Pison tenuisculptum Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 452-454

Pison tenuisculptum Pulawski , species nova

Figures 1099-1107.

NAME DERIVATION.– Tenuisculptum is derived from two Latin words: tenuis, meaning fine, delicate, and sculptum, sculptured; with reference to the fine body sculpture, particularly on the propodeal dorsum.

RECOGNITION.– Pison tenuisculptum is a species with the second recurrent vein received near the middle of the second submarginal cell, an all black gaster, and ferruginous femora, tibiae, and tarsi. It differs from most other such species in having a finely punctate propodeal dorsum (also with microscopic, inconspicuous ridges). In addition, the emargination of the inner orbit is the usual shape (not unusually shallow) and tergum I is sessile (its length about equal to apical width). In the female, the distance between the antennal socket and the orbit is about half of the socket width, and in the male the free margin of the clypeal lamella is broadly arcuate (Fig. 1100).

The species is similar to P. breviclypeatum (of which only the female is known). In the female of P. tenuisculptum , however, the frons is not swollen above the antennal base and the head is not subspherical in dorsal view (Fig. 1101), the clypeal lamella is conspicuously protruding beyond the free margin of the lateral section (Fig. 1099), the dorsal length of flagellomere I is 2.8 × apical width, the terga are covered with golden setae, and the length is 8.1-9.3mm. In the female of P. breviclypeatum , the frons is swollen above the antennal base ( Fig. 211 ), the head is subspherical in dorsal view ( Fig. 211 ), the clypeal lamella only insignificantly protrudes beyond the free margin of the lateral section ( Fig. 211 ), the dorsal length of flagellomere I 1.6 × apical width, the terga are covered with inconspicuous silvery setae, and the length is about 5.5 mm.

DESCRIPTION.– Frons dull, minutely punctate, punctures ill defined, about one diameter apart. Distance between antennal socket and orbit equal to about half socket width in female, about equal to socket width in male. Gena narrow in dorsal view (Fig. 1101). Labrum emarginate. Anteromedian pronotal pit transversely elongate, about as long as half midocellar diameter. Scutum finely foveate along flange, with short longitudinal ridges adjacent to posterior margin; scutal punctures minute, less than one diameter apart; interspaces dull (Fig. 1102). Tegula slightly enlarged. Mesopleural punctures minute, even, about one diameter apart. Postspiracular carina present, almost twice as long as midocellar diameter. Metapleural sulcus not costulate between dorsal and ventral metapleural pits. Propodeum with longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum minutely punctate, punctures less than one diameter apart laterally, but becoming sparser toward midline (Fig. 1103), many interspaces merging into fine to microscopic ridges; side microsculptured, minutely punctate, impunctate anteriorly; posterior surface finely punctate. Second recurrent vein received near middle of second submarginal cell. Posteroventral forefemoral surface microscopically, closely punctate. Hindcoxal dorsum with outer margin obtusely carinate. Outer surface of hindtibia with evanescent spines. Punctures of tergum I minute, in some specimens up to several diameters apart on horizontal part. Sternum II except laterally and sternum III mesally minutely microareolate, with microscopic punctures that are many diameters apart.

Setae golden or silvery with golden tinge, appressed on upper frons, scutum, and tergum I, oriented ventrally between midocellus and antenna; completely concealing integument on clypeus (except lamella); on lower gena suberect, about two-thirds of midocellar diameter. Tergal setae golden in female, in male silvery with golden tinge, forming fasciae on apical depressions.

Head, thorax, propodeum, and gaster black, female clypeus yellowish next to lobe free margin; mandible black basally, then yellowish, brown subbasally, and dark brown apically; antenna ferruginous, apical flagellomere dark. Femora, tibiae, and tarsi ferruginous, forefemur dark basodorsally in most specimens.

♀.– Upper interocular distance equal to 1.04-1.06 × lower interocular distance; ocellocular distance equal to 0.5 × hindocellar diameter, distance between hindocelli equal to 0.6-0.7 × hindocellar diameter; eye height equal to 1.12-1.14 × distance between eye notches. Free margin of clypeal lamella approximately truncate (Fig. 1099). Dorsal length of flagellomere I 2.8 × apical width, of flagellomere IX 1.0-1.1 × apical width. Mandible: trimmal carina with small incision at about one third length. Length 8.1-9.3 mm; head width 2.1-2.2 mm.

♂.– Upper interocular distance equal to 0.94 × lower interocular distance; ocellocular distance equal to 0.8 × hindocellar diameter, distance between hindocelli equal to 0.9 × hindocellar diameter; eye height equal to 1.16 × distance between eye notches. Free margin of clypeal lamella broadly arcuate (Fig. 1000). Dorsal length of flagellomere I 2.5 × apical width, of flagellomere X 0.8 × apical width. Sternum VIII shallowly emarginate apically (Fig. 1104). Genitalia are unusual in having gonocoxite markedly shorter than penis valve: Fig. 1105, 1106. Length 7.2 mm; head width 2.0 mm.

GEOGRAPHIC DISTRIBUTION (Fig. 1107).–

Eastern New South Wales, eastern Queensland.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA: New

South Wales: Lorien 3 km N Landowne near Taree,

25-27 Nov 1987, D.J. Bickel (AMS).

PARATYPES: AUSTRALIA: New South Wales :

same data as holotype (1 ♂, AMS); Catherine Hill

Bay, 2 Dec 1962, E.S. Ross and D.O. Cavagnaro

(1 ♀, CAS); Elizabeth Bay [in Sydney], 24 Jan

1952, no collector (2 ♀, AMS). Queensland: Crediton State Forest at 21°11.8ʹS 148°29.9ʹE, 31 Oct

2006, V. Ahrens and W.J. Pulawski (1 ♀, CAS) ;

Eungella National Park at 21°10.5ʹS 148°30.3ʹE,

6 Nov 2012, V. Ahrens and W.J. Pulawski (1 ♀,

CAS); Kroombit Tops State Forest: Kroombit Creek FIGURE 1107. Collecting localities of Pison tenuisculpat 24°23ʹS 151°02ʹE, 31 Oct – 2 Nov 1999, D.J. tum Pulawski, sp. nov.

Bickel (1 ♀, AMS); 18 km S Ravenshoe, 16 Oct 1984, N.W. Rodd (1 ♀, AMS) .

Figures

FIGURES 197–299. Pison bimbi Pulawski, sp. nov., female. (197) Clypeus and mandibles; (198) Head in dorsal view; (199) Pronotum in dorsal view. FIGURE 200. Collecting localities of Pison bimbi Pulawski, sp. nov.