Creagrutus, Gunther, 1864

Toledo-Piza, Mônica, Baena, Eduardo G., Dagosta, Fernando C. P., Menezes, Naércio A., Ândrade, Marcelo, Benini, Ricardo C., Bertaco, Vinicius A., Birindelli, José Luís O., Boden, Gert, Buckup, Paulo A., Camelier, Priscila, Carvalho, Fernando R., Castro, Ricardo M. C., Chuctaya, Junior, Decru, Eva, Derijst, Eddy, Dillman, Casey B., Ferreira, Katiane M., Merxem, Dimitri G., Giovannetti, Victor, Hirschmann, Alice, Jégu, Michel, Jerep, Fernando C., Langeani, Francisco, Lima, Flávio C. T., Lucena, Carlos A. S., Lucena, Zilda Margarete S., Malabarba, Luiz R., Malabarba, Maria Cláudia S. L., Marinho, Manoela M. F., Mathubara, Kleber, Mattox, George M. T., Melo, Bruno F., Moelants, Tuur, Moreira, Cristiano R., Musschoot, Tobias, Netto-Ferreira, André L., Ota, Rafaela P., T. Oyakawa, Osvaldo, Pavanelli, Carla S., Reis, Roberto E., Santos, Osmar, Serra, Jane Piton, Silva, Gabriel S. C., Silva-Oliveira, Cárlison, Souza-Lima, Rosana, Vari, Richard P. & Zanata, Angela M., 2024, Checklist of the species of the Order Characiformes (Teleostei: Ostariophysi), Neotropical Ichthyology (e 230086) 22 (1), pp. 1-548 : 133-143

CREAGRUTUS Günther, 1864

FCP Dagosta

Creagrutus Günther, 1864:339 . Type species: Leporinus muelleri Günther, 1859 . Type by monotypy. Gender: Masculine.

Creagrudite Myers, 1927:117 . Type species: Creagrudite maxillaris Myers, 1927 . Type by original designation. Gender: Masculine.

Creagrutops Schultz, 1944:327 . Type species: Creagrutops maracaiboensis Schultz, 1944 . Type by original designation. Gender: Masculine.

Creagrutus affinis Steindachner, 1880 Creagrutus affinis Steindachner, 1880:79 [27]. Type locality: Cauca. [ Colombia ]. Lectotype: NMW 67120.1. [Designated by Harold, Vari (1994:25)]. Creagrutus notropoides Meek, Hildebrand, 1912:68 . Type locality: Rio Indio, Panama. Holotype: FMNH 7574.

Creagrutus simus Meek, Hildebrand, 1913:85 . Type locality: Rio Cupe , Cituro, Panama. [Rio Tuira Basin]. Holotype: FMNH 7591 .

Creagrutus leuciscus Regan, 1913b:463 . Type locality: Rio Condoto, a tributary of the San Juan, a river of the Pacific slope in S. W. Colombia . Lectotype: BMNH 1913.10.1:6. [Designated by Harold, Vari (1994:26)]. Synonym of Creagrutus affinis following Harold, Vari (1994) and Vari, Harold (2001) .

Creagrutus londonoi Fowler, 1945a:3 , figs. 1–3. Type locality: Honda, Colombia . Holotype: ANSP 71610.

Distribution: 301, 302 (Harold, Vari , 1994; Vanegas-Rios et al., 2021).

Creagrutus amoenus Fowler, 1943

Creagrutus amoenus Fowler, 1943d:239 , fig. 18. Type locality: Florencia , Rio Orteguasa, Colombia . Holotype: ANSP 70499 .

Creagrutus boehlkei Géry, 1972b:63 , Pl. 4 (fig. 2). Type locality: Oriente del Ecuador (basin amazonien, localité non précisée). Holotype: ZSM 28428 a [not 28428, see Neumann, 2011:244].

Distribution: 306, 313, 316 ( Vari, Harold, 2001 ).

Remarks: Description based on four specimens, one holotype and three paratypes. One of the paratypes of the species (ANSP 70500, 62.8 mm SL) is a specimen of C. flavescens ( Vari, Harold, 2003) .

