Colastes (Fungivenator), Achterberg & Shaw, 2008
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publication ID |
https://doi.org/ 10.1080/00222930802114280 |
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persistent identifier |
https://treatment.plazi.org/id/E369983E-840C-FFF2-FE49-A809FB8C504A |
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treatment provided by |
Felipe |
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scientific name |
Colastes (Fungivenator) |
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Subgenus Fungivenator nov.
( Figures 1 View Figure 1 , 2A–C View Figure 2 )
Type species
Colastes sandei sp. nov.
Etymology
From ‘‘fungis’’ (Latin for ‘‘fungus’’) and ‘‘venator’’ (Latin for ‘‘hunter’’), because the females search for fungi in which to hunt hosts. Gender: masculine.
Diagnosis
Antenna of female with 20–31 segments, third segment 4–5 times as long as wide ( Figures 1K View Figure 1 , 2B View Figure 2 ); occipital carina narrowly reduced medio-dorsally and remaining far from hypostomal carina ventrally ( Figure 2B View Figure 2 ); clypeus convex, distinctly differentiated from face ( Figures 1D View Figure 1 , 2B View Figure 2 ); ventral rim of clypeus thin, protruding beyond level of face and apex of mandible in lateral view ( Figures 1L View Figure 1 , 2B View Figure 2 ); hypoclypeal depression deep and rather large ( Figure 1D View Figure 1 ); orbits reddish-brown or brown dorsally; pronope absent or small (but distinct in one of the paratypes of C. sandei from Monks Wood), pronotum dorsally often with a more or less crenulate transverse depression anteriorly, which can be open anteriorly; mesoscutum glabrous and strongly shiny, but densely setose (except posteriorly) in C. fritzeni ; notauli finely crenulate, posteriorly notauli not strongly converging and ending in a broad more or less rugose area ( Figure 1B View Figure 1 ); scutellum often finely rugulose medioposteriorly ( Figure 1B View Figure 1 ); propodeum coarsely reticulate, at most anteriorly narrowly smooth; vein r shorter than width of pterostigma ( Figures 1A, 1E View Figure 1 , 2A View Figure 2 , 3A View Figure 3 ); shape and pigmentation of pterostigma of male similar to pterostigma of female ( Figures 1A, 1E View Figure 1 ); vein 1-SR of forewing and vein 1-M of hind wing medium-sized ( Figures 1A View Figure 1 , 2A View Figure 2 ); vein r of forewing usually distinctly oblique and from basal 0.3–0.5 of pterostigma ( Figures 1A, 1E View Figure 1 , 2A View Figure 2 , 3A View Figure 3 ); first subdiscal cell of forewing robust ( Figures 1A View Figure 1 , 2A View Figure 2 ); dorsal carina of first metasomal tergite not united medially ( Figure 2D View Figure 2 ) or united and without distinct median crest ( Figure 1H View Figure 1 ); second metasomal tergite distinctly striate basally; second and third tergites about of equal length and second tergite less transverse than third tergite ( Figure 1H View Figure 1 ); fourth and fifth tergites smooth ( Figure 1L View Figure 1 ); fifth tergite narrowed posteriorly; ovipositor slender ( Figure 1L View Figure 1 ); length of ovipositor sheath 0.3–0.5 times as long as forewing and 2–4 times as long as first tergite.
Biology
Host unknown, but reared from bracket fungi; considering the biology of other species of the genus Colastes likely to be an idiobiont ectoparasitoid of holometabolous larvae in the bracket fungi, with strong circumstantial evidence for one species to be a parasitoid of Dorcatoma species ( Coleoptera : Anobiidae ).
Distribution
Holarctic, five species.
Key to species of the subgenus Fungivenator nov.
1. Length of ovipositor sheath about 0.5 times forewing; pterostigma elongate (( Figure 3A View Figure 3 ); middle lobe of mesoscutum and scutellum densely setose; hind femur micro-sculptured and rather robust ( Figure 3D View Figure 3 ); outer side of hind coxa rugulose; mesosternum with widely crenulate and deep sulcus medially and sublaterally superficially crenulate; Finland................... fritzeni View in CoL
- Length of ovipositor sheath 0.3–0.4 times forewing; pterostigma robust (( Figures 1A View Figure 1 , 2A View Figure 2 ); middle lobe of mesoscutum and scutellum glabrous; hind femur largely smooth and more slender ( Figure 1C View Figure 1 ); outer side of hind coxa at least partly smooth; mesosternum with narrowly crenulate and rather deep sulcus medially and sublaterally smooth.......................... 2
2. First metasomal tergite more finely, densely and very regularly striate, tergite evenly strongly convex; length of ovipositor sheath about 0.4 times forewing; antenna with 28–31 segments; Nearctic...................... polypori View in CoL
- First tergite more coarsely, less densely and rather irregularly striate or rugose ( Figures 1H View Figure 1 , 2D View Figure 2 ), tergite more or less flattened medially; length of ovipositor sheath about 0.3 times forewing; antenna with 21–30 segments; Palaearctic............................................... 3
3. Vein 3-SR of forewing 0.9–1.0 times as long as vein 2-SR ( Figure 1A View Figure 1 ); basally vein r of forewing somewhat wider than vein 2-SR ( Figures 1A, 1E View Figure 1 ); pterostigma distinctly triangular ( Figure 1A View Figure 1 ); West Palaearctic...... sandei View in CoL
- Vein 3-SR of forewing 1.3–1.5 times as long as vein 2-SR ( Figure 2A View Figure 2 ); basally vein r of forewing slender, about as wide as vein 2-SR ( Figure 2A View Figure 2 ); pterostigma elliptical ( Figure 2A View Figure 2 ); Central and East Palaearctic................... 4
4. Ventral half of face rugulose; first subdiscal cell of forewing weakly narrowed basally; vein 3-SR of forewing about 1.3 times as long as vein 2-SR.............................................. effectus
- Ventral half of face densely coriaceous; first subdiscal cell of forewing strongly narrowed basally ( Figure 2A View Figure 2 ); vein 3-SR of forewing about 1.5 times as long as vein 2-SR ( Figure 2A View Figure 2 )................................. aciculatus View in CoL
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