Aetana ondawamei Huber

Huber, Bernhard A. & Carvalho, Leonardo S., 2019, Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae), Zootaxa 4546 (1), pp. 1-96: 78-79

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Aetana ondawamei Huber

sp. n.

Aetana ondawamei Huber  sp. n.

Figs 270–271View FIGURES 264–271, 286–287View FIGURES 283–287, 290View FIGURES 288–293, 297–299View FIGURES 294–299

Aetana Ind  101: Eberle et al. 2018 (molecular data); Huber et al. 2018: fig. 7.

Type material. INDONESIA: ♂ holotype, ZFMK (Ar 20657), West Papua, Manokwari, Gunung Meja (0.860°S, 134.084°E), 190 m a.s.l., among rocks, 8.xi.2009 (S. Sutono)GoogleMaps  .

Other material examined. INDONESIA: 4♂ 7♀, ZFMK (Ar 20658–59), and  2♀ 1 juv. in pure ethanol, ZFMK (Ind198), same data as holotypeGoogleMaps  .

Etymology. The species is named for Papuan activist John Ondawame (1953–2014) who promoted a peaceful solution to the conflict in West Papua.

Diagnosis. Males differ from most known congeners (except A. mokwam  ) by pair of finger-shaped apophyses on male palpal trochanter (cf. Fig. 281View FIGURES 281–282) and by shape of procursus ( Fig. 286View FIGURES 283–287; three distal elements: whitish oval element with two heavily sclerotized rounded apophyses, and two long hinged elements); from A. mokwam  by much smaller palps (compare Figs 284 and 286View FIGURES 283–287) and by wider dorsal hinged element of procursus (12 x longer than wide versus 19 x in A. mokwam  ). Females differ from most congeners (except A. mokwam  and A. ternate  ) by elongated female internal genitalia and pore plates ( Fig. 287View FIGURES 283–287); females differ from A. mokwam  and A. ternate  by distinctive curved internal folds visible through cuticle ( Fig. 297View FIGURES 294–299); from A. mokwam  also by anterior part of internal genitalia barely narrower than posterior part ( Figs 298–299View FIGURES 294–299). Males and females also differ from A. mokwam  by pair of marks ventrally on abdomen (unpaired median mark in A. mokwam  ; compare Figs 289 and 290View FIGURES 288–293).

Description. Male (holotype). MEASUREMENTS. Total length 3.5, carapace width 1.1. Distance PME-PME 340 µm; diameter PME 110 µm; distance PME-ALE 40 µm; AME absent. Leg 1: 38.5 (9.0 + 0.5 + 9.0 + 16.7 + 3.3), tibia 2: 5.5, tibia 3: 3.6, tibia 4: 5.5; tibia 1 L/d: 90.

COLOR (in ethanol). Carapace ochre yellow with narrow dark brown lateral marginal bands and narrow median band including posterior part of ocular area; clypeus with dark brown pattern in distal half; sternum monochromous light brown; legs light brown, without darker rings, tips of femora and tibiae lighter. Abdomen pale gray, densely covered with dark internal marks dorsally and laterally; ventrally with pair of dark marks behind gonopore and median mark in front of spinnerets.

BODY. Habitus as in Fig. 270View FIGURES 264–271. Eye triads on very short stalks directed towards lateral. Thoracic furrow absent (only black median line). Clypeus with pair of short obtuse apophyses close to distal margin (cf. Fig. 283View FIGURES 283–287). Sternum wider than long (0.72/0.56), unmodified.

CHELICERAE. As in A. mokwam  (cf. Fig. 283View FIGURES 283–287), with pair of long lateral apophyses directed towards lateral and anterior, and pair of small proximal lateral processes.

PALPS. Almost identical in shape to A. mokwam  (cf. Figs 281–282View FIGURES 281–282) but much smaller (compare Figs 284 and 286View FIGURES 283–287); dorsal hinged element of procursus relatively thicker (12 x as long as thick in mid-section versus 19 x in A. mokwam  ).

LEGS. Without spines; few vertical hairs; with curved hairs on tibiae and metatarsi 2; retrolateral trichobothrium of tibia 1 at 3%; tibia 1 without prolateral trichobothrium (present on other tibiae); tarsus 1 with ~30 pseudosegments, distally distinct.

Male (variation). Tibia 1 in 5 males (including holotype): 8.3–9.6 (mean 9.1).

Female. In general similar to male ( Fig. 271View FIGURES 264–271) but eye triads on low humps and closer together (distance PME- PME 200 µm), clypeus unmodified, abdomen anteriorly above pedicel with slightly sclerotized area acting against barely elevated small brown area medially on carapace; sometimes with indistinct darker rings on leg femora (subdistal) and tibiae (proximal and subdistal). Tibia 1 in 6 females: 7.9–8.6 (mean 8.3). Epigynum long, resulting in relatively posterior position of genital opening ( Fig. 290View FIGURES 288–293); anterior plate barely modified in anterior part, internal valve visible through cuticle ( Fig. 297View FIGURES 294–299), with strong transversal ridges in posterior part and dark internal folds visible through cuticle; posterior plate simple, pockets possibly present in the membrane on both sides of genital opening (arrows in Fig. 287View FIGURES 283–287). Internal genitalia with very long pore plates (actually fields of pores), apparently without internal pockets ( Figs 287View FIGURES 283–287, 299View FIGURES 294–299).

Distribution. Known from type locality only ( Fig. 349View FIGURE 349).

Natural history. This species was found among karstic rocks near the ground, in domed sheet webs of ~ 30 cm diameter. The webs were rather exposed and the spiders moved slowly when disturbed.