Creagrutus anary Fowler, 1913

Creagrutus anary Fowler, 1913b:552 , fig. 16. Type locality: Madeira River, about 200 miles east of Long. 62°20’W., Brazil. [Böhlke (1955a) proposed that the actual distance was 20 rather than 200 miles east of longitude 62°20’W. According to Vari, Harold (2001) such an interpretation shifts the type locality of C. anary further upstream along the Rio Madeira and places it significantly closer to the other collecting sites with exact locality information provided by the collector, E.A. Smith. The known distribution of C. anary is in agreement with Böhlke’s suggestion]. Holotype: ANSP 39290 (missing).

Distribution: 316 ( Vari, Harold, 2001 ).

Remarks: Description was based on the holotype and a single paratype, both collected in the Rio Madeira system. Böhlke (1984) reported that the holotype (ANSP 39290) was missing and that the single paratype (ANSP 39291) had been cleared and stained for bone. According to Vari, Harold (2001) the paratype (ANSP 39291) apparently had desiccated at some point prior to being stained and it is not clear whether it also had been cleared.

Creagrutus andaki Albornoz-Garzón, Acosta-Santos, Bogotá-Gregory, Agudelo-Córdoba, 2020

Creagrutus andaki Albornoz-Garzón, Acosta-Santos, Bogotá-Gregory, Agudelo-Córdoba, 2020:366 , figs. 1–3. Type locality: Colombia , Caquetá, Florencia, Río Caquetá basin, Río Hacha drainage, Río Sucre , km 61, bridge at old road to Neiva , 01°47’43.2”N, 75°38’48.9”W. Holotype: CIACOL 3462.

Distribution: 313 (Albornoz-Garzón et al., 2020).

Creagrutus ardilai Ardila Rodríguez, 2021

Creagrutus ardilai Ardila Rodríguez, 2021:10 , fig. 2B, 4A, 5C, 7A, 8C, 9C. Type locality: Colombia . Quebrada a 8 kilómetros del Municipio El Playón, Departamento de Santander. Holotype: CAR 919.

Distribution: 302 (Ardila Rodríguez, 2021).

Remarks: Named in honor to the author’s brother.

Creagrutus argos Ardila Rodríguez, 2020

Creagrutus argos Ardila Rodríguez, 2020b:7 , figs. 1–3, 5A, 7A, 8A, 9B–B1, 10A, 11A, 12A, 13, 14A. Type locality: Colombia, Quebrada El Laurel , 317 msnm, afluente del río Saldaña , cuenca alta del río Magdalena, municipio del Guamo, Departamento del Tolima. Holotype: CAR 900.

Distribution: 302 (Ardila Rodríguez, 2020b).

Creagrutus atratus Vari, Harold, 2001

Creagrutus atratus Vari, Harold, 2001:66 , figs. 24–25. Type locality: Colombia . Cundinamarca: Rio Meta basin, Rio Caqueza, tributary to Rio Negro , about 2 km upstream of village of Caqueza (latter at 4°25’N, 73°57’W). Holotype: ICN-MHN 4158. [Mojica, Agudelo-Zamora (2018:14)].

Distribution: 307 ( Vari, Harold, 2001 ).

Creagrutus atrisignum Myers, 1927

Creagrutus atrisignum Myers, 1927:116 . Type locality: Brazil: Rio Maranhão ( Upper Tocantins), Goyaz. Holotype: CAS 41339.

Distribution: 324 ( Vari, Harold, 2001 ).

Remarks: One of the paratypes is not a Creagrutus but rather another Stevardiinae .

Creagrutus barrigai Vari, Harold, 2001

Creagrutus barrigai Vari, Harold, 2001:71 , fig. 27. Type locality: Ecuador. Sucumbios: Rio Aguarico, near San Pablo de Kantesiya (0°15’18”S, 76°25’30”W). Holotype: MEPN 4621 .

Distribution: 313, 316 ( Vari, Harold, 2001 ).

Creagrutus beni Eigenmann, 1911

Creagrutus beni Eigenmann, 1911c:172 , Pl. 6 (fig. 2). Type locality: Villa Bella on Rio Beni . [ Bolivia]. Holotype: FMNH 54585 . [Holotype is in poor condition].

