Bromus lepidus Holmb., Bot. Not. 1924: 326 (1924)

Llamas, Felix & Acedo, Carmen, 2019, Typification of eight current and seven related names and a new section in the genus Bromus (Bromeae, Pooideae, Poaceae), PhytoKeys 121, pp. 53-72 : 59-60

Bromus lepidus Holmb., Bot. Not. 1924: 326 (1924)

Bromus lepidus Holmb., Bot. Not. 1924: 326 (1924).Type Protologue: "Nach ROUY kommt sie in Frankreich hier und da vor,aber ziemlich selten; Krösche hat sie nur von einem Ståndort in Norddeutschland. In Schweden kommt sie besonders in Schonen vor; ich habe sie mehrmals eingesammelt, wie in der Gegend von Lund und Malmö an mehreren Orten; Svenshög in Wallkärra; Skartofta in Öved; Gudmundtorp. Außerdem sah ich Exemplare aus Blommeröd in Öved (leg. P. Boren 1903), Skelderviken (leg. Sten Selander), Kalmar (leg. N. Blomgren), Borås (leg. C. Sandberg), Fyen, Stenlose (leg. G. Samuelsson). Kommt in Klee- und Grasfeldern, auf Rainen, Wegrändern etc. vor, oft mit B. mollis und B. commutatus zusammen; bei Malmö auch als Ruderat". Type: SWEDEN. Scania: Malmö, in ruderatis, 18-06-1920, Otto R.Holmberg, (lectotype, designated here: LD [LD1136595 image!]; isolectotype: K [K000913599!]). (Figure 7)

Bromus lepidus Holmb. f. lasiolepis Holmb., Bot. Not. 1924: 326 (1924). Type Protologue: not indicated. Type: Sweden. Scania: Vallkörra. Svenenshög. 4-07-1923. Otto R.Holmberg, (lectotype, designated here: LD [LD 1136235 image!]; isolectotype: K [K000913598!]). (Figure 8)

Remarks.

This taxon has a long history full of nomenclatural problems. Duval-Jouve (1865: 208), who only lists, without naming them, the variations in spikelet size and hairiness of Bromus species, e.g. Bromus mollis " microstachys glabre" and Bromus mollis " microstachys pubescent". This is the first mention of this taxon. Later Rouy (1913: 236) proposed a named variety under the genus Serrafalcus Parl.: S. mollis ß microstachys Rouy, giving " Bromus microstachys Duval-Jouve" (1865: 207) as a synonym and adding a diagnosis. Therefore, there is no doubt this is the same plant cited by Duval-Jouve (l. c.). Afterwards, Krösche (1924: 329) described B. gracilis Krösche, which unfortunately is a posterior homonym as Weigel (1772: 15) previously proposed this name for a different plant.

Finally, Holmberg (1924: 326) solves this unfortunate situation and names simultaneously the new taxa Bromus lepidus Holmb. and B. lepidus f. lasiolepis Holmb. The herbaria having Holmberg material are K, LD and S. There are 13 sheets in LD collected before 1924, five in S and three in K. It is reasonably certain that Holmberg studied all those plants before the description of the species and form and that all of them are original material. Therefore, we decided to limit our choices to sheets with the annotations " B. lepidus mihi" and " B. lepidus f. lasiolepis Holmb.", as this annotation indicates that Holmberg is interpreting them as the new taxa he is going to describe.