Remexudes toompani, Błażewicz-Paszkowycz, Magdalena & Bamber, Roger N, 2007

Błażewicz-Paszkowycz, Magdalena & Bamber, Roger N, 2007, Parapseudid tanaidaceans (Crustacea: Tanaidacea: Apseudomorpha) from Eastern Australia, Zootaxa 1401, pp. 1-32 : 20-31

Remexudes toompani sp. nov. ( Figs. 10–12 )

Diagnosis. As for the genus (currently monotypic).

Material examined: Holotype: female, NMV J53140 , BSS 110, Central Bass Strait, S km E of Cape Edie, Robbins Island, 40°41.8'S, 145°07'E, depth 16m, fine shelly sand, 0 3 Nov 1980. Paratypes: 49 females, NMV J55294 ; 1 female dissected on slides, NMV J55295 , same locality as holotype. Other Material: 1 specimen, BSS 66, Western Bass Strait, 82 km SSE of Port Fairy, 39°06’S, 142°29’E), depth 630 m, very fine sand, 9 Oct 1980; 1 specimen, BSS 55, Western Bass Strait, 32 km SSW of Cape Otway, 39°06’S, 43°26’E, coarse carbonate sand, depth 85 m, 8 Oct 1980; 1 specimen, BSS 33 I, Eastern Bass Strait, 50 km NE of Babel Island, 39°40.3’S, 148°46.5’E, depth 293 m, rock, coarse 27 Mar 1979; 2 specimens, BSS 35 S, Eastern Bass Strait, 70 km ENE of North Point, Flinders Island, 39°28.4’S, 148°41.8’E, depth 110 m, coarse sand, 28 Mar 1979; 2 specimens, BSS 36, Eastern Bass Strait, 82 km ENE of North Point, Flinders Island, 39°27.7’S, 148°41.4’E, depth 293 m, coarse sand, 28 Mar 1979; 3 specimens, BSS 46 I, Eastern Bass Strait, 116 km ESE of Seaspray, 38°52.2’S, 148°20.6’E), depth 110 m, coarse sand, 31 Mar 1979; 3 specimens, BSS 76, Western Bass Strait, 42 km NW of Cape Farewell, King Island, 39°19’S, 143°38’E), depth 95 m, coarse sand, carbonate, 10 Oct 1980; 4 specimens, BSS 52, Western Bass Strait, 11 km SW of Cape Otway, 38°57’S, 143°27’E, depth 49 m, coarse sand, 8 Oct 1980; 4 specimens, BSS 61, Western Bass Strait, 32 km WSW of Cape Otway, 39°00’S, 143°11’E, depth 68 m, coarse sand, 8 Oct 1980; BSS 64, Western Bass Strait, 79 km SSE of Port Fairy, 39°02’S, 142°38’E, depth 119, fine sand, 8 Oct 1980; 7 specimens, BSS 85, Western Bass Strait, 48 km WNW of Stokes Point, King Island, 40°3’S, 143°24’E, depth 122 m, coarse sand, 11 Oct 1980; 8 specimens, BSS 183, Western Bass Strait, 35 km SSW of Cape Otway, 39°07.0’S, 143° 14.6’E, depth 84 m, coarse sand, 20 Nov 1981; 14 specimens, BSS 196, Western Bass Strait, 6 km W of Currie, King Island (39°54.7’S, 143°43.4’E), depth 49 metres, coarse sand, 21 Nov 1981; 55 specimens, BSS 201, Central Bass Strait. 65 km S of Cape Schanck. 39°08.3’S, 144°43.9’E, depth 66 m, coarse sand, 23 Nov 1980; numerous specimens, BSS 203, Central Bass Strait, 44 km NE of Cape Wickham, King Island, 39°22.0’S, 144°18.3’E, depth 60 m, coarse sand, 23 Nov 1981; numerous specimens BSS 75, Western Bass Strait, 42 km NW of Cape Farewell, King Island (39°17’S, 143°39’E), depth 86 m, coarse sand, carbonate, 10 Oct 1980; numerous specimens, BSS 76, Western Bass Strait, 42 km NW of Cape Farewell, King Island (39°19’S, 143°38’E), depth 95 m, coarse sand, 10 Oct 1980; numerous specimens, BSS 77, Western Bass Strait, 47 km NW of Cape Farewell, King Island, 39°20’S, 143°34’E, depth 95 m, coarse sand, carbonate, 10 Oct 1980. Moreton Bay: 1 female with oostegites, 3 females without oostegites, 2 males, paratypes (QM W28150), 1 female with oostegites, dissected (NHM. 2006.1506) clean medium sand with some shell and ophiuroids, off South Passage East of Moreton Island, 27º17.65’S 153º26.95’E, depth 19.4 m, 35‰, 28.4ºC; 2 males, 5 females, paratypes (QM W28151), clean medium sand WNW of Flat Rock, off South Passage East of Moreton Island, 27º24.10’S 153º29.18’E, depth 10.7 m, 35‰, 28ºC; 4 males, 4 female without oostegites, paratypes (NHM 2006.1507– 1514), clean medium sand WNW of Flat Rock, off South Passage East of Moreton Island, 27º23.46’S 153º30.08’E, depth 20 m, 35‰, 28ºC; 1 male, 1 brooding female, 1 female with oostegites, 1 female without oostegites, paratypes (QM W28152) slightly muddy medium sand with holothurians, WNW of Flat Rock, off South Passage East of Moreton Island, 27º22.95’S 153º30.79’E, depth 26.6 m, 35‰, 28ºC. All 19 Feb 2005, van Veen grab; coll. R.N.B., 2 males, 5 females (4 of these with oostegites), paratypes, (QM W28153) clean medium sand East of Moreton Island, 27º18.19’S 153º27.56’E, depth 20.6 m, 35‰; 1 female without oostegites, clean medium sand East of Moreton Island, 27º18.37’S 153º26.77’E, depth 9.9 m, 35‰; both 22 Feb 2005, van Veen grab; coll. R.N.B.

