Acalypha urophylla Boivin ex Baill., Adansonia 1: 273. 1861.

Munoz, Iris Montero, Cardiel, Jose Maria & Levin, Geoffrey A., 2018, Nomenclatural review of Acalypha (Euphorbiaceae) of the Western Indian Ocean Region (Madagascar, the Comoros Archipelago, the Mascarene Islands and the Seychelles Archipelago), PhytoKeys 108, pp. 85-116 : 85

33. Acalypha urophylla Boivin ex Baill., Adansonia 1: 273. 1861.

Acalypha reticulata var. urophylla (Boivin ex Baill.) Müll.Arg. Linnaea 34: 32. 1865.

Acalypha filiformis var. urophylla (Boivin ex Baill.) Govaerts, World Checkl. Bibliogr. Euphorbiaceae 63. 2000.

Type. Madagascar. Prov. Antsiranana: Nossibé, plateau de Hell-Ville, Jun 1847, L. H. Boivin 2178 (lectotype, designated here: P [P00536752]!; isolectotypes: G [G00034246]!, GDC [G00324519]!, P [P00536751!, P00536753!]).

Syntypes. Madagascar: s.l., s.d., L. M. A. Du Petit-Thouars s.n. P [P00536748]!; Madagascar. Prov. Antsiranana: Nossibé, Dec 1840, A. Pervillé 364 P [P00536749]!; Madagascar. Prov. Antsiranana: Nossibé, M. Richard 385 P [P00536750]!.

Acalypha goudotiana Baill., Adansonia 1: 268. 1861. Acalypha reticulata var. goudotiana (Baill.) Müll.Arg., Linnaea 34: 32. 1865. Acalypha filiformis var. goudotiana (Baill.) Govaerts, World Checkl. Bibliogr. Euphorbiaceae 62. 2000. Type: Madagascar: s.l., 1830, J. P. Goudot s.n. (holotype: G!; isotype: P [P00536727]! fragment), syn. nov.

Acalypha reticulata var. urophylla f. meeusei Leandri, Notul. Syst. (Paris) 10: 264. 1942. Type: Madagascar, Prov. Fianarantsoa, base Est du Pic d’Ivohibe, 19 Sep 1926, R. Decary 5352 P [P00224690]!; Prov. Toamasina, Analamazaotra forest, 1912, H. Perrier de la Bâthier 9741 (lectotype, designated here: P [P05604377]!, syn. nov.).

Syntypes. Madagascar, Prov. Toamasina, Soanierana, Antasibé, 9 Dec 1938, H. J. Lam & A. D. J. Meeuse 5814 WAG [WAG0133229]!, L [L0242110]!; Moramanga, 900 m, 11 Nov 1938, H. J. Lam & A. D. J. Meeuse 5363 L [L0242105]!; Analamazaotra, 1000 m, 10 Nov. 1938, H. J. Lam & A. D. J. Meeuse 5290 L [L0242106]!.

Acalypha reticulata var. urophylla f. longa Leandri, Notul. Syst. (Paris) 10: 265. 1942. Type: Madagascar, Prov. Toliara, bassin de la Manampanihy, Col de Fitana, 700 m, 15 Oct 1928, H. Humbert 6015 (lectotype, designated here: P [P05604383]!, syn. nov.).

Syntypes. Madagascar, Prov. Toliara, Col d’Ivolo (District de Fort-Dauphin), 500 m, 5 Sept 1932, R. Decary 10558 P [P05604384]!; Fort-Dauphin, col de Tsitongabarika, 600 m, 9 Sept 1932, R. Decary 10595 P [P05604399]!.

Acalypha reticulata var. urophylla f. vohitrae Leandri, Notul. Syst. (Paris) 10: 265. 1942. Type: Madagascar, Prov. Toamasina, Andevorante, rive droite de la Vohitra près de Lohariandava, 200-250 m, 10 Oct 1912, R. Viguier & H. Humbert 661 (lectotype, designated here: P [P05604437]!; isolectotypes: P [P05604440!, P05604441!], syn. nov.).

Acalypha reticulata var. urophylla f. glabrescens Leandri, Notul. Syst. (Paris) 10: 266. 1942. Type: Madagascar, Prov. Antsiranana, Massif du Tsaratanana, 2200 m, H. Perrier de la Bâthie 16180 (lectotype, designated here: P [P05604386]! syn. nov.).

Syntypes. Madagascar, Prov. Antsiranana, Sambirano, Nov-Dec 1937, H. Humbert 18659 P [P05604397!, P05604400!]; Massif du Tsaratanana, Sep 1912, H. Perrier de la Bâthie 18614 P [P05604382]!; Massif du Tsaratanana, 1600 m, H. Perrier de la Bâthie 15371 P [P05604385]!; Madagascar, Prov. Fianarantsoa, Ranohira, Isalo, 30 Jul 1928, H. Humbert 5014 P [P05604401!, P05604402!]; Madagascar, Prov. Mahajanga, Beritsoka, H. Perrier de la Bâthie 413 P [P05604392!, P05604390!]; Beritsoka, Dec 1897, H. Perrier de la Bâthie 422 P [P05604387!, P05604388!, P05604391!]; Beritsoka, H. Perrier de la Bâthie 9822 P [P05604389]!; Madagascar, s. l., R. Baron 5987 P [P05604407]!; s. l., L. Humblot 335 P [P05604393!, P05604394!, P05604395!, P05604396!].

Acalypha reticulata var. urophylla f. typique Leandri, Notul. Syst. (Paris) 10: 262. 1942 nom. inval.

Distribution.

Madagascar (Antananarivo, Antsiranana, Fianarantsoa, Mahajanga, Toamasina and Toliara). Comoros Archipelago (Grande Comore, Anjouan, Mohéli and Mayotte).

References.

Müller Argoviensis (1866: 260, 852) as A. reticulata var. goudotiana and A. reticulata var. urophylla ; Baron (1889: 262); Baillon (1891: 189); Baillon (1892: 1004); De Candolle (1901: 567); Palacký (1907: 25 as A. goudotiana , 26); Pax and Hoffmann (1924: 105, 260) as as A. reticulata var. goudotiana and A. reticulata var. urophylla ; Leandri (1942: 258, 260, 262) as A. reticulata var. goudotiana and A. reticulata var. urophylla ; Leandri (1942: 281) as A. emirnensis ; Leandri (1948: 186) as A. reticulata var. goudotiana ; Govaerts et al. (2000: 62, 63, 100, 105, 108).

Notes.

Acalypha urophylla has been considered to be a variety of A. reticulata , but A. urophylla can be distinguished by its sessile female bracts with dentate margins vs. pedicellate female bracts with crenate to subentire margins in A. filiformis . (see notes in A. filiformis ). See notes under A. gracilipes , A. lamiana and A. paxii for the differences between those species and A. urophylla .