Panjange thomi Huber

Huber, Bernhard A. & Carvalho, Leonardo S., 2019, Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae), Zootaxa 4546 (1), pp. 1-96: 80-81

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Panjange thomi Huber

sp. n.

Panjange thomi Huber  sp. n.

Figs 300–303View FIGURES 300–307, 308–311View FIGURES 308–309View FIGURES 310–311, 316–318View FIGURES 316–321, 333–335View FIGURES 329–335

Panjange Ind  103: Eberle et al. 2018 (molecular data); Huber et al. 2018: fig. 10.

Note. Females of this species were compared directly with the female holotype of Panjange dubia ( Kulczyński, 1911)  : the epigynum is indistinguishable externally. However, several species in the Panjange cavicola  group have very similar female external genitalia (e.g., P. alba Deeleman-Reinhold & Deeleman, 1983  ; P. madang Huber, 2011  ), and the dark mark in the ocular area is smaller in all females below than in the P. dubia  holotype. Thus, the specimens below are not considered to be P. dubia  even though they originate from closer to the type locality of P. dubia  than the male tentatively described as P. dubia in Huber (2011a)  (175 km vs. 400 km; Fig. 350View FIGURE 350).

Type material. INDONESIA: ♂ holotype, ZFMK (Ar 20660), West Papua, Manokwari, Gunung Meja (0.860°S, 134.084°E), 190 m a.s.l., 8.xi.2009 (S. Sutono)GoogleMaps  .

Other material examined. INDONESIA: 1♂ 9♀ 2 juvs, ZFMK (Ar 20661–62), and  2♀ 4 juvs in pure ethanol, ZFMK (Ind199), same data as holotypeGoogleMaps  .

Etymology. The species is named for Thomas Wapai N.S. Wainggai (also known as Dr. Thom) (1937–1996), West Papuan scholar and political leader.

Diagnosis. Males can be distinguished from other representatives of the cavicola  group with strongly elongated palps ( P. alba  , P. dubia  , P. madang  , P. mirabilis  ) by shape of procursus ( Figs 308–309View FIGURES 308–309; only one ventral ridge, distinctive flat distal elements) and by extremely elongated male palpal tibia (Figs; 308–309> 7 x longer than wide; other species: <5 x longer than wide). Females of these five species may be indistinguishable externally, but internal genitalia of P. thomi  with distinctive structures diverging towards anterior ( Figs 311View FIGURES 310–311, 318View FIGURES 316–321).

Description. Male (holotype). MEASUREMENTS. Total length 3.7, carapace width 0.87. Distance PME-PME 400 µm; diameter PME 90 µm; distance PME-ALE 40 µm; AME absent. Leg 1: 38.2 (8.8 + 0.4 + 9.0 + 18.1 + 1.9), tibia 2: 5.8, tibia 3: 3.2, tibia 4: 4.7; tibia 1 L/d: 113.

COLOR (in ethanol). Carapace pale whitish with dark median mark posteriorly and dark brown mark in ocular area including eye stalks; clypeus not darkened; sternum whitish; legs ochre-yellow, with brown rings in patella area and at tibia-metatarsus joints. Abdomen pale gray, with dark marks dorsally and laterally.

BODY. Habitus as in Fig. 300View FIGURES 300–307. Eye triads on stalks ( Fig. 310View FIGURES 310–311), each stalk with single pointed process curved towards anterior. Thoracic furrow absent. Clypeus unmodified. Sternum wider than long (0.54/0.44), unmodified. Abdomen cylindrical, angular dorso-posteriorly.

CHELICERAE. As in Fig. 310View FIGURES 310–311, with pair of simple weakly sclerotized processes proximally.

PALPS. As in Figs 308–309View FIGURES 308–309, symmetric; coxa with small but distinct retrolateral-ventral apophysis, trochanter with retrolateral-dorsal process, femur with rounded prolateral process proximally; femur-patella joints strongly shifted towards prolateral side; tibia strongly elongated; tarsus with long whitish dorsal process with tarsal organ at tip; procursus with only one ventral ridge, without ventral pointed process, with distinctive distal flat elements; genital bulb oval, with two very long processes lying parallel to each other: semitransparent embolus and slightly sclerotized bulbal apophysis (‘appendix’; ba in Fig. 308View FIGURES 308–309).

LEGS. Without spines and curved hairs, few vertical hairs; retrolateral trichobothrium of tibia 1 at 3%; tibia 1 without prolateral trichobothrium (present on other tibiae); tarsus 1 with ~30 pseudosegments, only distally a few fairly distinct.

Male (variation). Tibia 1 in other male: 9.1.

Female. In general similar to male ( Figs 302–303View FIGURES 300–307) but eye triads on low humps, closer together (distance PME-PME 210 µm) and dark mark in ocular area smaller (only dark band between eye triads). Tibia 1 in 8 females: 7.6–9.0 (mean 8.2). Tarsus 4 comb-hairs as in Fig. 334View FIGURES 329–335. Epigynum weakly sclerotized ( Fig. 316View FIGURES 316–321), with long folded scape ending in rounded ‘knob’ ( Figs 317View FIGURES 316–321, 333View FIGURES 329–335). Internal genitalia with pair of roundish pore plates and distinctive structures diverging towards anterior ( Figs 311View FIGURES 310–311, 318View FIGURES 316–321). ALS with one widened, one pointed, and six cylindrically shaped spigots ( Fig. 335View FIGURES 329–335).

Distribution. Known from type locality only ( Fig. 350View FIGURE 350).

Natural history. The spiders were collected from the undersides of green leaves.