Etymology. “ Toompani ” is the Australian aboriginal name for a beach on Moreton Island, in the vicinity of the Queensland type locality.

Description of female. Body ( Fig. 10 ), dorsoventrally flattened, small, holotype 7.2 mm long (tip of rostrum to posterior of pleotelson), 5.5 times as long as wide, tapering towards posterior. Cephalothorax subrectangular, just longer than wide, with pronounced blunt rostrum, naked; eyelobes distinct, eyes present; anterolateral margin of sternal carapace with three or four prominent spine­like apophyses directed anteriorly. Six free pereonites; pereonite 1 shortest, about one­third as long as cephalothorax, pereonites 2 and 3 progressively longer; pereonite 5 longest, twice as long as pereonite 1, pereonites 4 and 6 subequal, just shorter than pereonite 5 (all pereonites respectively 2.7; 2.2; 1,6; 1.3; 1.1 and 1.1 times as wide as long); pereonites generally laterally smoothly convex, tapering distally, naked; ventral hyposphenia present on posterior segment of cephalothorax (cheliped segment) and pereonite 1. Pleon twice as long as pereonite 6, of five free subequal pleonites bearing pleopods; pleonites five times as wide as long, pleonite 1 bearing continuous row of plumose setae, other pleonites without setae. Pleotelson rectangular, elongate, just less than half length of pleon, 1.4 times as long as wide, with one seta on each side and occasional pairs of dorsal setae.

Antennule ( Fig. 11 A). Peduncle 4­articled, article 1 stout, just over twice as long as wide, with numerous median and distal setae as figured; article 2 less than half as long as article 1, with outer median and inner distal tufts of setae; article 3 one third length of article 2, as long as wide, with distal setae three times as long as article length; fourth article just smaller than third, naked. Main flagellum of 12 segments, aesthetascs on segments 3, 4, 6, 7, 9; accessory flagellum of 10 segments.

