Arnapa arfak Huber

Huber, Bernhard A. & Carvalho, Leonardo S., 2019, Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae), Zootaxa 4546 (1), pp. 1-96: 16-19

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Arnapa arfak Huber

sp. n.

Arnapa arfak Huber  sp. n.

Figs 39–40View FIGURES 39–44, 45–49View FIGURES 45–49, 70View FIGURES 70–74, 75–77View FIGURES 75–80

Gen.n. Ind96: Eberle et al. 2018 (molecular data); Huber et al. 2018: fig. 2.

Type material. INDONESIA: ♂ holotype, ZFMK (Ar 20604), West Papua, Arfak Mountains, forest above Mokwam (1.112°S, 133.911°E), 1600 m a.s.l., near ground, 5–6.xi.2009 (S. Sutono)GoogleMaps  .

Other material examined. INDONESIA: 2♂ 7♀ 2 juvs, ZFMK (Ar 20605), and  1♀ 4 juvs in pure ethanol,

ZFMK (Ind196), same data as holotype. 3♂ 1♀, ZFMK (Ar 20606), West Papua, Arfak Mountains, forest along stream above Syobri village (1.114°S, 133.906°E), 1680 m a.s.l., among rocks, 7.xi.2009 (S. Sutono).

Etymology. The species name is derived from the type locality; noun in apposition.

Diagnosis. Easily distinguished from known congeners by wide procursus (especially in dorsal view; Fig. 47View FIGURES 45–49) and by roundish epigynum with strong transversal ridges anteriorly ( Fig. 75View FIGURES 75–80); from most congeners also by armature of male chelicerae ( Fig. 48View FIGURES 45–49; two distinctive distal apophyses on each side directed towards median; similar only in A. manokwari  ); also by bulbal process ( Fig. 49View FIGURES 45–49; with wide flat curved sclerite) and by distal elements of procursus ( Fig. 47View FIGURES 45–49).

Description. Male (holotype). MEASUREMENTS. Total length 2.2, carapace width 1.05. Distance PME-PME 110 µm; diameter PME 110 µm; distance PME-ALE 120 µm; distance AME-AME 20 µm, diameter AME 50 µm. Leg 1: 28.2 (6.7 + 0.4 + 7.0 + 11.8 + 2.3), tibia 2: 3.9, tibia 3: 2.8, tibia 4: 4.0; tibia 1 L/d: 78.

COLOR (in ethanol). Carapace ochre-orange with wide dark brown bands laterally and medially, including posterior part of ocular area; clypeus with pair of dark bands between eye triads and rim; sternum light brown to orange; legs brown, with indistinct darker rings on femora (distally) and tibiae (proximally). Abdomen gray, dorsally densely covered with dark marks, ventrally with slightly darker mark in gonopore area, dark median band behind gonopore, light brown area and pair of large dark ventro-lateral marks in front of spinnerets.

BODY. Habitus as in Fig. 39View FIGURES 39–44. Ocular area elevated, thoracic furrow present; clypeus unmodified, only strongly protruding towards rim. Sternum wider than long (0.70/0.46), unmodified.

CHELICERAE. As in Fig. 48View FIGURES 45–49, with several sclerotized cones in distinctive arrangement, no modified hairs, with stridulatory ridges.

PALPS. As in Figs 45–46View FIGURES 45–49; coxa unmodified, trochanter with short ventral apophysis, femur strongly widened, with large rounded retrolateral process proximally, with stridulatory pick (modified hair) proximally, patella ventrally reduced (not closed), tarsus small, procursus with large flat dorsal apophysis proximally, with ventral groove (rather than distinct ventral pocket), distally with sclerotized and membranous elements ( Fig. 47View FIGURES 45–49); genital bulb large, with complex process ( Fig. 49View FIGURES 45–49).

LEGS. Without spines, few vertical hairs; with weakly curved hairs on all tibiae and metatarsi (especially anterior legs); retrolateral trichobothrium of tibia 1 at 4.5%; prolateral trichobothrium present on all tibiae; tarsus 1 with>30 pseudosegments, distally distinct.

Male (variation). Number of sclerotized cones on chelicerae slightly variable but pattern consistent. Tibia 1 in 5 males (incl. holotype): 6.5–7.0 (mean 6.8).

Female. In general similar to male but sternum darker brown. Tibia 1 in 7 females: 4.2–4.9 (mean 4.7). Epigynum anterior plate roundish, with strong transversal ridges in anterior part ( Fig. 75View FIGURES 75–80); posterior plate large, also with transversal ridges. Internal genitalia with distinctive rounded frontal processes at both sides of ‘valve’ and with internal pockets, with large pore plates and pair of lateral sclerites ( Figs 70View FIGURES 70–74, 76–77View FIGURES 75–80).

Distribution. Known from type locality only ( Fig. 343View FIGURE 343).

Natural history. This species was found in humid forest ( Fig. 336View FIGURES 336–341) in small domed webs at the basis of trees and in small holes in the ground.