Ontherus (Caelontherus) trituberculatus Balthasar, 1938

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 169

Ontherus (Caelontherus) trituberculatus Balthasar, 1938 Plate 38A

Ontherus trituberculatus Balthasar, 1938: 220 (original description. Type locality: Amerika merid. [= South America.], Cachabé).

Ontherus trituberculatus : Blackwelder 1944: 206 (list of species from Latin America); Medina et al. 2001: 139 (cited for Colombia); Krajcik 2012: 174 (complete list of species).

Ontherus (Caelontherus) trituberculatus : Génier 1996: 53 (cited as new combination, redescription); Carvajal et al. 2011: 318-319 (cited for Ecuador); Chamorro et al. 2018: 96 (cited for Ecuador).

Type specimens.

Ontherus trituberculatus Balthasar, 1938 Génier, 1996. The holotype (♂) is deposited at the SMTD (see Génier 1996: 54). Locality: Cachabé, not examained.

Distribution.

Colombia and Ecuador.

Records examined.

CARCHI: Maldonado, 1830 m (10 specimens CEMT; 8 specimens MQCAZ); Quinjul, 1700 m (2 specimens CEMT; 4 specimens MQCAZ); Tobar Donoso, 300 m (16 specimens MECN). EL ORO: Reserva Jocotoco, 1250 m (3 specimens MQCAZ). ESMERALDAS: Charco Vicente (21 specimens MECN; 17 specimens MQCAZ); Palma Real (14 specimens MECN; 11 specimens MQCAZ); Playa de Oro, 200 m (1 specimen MUTPL; 18 specimens MECN; 26 specimens MQCAZ); Playa de Oro, La Tabla (37 specimens MECN; 21 specimens MQCAZ); Playa de Oro, Padre Santo (21 specimens MECN; 28 specimens MQCAZ); Playa de Oro, Playa Rica (13 specimens MECN; 9 specimens MQCAZ); Playa de Oro, Pote (8 specimens CEMT; 20 specimens MQCAZ); Salto del Bravo (17 specimens MECN; 10 specimens MQCAZ); Tsejpi (28 specimens MECN; 17 specimens MQCAZ). IMBABURA: Río Getsemani, 600 m (4 specimens MQCAZ); Lita, 680 m (7 specimens MECN; 5 specimens MQCAZ). PICHINCHA: Choconde, 1200 m, San Miguel de los Bancos (1 specimen MUTPL); El Encuentro, 620 m, San Miguel de los Bancos (1 specimen MUTPL); El Tigre Río Guayllabamba, Pedro Vicente Maldonado (2 specimens MUTPL); Guayabilla Río Guayllabamba, 520 m, Manduriacus (1 specimen MUTPL); Llurimaguas Río Guayllabamba, 290 m, Pedro Vicente Maldonado (2 specimens MUTPL); Mangaloma, 820 m, San Miguel de los Bancos (1 specimen MUTPL); Pedro Vicente Maldonado, 640 m (3 specimens CEMT; 5 specimens MCAZ); Tortugo Río Guayllabamba, 450 m, Pedro Vicente Maldonado (2 specimens MUTPL).

Literature records.

CARCHI: Chical, 1250 m ( Génier 1996: 55); 18 km SE Maldonado, 2420 m ( Génier 1996: 55). ESMERALDAS: Cachabé ( Génier 1996: 55). PICHINCHA: 5.3 km road to Pachija [= Pachijal], 2800-3000m ( Génier 1996: 55); 85 km NW de Quito, on Puerto Quito Rd, 1520 m ( Génier 1996: 55); 113 km NW Quito, on Puerto Quito Rd, 790 m ( Génier 1996: 55). PICHINCHA: [= SANTO DOMINGO DE LOS TSÁCHILAS]: 4 km SE Santo Domingo, 500 m ( Génier 1996: 55); 16 km SE Santo Domingo, Tinalandia, 680 m ( Génier 1996: 55).

Temporal data.

Collected in February, March, April, May, July, September, October, November, and December.

Remarks.

Inhabits the coastal lowland evergreen forests and coastal evergreen foothill forests from 200-1250 m a.s.l. Additionally, there is a single record for this species for a locality 5.3 km along the road to Pachija [= Pachijal]. However, because this is the only record from the Andean region cited by Génier (1996), it may be erroneous. Collected with pitfall traps baited with carrion and human feces.