Antenna ( Fig. 11 B). Inner process on proximal peduncle article with smooth margin; article 2 with two distal setae adjacent to attachment of elongate squama bearing up to 14 marginal setae; peduncle article 3 shorter than wide, with one seta; article 4 as long as wide; article 5 longer than wide. Flagellum of eleven segments bearing long outer setae.

Mouth parts. Labrum depressed centrally, laterally setose ( Fig. 11 C). Left mandible ( Fig. 11 D) with strong, crenulated pars incisiva, lacinia mobilis crenulated, setiferous lobe with five, distally­serrated setae, pars molaris slender, blunt with fine denticulation; mandibular palp of three articles, proximal article just longer than wide with numerous distal setae, article 2 twice as long as wide with four simple setae, article 3 with 5 short spines, 2 long and two simple setae. Right mandible ( Fig. 11 E) as left but without lacinia mobilis. Maxillule ( Fig. 11 F) inner endite with outer apophysis and inner margins finely setose, with five variously armed setae distally; outer endite with nine distal spines and two subdistal setae; palp of two articles, distally with six setae (Fig. F’). Maxilla ( Fig. 11 G) outer lobe of outer endite with two distally setulated setae on outer margin; inner lobe of movable endite with two setulated and five serrated setae. Outer lobe of inner endite with four simple, four setulated, and three multifurcate spiniform setae. Inner lobe of fixed endite with row of six setulated setae and row of numerous bifurcated setae. Labium ( Fig. 11 H) with serrated outer margin and setulose inner margin, palp with fine lateral setules and three simple distal setae. Epignath distal seta simple ( Fig. 11 J).

Maxilliped ( Fig. 11 I). Basis extended into outer acute distal apophysis; palp article 1 with one spine on outer margin and with two simple setae on inner margin; palp article 2 little longer than wide, with ten longer simple setae and numerous rows of short setae on inner margin; outer margin with five simple setae distally (one very short); palp article 3 as long as wide, with three vertical rows of simple setae; palp article 4 mounted subdistally, with two vertical rows of setae.

Cheliped ( Fig. 12 A) slender. Basis more than twice as long as wide, dorsally with five setae, ventrally with proximal seta and pair of distal setae. Exopodite present, 3­articled, second article naked, short, distal article with four plumose setae. Merus elongate, distal ‘shoulder’ well developed bearing brush of numerous distal setae. Carpus slender, over 7 times as long as wide, with some setae all along article. Chela slender, ventral margin with one medial seta and group of setae distally; six setae near articulation of fixed finger, six setae ventrally and nine setae on inner margin; dactylus with no apophyses on cutting edge, curved distally.

Pereopod 1 ( Fig. 12 B) of swimming­type. Basis stout, twice as long as wide, dorsal margin bearing five simple setae, row of nine simple setae distally, small ventrodistal spine; exopodite present, 3­articled, article 2 naked, article 3 with four plumose setae. Ischium with three simple ventrodistal setae. Merus widening distally, four ventral setae, two stout ventrodistal spines and more slender dorsodistal spine adjacent to three simple setae. Carpus compact, slightly wider than long, with dorsal and ventral marginal setae, two dorsodistal blunt spines and three ventral blunt spines. Propodus as long as basis and 1.6 as long as wide, tapering distally with two dorsal, two dorsodistal slender blunt spines, six ventral slender blunt spines interspersed with single tapering setae. Dactylus stout, with middorsal fine seta, three ventral denticulations and one seta at claw insertion; claw short.

Pereopod 2 ( Fig. 12 C) of swimming­type. Basis twice as long as wide with long ventrodistal setae. Ischium half as long as wide with long ventrodistal setae. Merus as long as carpus, triangular and with long ventral setae and one ventrodistal slender blunt spine; two ventral rows of setae, distal row with blunt spine. Carpus similar size to merus, with five blunt spines in ventral row and four spines with dorsodistal row interspersed with single seta. Propodus oval, twice as long as wide, with five blunt spines ventrally, 13 setae dorsally and two spines dorsodistally. Dactylus with small distal seta, claw simple.

