Coniceromyia systenopoda, Ament & Kung & Brown, 2020

Ament, Danilo César, Kung, Giar-Ann & Brown, Brian V., 2020, Forty-one new species of Coniceromyia Borgmeier (Diptera: Phoridae), an identification key, and new distributional records for the species of the genus, Zootaxa 4830 (1), pp. 1-61 : 47

Coniceromyia systenopoda sp. nov.

( Figs. 7C , 9I , 16C , 22K , 25P , 26G )

Diagnosis (male). Base of forefemur with a triangular ventral projection bearing short dark setulae posteriorly and long thick setae along anteroventral margin ( Fig. 26G ).

Material examined. Holotype ♂, COLOMBIA: Valle de Cauca: PNN Farallones de Cali: Anchicaya , 3.43ºN, 76.80ºW, 21.xi–19.xii.2000, Malaise trap, 900m, S. Sarria col. ( IAVH) [LACM ENT 165165].

Description. Male. Body length 2 mm. Head. Frons brown, sparsely setulose, with well-defined median furrow. Flagellomere 1 light brown, conical-elongate, with abruptly thinner elongation two times longer than base. Arista apical, short. Palpus light brown; gena with two setae. Thorax. Scutum, scutellum, and pleural regions brown; anepisternum bare. Legs light brown, fore and midtibiae and tarsi lighter. Base of forefemur with a triangular ventral projection bearing short dark setulae posteriorly and long thick setae along anteroventral margin ( Fig. 26G ). Forefemur with ventral row of thin setae. Foretibia with three dorsal setae and anterodorsal row of strong setulae. Foremetatarsus with anterior apparently sculptured excavation, short setulae at base of excavation, row of setulae on ventral margin of excavation, and without anteroapical process, but with a strong seta in its place ( Fig. 9I ). Foretarsomeres 2 and 3 short. Foremetatarsus ratio 0.25. Posterior face of hind femur with narrow basoventral band of tiny blunt setulae ( Fig. 22K ). Hind femur height/length ratio, 0.37. Hind tibia with one dorsal seta in basal half. Wing ( Fig. 16C ). Costa swollen, 0.47 of the wing length; R 2+3 absent; M 1 concave, curved posteriorly at apex; M 2 approximately concave, CuA 1 -M 2 /M 2 -M 1 ratio 1.53; CuA 1 sinuous. Halter dark brown at base, light brown at apex.

Abdomen. Tergites brown, with shining pubescence. Hypopygium light brown ( Fig. 25P ), left epandrial process with only one discernible process: a subepandrial wall extending from the hypoproct to its apex. Margin of right lobe of hypandrium without lateral projection. Hypoproct with one seta.

Distribution. Known from a single low-elevation site in Colombia.

Etymology. Name derived from the Greek roots systenos (running or tapering to a point) and podos (foot), referring to the base of forefemur with a triangular ventral projection.

Figures

FIGURE 7. Habitus of the holotypes of new species of Coniceromyia, continuation (scale bars=1mm). A. C. stichochaeta, B. C. strongyla, C. C. systenopoda, D. C. trilobata, E. C. xiei.

FIGURE 9. Male foremetatarsus of new species of Coniceromyia, continuation (scale bars=0.1mm). A. C. platypoda, B. C. polychaeta, C. C. puntarenensis, D. C. riccardiae, E. C. setosa, F. C. solaris, G. C. stichochaeta, H. C. strongyla, I. C. systenopoda, J. C. trilobata, K. C. xiei.

FIGURE 16. Wings of the males of Coniceromyia new species, continuation. A. C. stichochaeta, B. C. strongyla, C. C. systenopoda, D. C. trilobata, E. C. xiei.

FIGURE 22. Hind femora of the males of Coniceromyia new species, continuation (scale bars on figures of the entire femur=0.3 mm, on figures of ventrobasal part of the femur=0.1 mm). A. C. polychaeta, B. C. puntarenensis, C. C. riccardiae, D–E. C. setosa, F–G. C. solaris, H–I. C. stichochaeta, J. C. strongyla, K. C. systenopoda.

FIGURE 25. Hypopygia of Coniceromyia new species in right lateral view, continuation. A. C. megalosoma, B. C. memorialis, C. C. niemeyeri, D. C. numerosa, E. C. ochyrosticha, F. C. parallela, G. C. pectinella, H. C. platypoda, I. C. polychaeta, J. C. puntarenensis, K. C. riccardiae, L. C. setosa, M. C. solaris, N. C. stichochaeta, O. C. strongyla, P. C. systenopoda, Q. C. trilobata, R. C. xiei.

FIGURE 26. Unique or uncommon features of males of Coniceromyia species. A. Curved forecoxa of C. broadheadi, B–C, Sculpturing in dorsum of forefemur of C. broadheadi, D. Anterior view of foretibia of C. leia, showing the ventral face mostly bare and shiny but with differentiated thin setulae, E. Hind tibia of C. piricornis with dorsal patch of differentiated setulae, F. Hind tibia of C. diadella with dorsal carina with differentiated setulae, G. Posteroventral face of forefemur of C. systenopoda with a triangular ventral projection, H. Forefemur of C. pectinella with dorsal excavation bearing a comb of apically-directed curved setae, I. Foretrochanter of C. circulata with ventral process bearing apical group of curved setae, J. Foretrochanter of C. bellatula with ventral projection bearing apical group of curved setulae, K. Forefemur of C. chrysopa with bare oval ring surrounding an oval patch with small setulae, L. Forefemur of C. solaris with bare oval ring surrounding an oval patch with small setulae, M. Anterior face of midfemur of C. piricornis with ventral patch of dense setulae, N. Anterior face of midfemur of C. setosa with band of tiny setulae along basal two-thirds of ventral margin.