Distribution: 318, 319 ( Vari, Harold, 2001 ).

Creagrutus bolivari Schultz, 1944

Creagrutus bolivari Schultz, 1944:334 , fig. 49. Type locality: Rio Guárico and tributaries between San Sebastián and San Casimiro, Estado de Aragua. [ Venezuela]. Holotype: USNM 121497 .

Distribution: 307, 308 ( Vari, Harold, 2001 ).

Creagrutus brevipinnis Eigenmann, 1913

Creagrutus brevipinnis Eigenmann, 1913:10 . Type locality: Piedra Moler. [ Colombia ]. Holotype: FMNH 56095.

Distribution: 302 (Harold, Vari , 1994).

Creagrutus britskii Vari, Harold, 2001

Creagrutus britskii Vari, Harold, 2001:83 , fig. 31. Type locality: Brazil. Goias: Iaciara, Ribeirao Macambira (approximately 14°20’S, 46°45’W), near bridge on highway GO 112. Holotype: MZUSP 40537 .

Distribution: 322a, 324 (Meza-Vargas, 2014; Vari, Harold, 2001 ).

Creagrutus cacique Flausino, Lima, 2019

Creagrutus cacique Flausino, Lima, 2019:2, figs. 1,2. Type locality: Brazil: Mato Grosso: Campinápolis : rio Culuene , fish ladder of Paranatinga II hydroelectric dam, 13°50’58”S, 53°15’22”W. Holotype: ZUEC 15770 .

Distribution: 322 (Flausino, Lima 2019).

Creagrutus calai Vari, Harold, 2001

Creagrutus calai Vari, Harold, 2001:85 , fig. 32. Type locality: Colombia . Meta: Unnamed stream on Rancho El Viento, across Rio Meta from Puerto Lopez (4°08’N, 72°39’W). Holotype: ANSP 130527 .

Distribution: 307 ( Vari, Harold, 2001 ).

Creagrutus caucanus Eigenmann, 1913

Creagrutus caucanus Eigenmann, 1913:9 . Type locality: Paila. [ Colombia ]. Holotype: FMNH 56104.

Distribution: 302 (Harold, Vari , 1994).

Creagrutus cerritulus Netto-Ferreira, Vari, 2017

Creagrutus cerritulus Netto-Ferreira, Vari, 2017:369 , figs. 1–4. Type locality: Peru, San Martin, Tocache, Shapaja, Río Cachiyacu , tributary of Río Huallaga , 520 m elevation, 8°17’37”S, 76°21’05”W. Holotype: MUSM 54501 .

Distribution: 313 (Netto-Ferreira, Vari , 2017)

Creagrutus changae Vari, Harold, 2001

Creagrutus changae Vari, Harold, 2001:88 , fig. 34. Type locality: Peru. Huanuco: Provincia Pachitea, Rio San Alejandro, a tributary of Rio Sungarayacu , just above junction with Rio Sungarayacu (approximately 9°23’S, 75°11’W). Holotype: MUSM 8858 .

Distribution: 317 ( Vari, Harold, 2001 ).

Creagrutus cochui Géry, 1964

Creagrutus cochui Géry, 1964 h:56, fig. 12. Type locality: Upper Amazon region surrounding Iquitos. [ Peru]. Holotype: USNM 200426 .

Distribution: 316 ( Vari, Harold, 2001 ).

Creagrutus cracentis Vari, Harold, 2001

Creagrutus cracentis Vari, Harold, 2001:93 , fig. 36. Type locality: Brazil. Para : Rio Tapajos, sandy beaches on an island located 5 km S of Itaituba (latter locality at 4°17’S, 55°59’W). Holotype: MCP 15213 .

Distribution: 320, 322 (Dagosta, Pastana, 2014; Vari, Harold, 2001 ).

Creagrutus crenatus Vari, Harold, 2001

Creagrutus crenatus Vari, Harold, 2001:95 , fig. 37. Type locality: Venezuela. Lara: Quebrada Sanare in Yay (9°45’N, 69°43’W). Holotype: MHNLS 505 .