Pereopod 3 ( Fig. 12 D) similar to pereopod 2 but smaller, basis less setose, merus and carpus much more slender; merus with three spines ventrally and two spines dorsally, distal spines on carpus more elongated.

Pereopod 4 ( Fig. 12 E) similar to pereopod 3 but basis much stouter, carpus and merus more elongate.

Pereopod 5 as pereopod 4.

Pereopod 6 ( Fig. 12 F) similar to pereopod 4 but basis with row of over 20 plumose setae dorsally and three plumose setae ventrally. Ischium with five ventral setae. Merus with two dorsal setae. Carpus with three plumose setae dorsally. Propodus distally with row of leaf­like spines surrounding medial and distal part of article.

Pleopods ( Fig. 12 G) all alike. Basis with six inner and eight outer plumose setae. Endopod 0.7 times as long as exopod; both slender, respectively with 17 and 20 plumose setae.

Uropod ( Fig. 12 H) biramous. Basis with four setae distally. Exopod twice 0.2 times as long as endopod, reaching to end of third segment of endopod. Endopod elongate, filiform, multisegmented.

Male. Generally as female, no dimorphism of cheliped; sexual dimorphism shown in antennae and antennules. Antennule main flagellum, segment 1 with two bands of 16 (proximal) and 12 (distal) aesthetascs, succeeding segments respectively with 16, 20, 20, 16, 16, 10 and 4 aesthetascs; antenna flagellum proximal segment with two diffuse bands of 18 (proximal) and 9 (distal) aesthetascs, succeeding segment s respectively with 16, 20, 20, 18, 18, 16, 16, 14 and 10 aesthetascs. Pereonite 6 of the male with midventral penial tubercle.

Remarks. There are distinct affinities between the present taxon and the Pakistanapseudes ­group of genera, and with Saltipedis , not only the gross morphology but also notably in the slender cheliped and the wide, flattened distal articles of pereopod 1, the inflated bases to pereopods 4 to 6, with dense plumose setae on the dorsal margin of the basis of pereopod 6 ( Saltipedis ), the spine­like outer apophysis on the basis of the maxilliped (Australian Saltipedis ), the outer apophysis on the maxillule inner endite (both genera) and the inner apophysis on the labial palp ( Pakistanapseudes ­group); the lateral rows of plumose setae on pleonite 1 are also found in Saltipedis , although not in Pakistanapseudes sensu lato. Remexudes toompani gen. et sp. nov. is distinct from all of these taxa in having the propodus of pereopod 1 twice as long as wide, and from all other parapseudids in having a flattened “swimming­type” morphology to the distal articles of pereopod 2, after which the genus is named. These features of the elongate pereopod 1 propodus and the flattened pereopod 2 distal articles allow recognition of this species in samples, without dissection.

Genus Saltipedis Gu ţ u, 1995

Saltipedis nugoris n.sp. ( Figs 13–15 )

Diagnosis: Saltipedis with large hyposphenium between the chelipeds and a smaller one on each of the first two pereonites; first pleonite with entire dorsal row of setae. Second peduncle article of antenna 1.5 times as long as wide; basis and proximal palp article of maxilliped with sharp outer apophyses. Pereopod 2 with small spines on the basis and long ventrodistal spine on the merus. Dorsal edge of basis pereopod 1 with distinct proximal serration.

