Arnapa tolire Huber

Huber, Bernhard A. & Carvalho, Leonardo S., 2019, Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae), Zootaxa 4546 (1), pp. 1-96: 22-24

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Arnapa tolire Huber

sp. n.

Arnapa tolire Huber  sp. n.

Figs 41–42View FIGURES 39–44, 60–64View FIGURES 60–64, 73View FIGURES 70–74, 84–86View FIGURES 81–89, 90–94View FIGURES 90–97

Gen.n. Ind117: Eberle et al. 2018 (molecular data); Huber et al. 2018: fig. 2.

Type material. INDONESIA: ♂ holotype, ZFMK ( Ar 20611), Ternate , at Danau (Lake) Tolire (0.830°N, 127.311°E), 180–220 m a.s.l., 12.xi.2009 (S. Sutono)GoogleMaps  .

Other material examined. INDONESIA: 5♂ 11♀, ZFMK (Ar 20612–13) and  1 juv. in pure ethanol, ZFMK (Ind209), same data as holotypeGoogleMaps  .

Etymology. The species name is derived from the type locality; noun in apposition.

Diagnosis. Distinguished from known congeners by armature of male chelicerae ( Fig. 63View FIGURES 60–64; ~3–4 large lateral apophyses plus proximal patch of 3–4 smaller apophyses), by more slender palpal segments in general ( Figs 60– 61View FIGURES 60–64) and procursus in particular, the latter with distinctive prolateral sclerite distally ( Fig. 62View FIGURES 60–64; with pointed side branch), by shape of bulbal process ( Fig. 64View FIGURES 60–64), by shape of epigynum ( Figs 84View FIGURES 81–89, 94View FIGURES 90–97; pair of anterior areas with strong transversal ridges and pair of rounded posterior projections), and by elongate pore plates ( Figs 73View FIGURES 70–74, 86View FIGURES 81–89).

Description. Male (holotype). MEASUREMENTS. Total length 2.1, carapace width 0.95. Distance PME-PME 75 µm; diameter PME 85 µm; distance PME-ALE 60 µm; distance AME-AME 15 µm, diameter AME 40 µm. Leg 1: 24.6 (5.9 + 0.4 + 6.1 + 10.2 + 2.0), tibia 2: 3.4, tibia 3: 2.4, tibia 4: 3.3; tibia 1 L/d: 81.

COLOR (in ethanol). Carapace ochre-yellow with dark brown bands laterally and wide band medially including ocular area; clypeus with pair of dark bands below eye triads; sternum ochre-yellow with pair of light brown bands on both sides of median line; legs brown, with indistinct darker rings on femora (subdistally) and tibiae (proximally and subdistally), tips of femora and tibiae whitish. Abdomen gray, dorsally densely covered with dark marks, ventrally with large brown mark in front of gonopore, dark median band behind gonopore, pair of brown external marks and pair of larger internal ventro-lateral marks in front of spinnerets.

BODY. Habitus as in Fig. 41View FIGURES 39–44. Ocular area elevated, thoracic furrow present; clypeus unmodified. Sternum wider than long (0.58/0.36), unmodified.

CHELICERAE. As in Fig. 63View FIGURES 60–64, each side with 3–4 large sclerotized cones along lateral margin and 3–4 smaller cones frontally proximally; without modified hairs; with stridulatory ridges.

PALPS. As in Figs 60–61View FIGURES 60–64; coxa unmodified, trochanter ventrally only slightly protruding, femur widened, with rounded retrolateral process proximally, with prolateral stridulatory pick (modified hair) proximally and stronger than usual hairs distally on ventral-retrolateral side, patella ventrally reduced (not closed), tibia longer than in known relatives, retrolateral trichobothrium in very proximal position, tarsus small, procursus with dorsal process proximally, ventrally without pocket, distally with distinctive sclerotized and membranous elements ( Fig. 62View FIGURES 60–64); genital bulb large, with complex process ( Fig. 64View FIGURES 60–64).

LEGS. Without spines, few vertical hairs; with curved hairs on tibiae and metatarsi 1–2 (few on legs 2); retrolateral trichobothrium of tibia 1 at 11%; prolateral trichobothrium present on all tibiae; tarsus 1 with ~30 pseudosegments, distally distinct.

Male (variation). Size and number of sclerotized cones on chelicerae slightly variable but pattern consistent. Tibia 1 in 5 other males: 5.2–6.1 (mean 5.7).

Female. In general similar to male ( Fig. 42View FIGURES 39–44), also with curved hairs on legs. Chelicerae without stridulatory ridges ( Fig. 92View FIGURES 90–97). Tibia 1 in 9 females: 3.8–4.6 (mean 4.1). Palpal tarsal organ exposed ( Fig. 93View FIGURES 90–97). Epigynum anterior plate square-shaped ( Figs 84View FIGURES 81–89, 94View FIGURES 90–97), with two areas of deep transversal ridges anteriorly laterally, medially whitish, posterior rim with pair of rounded projections; with separate brown plate in front of epigynum; posterior plate very short but wide. Internal genitalia with distinctive lateral folds, narrow anterior ‘valve’, and elongated pore plates ( Figs 73View FIGURES 70–74, 86View FIGURES 81–89). ALS with only two spigots each (confirmed by SEM).

Distribution. Known from type locality only ( Fig. 343View FIGURE 343).

Natural history. The spiders were found in a degraded forest in a ravine leading into the lake; their domed webs were built among rocks near the ground and under large dead palm leaves lying on the ground.