʹE, 6 Dec 2006, W.J. Pulawski, sp. nov .

Pulawski (1 ♂, CAS); Brisbane : Karawatha Forest at

27°38.6ʹS 153°04.2ʹE, 12 Dec 2006, W.J. Pulawski (1 ♂, CAS) . South Australia: same locality and collectors as holotype : 21 Dec 2011 (1 ♀, 1 ♂, CAS) , 22 Dec 2010 (1 ♀, CAS) , 26 Jan 2011 (2 ♀, CAS) ; 3 km ENE Wilpena at 31°31.0ʹS 138°36.6ʹE , 26 Jan 2011, V. Ahrens and W.J. Pulawski (1 ♂, CAS) .

Figures

FIGURES 740-744. Pison ocellare Pulawski, sp. nov. (740) Female clypeus and mandibles; (741) Male clypeus and mandibles; (742) Upper frons of female showing orien- tation of setae; (743) Female head in dorsal view; (744) Male head in dorsal view.

FIGURES 745-747. Pison ocellare Pulawski, sp. nov., male. (745) Sternum VIII (ventral surface); (746) Genitalia in dorsal view; (747) Genitalia in lateral view.