Material examined: Holotype: female, NMV J55296 , BSS 169, Eastern Bass Strait, 85 km NE of North Point, Flinders Island 39 0 2 24 S, 148 30 36 E, depth 120 m, muddy sand, 15 Nov 1981. Paratypes: 85 females, NMV J55297 ; 1 female dissected on slides, NMV J47122 , same locality as holotype. Other material. 7 females, J53144 , Australia, Victoria, Crawfish Rock, Westernport Bay, depth 12 m, 25 Nov 1971; 1 specimen, BSS Eastern Bass Strait, 85 km NE of North Point, Flinders Island 39 0 2 24 S, 148 30 36 E, depth 95 m, coarse sand carbonate, 15 Nov 1981; 1 specimen, BSS 166, Eastern Bass Strait, 20 km SSW of Babel Island, 40°06.8’S, 148°24.3’E, depth 22 m, coarse shell, 14 Nov 1981; 1 specimen, BSS 168, Eastern Bass Strait, 45 km NE of North Point, Flinders Island, 39º31,2’S, 148°24.4’E, depth 40 m, medium sand, 15 Nov 1981; 1 specimen, BSS 47, Central Bass Strait, 36 km S of Cape Otway, 39º03’S, 143°51’E, medium sand, carbonate, depth 85 m, 7 Oct 1980; 1 specimen, BSS 204, Central Bass Strait, 35 km NNE of Cape Wickham, King Island, 39º16, 6’S, 144°05.4’E), sandy shell, depth 82 m, 23 Nov 1981; 1 specimen, BSS 157, Central Bass Strait, 65 km ENE of Cape Rochon, Three Hummock Island, 40°10.9’S, 145°44.3’E, shelly sand, depth 75 m, 13 Nov 1981; 2 specimens, BSS 159, Central Bass Strait, 57 km S of Rodondo Island, 39°43.5’S, 146°18.8’E, depth 80 m, muddy shell, 3 Nov 1981; 2 specimens, BSS 117, Central Bass Strait, 20 km NNE of North Point, 40°38.0’S, 145°23’E, depth 36 m, muddy shell grit, 4 Nov 1980; 2 specimens, BSS 158, Central Bass Strait, 66 km S of Rodondo Island, 39°49.5’S, 146°18.5’E, sand­silt­mud, depth 82 m, 13 Nov 1981; 2 specimens, BSS 65, Western Bass Strait, 76 km SSE of Port Fairy, 39°05’S, 142°33’E, depth 207 m, very fine sand, 9 Oct 1980; 2 specimens, BSS 179, Eastern Bass Strait, 15 km E of Cape Wellington, Wilsons Promontory, 39°03.2’S, 146°39.5’E, muddy fine, depth 55 m; 4 specimens, BSS 114, 14 Central Bass Strait, 6 km NE of Stanley, 40°48.8’S, 145°22’E, fine sand, depth 22 m, 4 Nov 1980; 4 specimens, BSS 115, Central Bass Strait, 5 km N of North Point, 40°40.3’S, 145°15’E, 33 medium shell, depth 33 m, 4 Nov 1980; 5 specimens, BSS 64, Western Bass Strait, 79 km SSE of Port Fairy, 39°02’S, 42°38’E, coarse sand, depth 119, 9 Oct 1980; 5 specimens, BSS 169, Eastern Bass Strait, 85 km NE of North Point, Flinders Island, 39°02.4’S, 148°30.6’E, muddy sand, depth 120 m, 15 Nov 1981; 5 specimens, BSS 160, Central Bass Strait, 25 km S of Deal Island, 39°43.7’S, 147°19.6’E, muddy shell, depth 59 m, 13 Nov 1981; 6 specimens, BSS 176, Eastern Bass Strait, 50 km SE of Port Albert, 38°54.3’S, 147°13.4’E, depth 58 m, coarse shell, 18 Nov 1981; 7 specimens, BSS 35, 35 Eastern Bass Strait, 70 km ENE of North Point, Flinders Island, 39°28.4’S, 148°41.8’E, depth 110 m, coarse sand, 28 Mar 1979; 7 