Distribution: 307 ( Vari, Harold, 2001 ).

Creagrutus dulima Albornoz-Garzón, Conde-Saldaña, García-Melo, Taphorn, Villa-Navarro, 2018

Creagrutus dulima Albornoz-Garzón, Conde-Saldaña, García-Melo, Taphorn, Villa-Navarro, 2018:342 , figs. 1, 2A, 3. Type locality: Colombia , Departamento Tolima, Ibagué, upper Río Magdalena drainage, quebrada Chumba , 4°29’06”N, 75°05’49”W, elevation 973 m a.s.l. Holotype: CZUT-IC 17663 .

Distribution: 302 (Albornoz-Garzón, Conde-Saldaña, García-Melo, Taphorn, Villa-Navarro, 2018).

Creagrutus ephippiatus Vari, Harold, 2001

Creagrutus ephippiatus Vari, Harold, 2001:98 , fig. 39. Type locality: Venezuela. Amazonas: Upper Rio Siapa, Campamento Siapa (Siapa Base Camp). Holotype: MBUCV-V-29068.

Distribution: 314 ( Vari, Harold, 2001 ).

Creagrutus figueiredoi Vari, Harold, 2001

Creagrutus figueiredoi Vari, Harold, 2001:101 , fig. 40. Type locality: Brazil. Distrito Federal: Rio Maranhao, upper Rio Tocantins basin, approximately 35 air km N of Brasilia (approximately 15°32’S, 47°49’W). Holotype: MZUSP 50542 .

Distribution: 324 ( Vari, Harold, 2001 ).

Creagrutus flavescens Vari, Harold, 2001

Creagrutus flavescens Vari, Harold, 2001:104 , fig. 41. Type locality: Ecuador. Pastaza: Rio Tiguino basin, Rio Tiguino N °3 (unnamed tributary of Rio Tiguino ; 1°07’35”S, 76°56’52”W). Holotype: MEPN 4622 .

Distribution: 313, 316 ( Vari, Harold, 2001 ).

Creagrutus gephyrus Böhlke, Saul, 1975

Creagrutus gephyrus Böhlke, Saul, 1975:25 , figs. 1–4. Type locality: Rio Aguarico at Santa Cecilia, backwater. [ Ecuador]. Holotype: ANSP 130516 .

Distribution: 313, 316 ( Vari, Harold, 2001 ).

Creagrutus gracilis Vari, Harold, 2001

Creagrutus gracilis Vari, Harold, 2001:111 , fig. 44. Type locality: Peru. Amazonas: Provincia Condorcanqui, Rio Santiago , at La Poza (4°01’S, 77°47’W). Holotype: LACM 41724-27 .

Distribution: 313, 316 ( Vari, Harold, 2001 ).

Creagrutus guanes Torres-Mejia, Vari, 2005

Creagrutus guanes Torres-Mejia, Vari, 2005:812 , fig. 1. Type locality: Colombia , Santander, Valle de San José, Río Magdalena system, Río Fonce , beside road from San Gil to Charalá , under bridge of road to Balneario Monas. Holotype: ICN-MHN 8520. [Mojica, Agudelo-Zamora, 2018:14].

Distribution: 302 (Torres-Meija, Vari , 2005).

Creagrutus gyrospilus Vari, Harold, 2001

Creagrutus gyrospilus Vari, Harold, 2001:113 , fig. 45. Type locality: Venezuela. Portuguesa: Rio Saguaz, Rio Guanare-Apure drainage, bridge near park on road to Chabasquen (= Paraiso de Chabasquen ; 9°23’30”N, 70°00’30”W). Holotype: INHS 69479 .

Distribution: 307 ( Vari, Harold, 2001 ).

Creagrutus hildebrandi Schultz, 1944

Creagrutus hildebrandi Schultz, 1944:330 , figs. 47–48. Type locality: Rio Táchira, 7 km north of San Antonio, Catatumbo system, Venezuela. Holotype: USNM 121482 .

Distribution: 303 (Harold, Vari , 1994).

