Panjange togutil Huber

Huber, Bernhard A. & Carvalho, Leonardo S., 2019, Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae), Zootaxa 4546 (1), pp. 1-96: 82-84

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Panjange togutil Huber

sp. n.

Panjange togutil Huber  sp. n.

Figs 304–307View FIGURES 300–307, 312–315View FIGURES 312–313View FIGURES 314–315, 319–321View FIGURES 316–321

Panjange Ind  109: Eberle et al. 2018 (molecular data); Huber et al. 2018: fig. 10. Type material. INDONESIA: ♂ holotype, ZFMK (Ar 20663), Halmahera, degraded forest along river near Sofifi (0.715°N, 127.586°E), 130–180 m a.s.l., 11.xi.2009 (S. Sutono).

Other material examined. INDONESIA: 3♂ 5♀ 7 juvs, ZFMK (Ar 20664–65), and 7 juvs in pure ethanol, ZFMK (Ind203), same data as holotype  .

Etymology. The species is named for the Togutil people in the interiors of north and central Halmahera; noun in apposition.

Diagnosis. Males are easily distinguished from other known representatives of the cavicola  group by strongly widening distal part of procursus ( Figs 312–313View FIGURES 312–313; without dorsal process as in P. cavicola  ) and by color pattern on carapace ( Fig. 304View FIGURES 300–307; anterior half pale whitish, posterior half dark brown); females differ (except from P. cavicola  ) by short epigynal scape ( Figs 315View FIGURES 314–315, 319–320View FIGURES 316–321); both males and females also distinguished by posteriorly widened abdomen (in dorsal view; Figs 304–307View FIGURES 300–307).

Description. Male (holotype). MEASUREMENTS. Total length 3.9, carapace width 1.0. Distance PME-PME 390 µm; diameter PME 90 µm; distance PME-ALE 30 µm; AME absent. Leg 1: 36.9 (8.5 + 0.4 + 8.7 + 17.4 + 1.9), tibia 2: 5.5, tibia 3: 3.0, tibia 4: 4.5; tibia 1 L/d: 109.

COLOR (in ethanol). Carapace anterior half pale whitish, posterior half dark brown; ocular area and clypeus also brown; sternum whitish; legs pale ochre-yellow, brown rings in patella area and at tibia-metatarsus joint. Abdomen pale ochre-yellow, with dark marks dorsally and laterally.

BODY. Habitus as in Fig. 304View FIGURES 300–307. Eye triads on stalks ( Fig. 314View FIGURES 314–315), each stalk with process ending in two tips each. Thoracic furrow absent. Clypeus unmodified. Sternum wider than long (0.62/0.50), unmodified. Abdomen cylindrical, widened posteriorly, pointed dorso-posteriorly.

CHELICERAE. As in Fig. 314View FIGURES 314–315, with pair of simple weakly sclerotized processes proximally.

PALPS. As in Figs 312–313View FIGURES 312–313, symmetric; coxa with small but distinct retrolateral-ventral apophysis, trochanter with retrolateral-dorsal process, femur with rounded dorsal process proximally; femur-patella joints strongly shifted towards prolateral side; tarsus with long whitish dorsal process with tarsal organ near tip; procursus complex, distal part apparently hinged, proximal part with two ventral processes, without ventral parallel ridges; brush of hairs dorsally between proximal and distal parts; distal part wide and flat; genital bulb oval, with two processes lying parallel to each other: semitransparent embolus and slightly sclerotized long bulbal apophysis (‘appendix’; ba in Fig. 312View FIGURES 312–313).

LEGS. Without spines and curved hairs, few vertical hairs; retrolateral trichobothrium of tibia 1 at 3%; tibia 1 without prolateral trichobothrium (present on other tibiae); tarsus 1 with ~30 pseudosegments, only distally a few fairly distinct.

Male (variation). Tibia 1 in 3 other males: 9.0, 9.5, 9.9.

Female. In general similar to male ( Figs 306–307View FIGURES 300–307) but eye triads on low humps, closer together (distance PME-PME 210 µm) and clypeus not darkened (only dark band between eye triads). Carapace pattern in most females like in males (but with light median line dividing dark mark), in one female different ( Fig. 307View FIGURES 300–307). Tibia 1 in 5 females: 6.7–8.0 (mean 7.5). Epigynum weakly sclerotized ( Fig. 319View FIGURES 316–321), with short folded scape. Internal genitalia with small median pocket and pair of large pore plates ( Figs 315View FIGURES 314–315, 321View FIGURES 316–321).

Distribution. Known from type locality only ( Fig. 350View FIGURE 350).

Natural history. The spiders were found in a disturbed forest ( Fig. 341View FIGURES 336–341), on the undersides of green leaves about 1–2 m above the ground. The indistinct domed webs were connected to the undersides of the leaves at their apex, with a few silk lines extending beyond the leaves. Most specimens were found in a dryer patch of forest higher up on the hill.