specimens, BSS 207, Eastern Bass Strait, 28 km SSW of Mario, 37°59.0’S 148°27.0’E), muddy sand and fine shell, depth 51 m, 30 Jul 1983; 10 specimens, BSS 180, Central Bass Strait, 8 km S of South East Point, Wilsons Promontory, 39°12.9’S, 146°27.3’E), medium sand, depth 65 m, 18 Nov 1981; 3 specimens BSS 112, 112 Central Bass Strait, 23 km E of Cape Rochon, Hummock Island, 40°22.2’S, 145°17’E, depth 40 m, mainly sand, 3 Nov 1980; 13 specimens., BSS 181, Central Bass Strait, 26 km SE of Aireys Inlet, 38°39.8’S, 144°18.2’E, depth 79 m, very fine sand, 19 Nov 1981; 14 specimens., BSS 177, ’77 Eastern Bass Strait, 43 km SE of Port Albert, 38°53.7’S, 147°06.5’E; depth 58 m, coarse shell, 18 Nov 1981; 15 specimens, BSS 161, Central Bass Strait, 33 km S of Deal Island, 39°48.3’S, 147°19.2’E, depth 60 m, muddy sand, 14 Nov 1981; 18 specimens, BSS 163, 163 Eastern Bass Strait, 24 km NNE of Eddystone Point, 40°43.9’S, 148°32.5’E, depth 56, muddy sand, 14 Nov 1981; 18 specimens., BSS 113, Central Bass Strait, 47 km E of Cape Rochon, Three Hummock Island,, 40°23.8’S, 145°32’E, muddy sand, 3 Nov 1980; 23 specimens, BSS 162, Central Bass Strait, 25 km SW of Cape Frankland, Flinders Island, 40°09.2’S, 147°31.9’E, depth 51 m, sandy shell, 14 Nov 1981; 35 specimens., BSS 167, Eastern Bass Strait, 63 km E of North Point, Flinders Island, 39°44.8’S, 148°40.6’E, depth 124 m, muddy sand, 14 Nov 1981; 42 specimens., BSS 155, 55 Central Bass Strait, 38 km SW of Cape Paterson, 38°55.5’S, 145°17.0’E, depth 70 m, fine sand, 12 Nov 1981; 229 specimens., BSS 165, Eastern Bass Strait, 42 km SW of Babel Island, 40°14.4’S, 148°40.0’E, depth 60 m, fine sand, 14 Nov 1981; numerous specimens, BSS 36, Eastern Bass Strait, 82 km ENE of North Point, Flinders Island, 39°27.7’S, 148°41.4’E, depth 293m, coarse sand, 28 Mar 1979; numerous specimens, BSS 46, Eastern Bass Strait, 116 km ESE of Seaspray, 38°52.2’S, 148°20.6’E, 110 metres, coarse sand. 31 Mar 1979; numerous specimens, BSS 183, Western Bass Strait, 35 km SSW of Cape Otway, 39°07.0’S, 143°14.6’E, depth 84 m, coarse sand, 20 Nov 1981; numerous specimens, BSS 170, Eastern Bass Strait, 100 km NE of North Point, Flinders Island, 31°51.8’S, 148°26.5’E, depth 130 m, fine sand, 15 Nov 1981; numerous specimens, BSS 173, Eastern Bass Strait, 30 km N of North Point, Flinders Island, 39°26.3’S, 147°48.7’E, depth 49 m, medium sand, 17 Nov 1981; numerous specimens, BSS 78, Western Bass Strait, 48 km WNW of Cape Farewell, King Island, 39°22’S, 143°28’E), depth 104 m, medium sand carbonate, 10 Oct 1980; numerous specimens, BSS 164, Eastern Bass Strait, 37 km NNE of Eddystone Point, 40°43.8’S, 148°37.2’E, depth 67 m, muddy sand, 14 Nov 1981; numerous specimens, BSS 38, Eastern Bass Strait, 67 km ENE of North Point, Flinders Island, 39°22.4’S, 148°38.7’E, coarse sand, depth 73 m, 29 Mar 1979.

Etymology: From the Latin nugoris , meaning buffoonish, antic, referring to an initial determination of this material as S. incognita Bamber, 2005

Description of female: body ( Fig. 13 ) dorsoventrally flattened, holotype 6.3 mm long, 6.3 times as long as wide, tapering towards posterior. Cephalothorax subrectangular, wider than long, with large pointed rostrum, apex down­turned; eyelobes and eyes present. Six free pereonites, with anterolateral and posterolateral setae; pereonites 1 and 2 subequal, about half as long as cephalothorax; pereonite 3 less than twice as long as pereonite 2; pereonites 4–6 subequal, longest (as long as cephalothorax), (all pereonites respectively 2.9, 3.4, 1.7. 1.2, 1.1 and 1.0 times as wide as long). Large, forward­pointing hyposphenium midventrally between chelipeds and small hyposphenium on pereonites 1 and 2. Pleon shorter than pereonite 6, tapering posteriorly, of five free, progressively longer pleonites bearing pleopods; first pleonite 0.4 times as long as pleonite 5 and bearing entire dorsal row of plumose setae. Pleotelson ( Figure 8 E) rectangular, subequal to combined length of all pleonites, 2.5 times as long as wide; laterally with three evenly spaced setae on each side.

Antennule ( Fig. 14 A). Peduncle article 1 stout, with numerous setae along article, article 2 little longer than wide, with setae as figured. Flagella missing.

Antenna ( Fig. 14 B). Inner process on proximal peduncle article with serrate margin and spine; article 2 with three setae along article; elongate squama (twice as long as wide) bearing up to 11 marginal setae; peduncle article 3 shorter than wide with one spine; article 4 as long as wide, article 5 longer than wide. Flagellum of eight segments bearing long outer setae.

Mouth parts. Left mandible ( Fig. 14 C, C’) with strong, crenulated pars incisive, lacinia mobilis crenulated, setiferous lobe with four bifurcated setae and one distally plumose seta; pars molaris slender, blunt with fine denticulation along edge; mandibular palp 3­articled, article 1 as long as wide with six setae; article 2 three times as long as wide, with two simple setae; article 3 with 3 short and one long simple setae distally. Right mandible ( Fig. 14 D) as left but without lacinia mobilis and with 6 setae on setiferous lobe. Labium ( Fig. 14 G) with serrated outer margin; palp with fine lateral setules on inner margin and distally, outer margin serrated; three simple distal setae. Maxillule ( Fig. 14 E) inner endite with outer apophysis and outer margins finely setose, with five variously armed distal setae; outer endite with twelve distal spines and two subdistal setae; palp of two articles, distally with six setae (not drawn). Maxilla ( Fig. 14 F) typical for the genus. Epignath ( Fig. 14 H) distal seta serrated.

Maxilliped ( Fig. 14 I). Basis extended into outer distal apophysis; palp article 1 extended into outer distal apophysis, with one seta on outer margin and with one simple seta on inner margin; palp article 2 little longer than wide, compact, three rows of setae on inner margin, and with five simple setae dorsodistally; palp article 3 just wider than long with simple setae distally; fourth palp article mounted subdistally, with simple setae distally.

Cheliped ( Fig. 15 A) slender. Basis three times as long as wide, dorsally with three setae, ventrally with spine and pair of distal setae. 3­articled exopodite, article 2 naked, long; article 3 with five plumose setae. Merus elongate, distal ‘shoulder’ poorly developed, rounded and bearing brush of numerous distal setae. Carpus slender about 4 times as long as wide, with four setae along ventral margin. Chela compound; dorsal margin with one medial seta; one setae near articulation of fixed finger; fixed finger with about ten setae ventrally and five setae on inner margin. Dactylus with 5 spines but without apophyses on cutting edge, curved distally; with paired setae dorsodistally.

Pereopod 1 ( Fig. 15 B) flattened (“swimming­type”). Basis stout, less than twice as long as wide, dorsal margin bearing four dorsodistal simple setae, and denticulation proximally; ventral margin with five spines and simple setae. 3­articled exopodite present; article 2 naked, article 3 with four plumose setae. Ischium naked. Merus widening distally, seven ventral setae, one stout ventrodistal spine and more slender dorsodistal spine adjacent to three simple setae. Carpus as long as merus, 1.5 times as long as wide, dorsal margin with five setae and one dorsodistal spine, ventral margin with three (occasionally four) ventral spines and six simple setae. Propodus as long as carpus, tapering distally, with dorsal setae and two dorsodistal slender blunt spines, six (occasionally five or seven) ventral slender spines interspersed with single tapering setae. Dactylus stout, with middorsal fine seta, three ventral denticulations and one seta at claw insertion; claw short.

Pereopod 2 ( Fig. 15 C). Basis twice as long as wide with long ventrodistal setae and three spines ventrally. Ischium half as long as wide with ventrodistal setae. Merus shorter than carpus, with three spines and eight long setae ventrally; one slender blunt spine and three setae ventrodistally. Carpus with two row of simple setae ventrally, one adjacent to four spines; two dorsodistal spines. Propodus oval, twice as long as wide, with row of setae, four blunt spines in ventral row and two dorsodistal spines; dactylus with small distal seta; claw simple.

Pereopod 3 ( Fig. 15 D) similar to pereopod 2, but ventral spines on basis smaller, no dorsodistal spine on merus.

Pereopod 4 ( Fig. 15 E) basis stouter, twice as long as wide, two ventrodistal setae. Ischium with eight simple seta and three spines ventrally (distalmost longest). Carpus nearly twice as long as merus, propodus as long as merus. Carpus and propodus with spines and simple setae in two rows ventrally, carpus with three long setae and one spine dorsodistally

Pereopod 5 ( Fig. 15 F) similar to pereopod 4.

Pereopod 6 ( Fig. 15 G) similar to pereopod 4 but basis with row of 18 plumose setae dorsally. Ischium with three ventral setae. Merus with six ventral setae.

Pleopods ( Fig. 14 H) all alike. Basis with five inner and five outer plumose setae, endopod little shorter than exopod, both slender, each with 12 plumose setae.

Uropod missing.

Remarks: Saltipedis nugoris sp. nov. shows close similarity to the southwest Australian species S. incognita Bamber, 2005 , and initially was identified as that species. Detailed analysis of the morphology has revealed characters which allow the distinction of these two species. The new species has three hyposphenia, a large one between the chelipeds and a smaller one on each of the first two pereonites ( S. incognita has only a single, large blade­like hyposphenium on the first pereonite). The antenna peduncle article 2 is longer at 1.5 times as long as wide (no longer than wide in S. incognita ) and there is a stronger spine on article 3. More conspicuous characters are the presence on pereopod 2 of small spines on the basis, and a long ventrodistal spine on the merus instead of the regular setae and a short spine found in S. incognita . The serrated dorsal edge of basis pereopod 1 has been never observed in any Saltipedis species, although S. forex Bamber, 2005 , also from southwestern Australia, has an irregular dorsal margin. These, the only three Australian species of Saltipedis , are clearly closely related.

Figures

FIGURE 8. Pakistanapseudes bassi sp. nov., paratype female, A, antennule; B, antenna; C, left mandible; D, right mandible incisor; E, mandible palp, F, maxillule; G, maxilla; H, labium; I, maxilliped; I’, maxilliped endite Scale line = 0.1 mm.

FIGURE 10. Remexudes toompani sp. nov., holotype female. Scale line = 1.0 mm.

FIGURE 11. Remexudes toompani sp. nov., paratype female, A, antennule; B, antenna; C, labrum; D, left mandible incisor; E, right mandible; F, maxillule; F’, maxillule palp; G, maxilla; H, labium; I, maxilliped; I’, maxilliped endite; J, epignath, Scale line = 0.1 mm.

FIGURE 12. Remexudes toompani sp. nov., paratype female, A, cheliped; B – D, pereopods 1 ­ 3 respectively; E and F, pereopods 4 and 6, respectively; G, pleopod, H, uropod. Scale line = 0.1 mm.

FIGURE 13. Salitipedes nugoris sp. nov., holotype female. Scale line = 1.0 mm

FIGURE 14. Salitipedes nugoris sp. nov., paratype female, A, proximal part of antennule; B, antenna; C, left mandible; C’, mandible palp; D, right mandible incisor; E, maxillule; F, maxilla; G, labium; H, epignath; I, maxilliped. Scale line = 0.1 mm.

FIGURE 15. Salitipedes nugoris sp. nov., paratype female, A, cheliped; B – G, pereopods 1 ­ 6 respectively; H, pleopod. Scale line = 0.1 mm.