Creagrutus holmi Vari, Harold, 2001

Creagrutus holmi Vari, Harold, 2001:116 , fig. 46. Type locality: Peru. Amazonas : Provincia Utcubamba , Bagua Grande , San Antonio, Quebrada Jaimito (approximately 5°47’S, 78°23’W). Holotype: MUSM 5670 .

Distribution: 312, 317 ( Vari, Harold, 2001 ).

Creagrutus hysginus Harold, Vari, Machado-Allison, Provenzano, 1994

Creagrutus hysginus Harold, Vari, Machado-Allison, Provenzano, 1994:975 , fig. 1. Type locality: Venezuela, Estado Sucre, Rio Guiria , near La Toma. Holotype: MBUCV-V-20310.

Distribution: 309 ( Vari, Harold, 2001 ).

Creagrutus ignotus Vari, Harold, 2001

Creagrutus ignotus Vari, Harold, 2001:122 , fig. 48. Type locality: Brazil. Mato Grosso: Riacho (small stream) 1, tributary to Rio Preto at road to San Francisco, Municipio de Diamantino ( Rio Arinos basin). Holotype: MZUSP 45310 .

Distribution: 320, 322a (Meza-Vargas, 2014; Vari, Harold, 2001 ).

Creagrutus jordanensis Ardila Rodríguez, 2021

Creagrutus jordanensis Ardila Rodríguez, 2021:9 , fig. 2A, 3B, 5B, 6B, 8B, 9B. Type locality: Colombia . Río Chicamocha , cuenca del río Magdalena, municipio de Jordán (425msmn), Departamento de Santander. Holotype: CAR 890.

Distribution: 302 (Ardila Rodríguez, 2021).

Creagrutus kunturus Vari, Harold, Ortega, 1995

Creagrutus kunturus Vari, Harold, Ortega, 1995:290 , figs. 1–2. Type locality: Peru, Departamento Amazonas , Provincia Condorcanqui , Cordílera del Condor , upper Río Comainas, 20 m upriver of Puesto de Vigilancia N °. 22 (hereafter cited as PV 22) (3°56’30”S, 78°24’20”W). Holotype: MUSM 5667 .

Distribution: 313, 316 ( Vari et al., 1995).

Remarks: Very similar to C. amoenus , but not syntopic with it.

Creagrutus lassoi Vari, Harold, 2001

Creagrutus lassoi Vari, Harold, 2001:128 , fig. 52. Type locality: Venezuela. Yaracuy: Rio Tupe, Rio Aroa drainage, approximately 12 km N of Aroa, on Highway 3 (10°30’19”N, 68°52’33”W). Holotype: INHS 60094 .

Distribution: 304 ( Vari, Harold, 2001 ).

Creagrutus lepidus Vari, Harold, Lasso, Machado-Allison, 1993

Creagrutus lepidus Vari, Harold, Lasso, Machado-Allison, 1993:352 , fig. 1. Type locality: Río Aroa basin, Quebrada El Charal, Finca El Jaguar, Sierra de Aroa, State of Yaracuy, Venezuela (approx. 10°32’N, 68°32’W). Holotype: MHNLS 9659 .

Distribution: 304 ( Vari et al., 1993).

Creagrutus machadoi Vari, Harold, 2001

Creagrutus machadoi Vari, Harold, 2001:134 , fig. 54. Type locality: Venezuela. Bolivar: Departamento Cedeno, upper Rio Caura, Cano Yumucukena , within 4 km of its mouth. Holotype: MCNG 18852 .

Distribution: 308 ( Vari, Harold, 2001 ).

Creagrutus maculosus Román-Valencia, García-Alzate , Ruiz-C., Taphorn B., 2010

Creagrutus maculosus Román-Valencia, García-Alzate , Ruiz-C., Taphorn B., 2010c:88, figs. 1–8. Type locality: Colombia : Meta State: Vista Hermosa municipality, Buenos Aires Village, Guadualito Creek drainage, 3°05’34”N, 73°51’53”W). Holotype: IUQ 2506 .

Distribution: 307 (Román-Valencia et al., 2010c).

Creagrutus magdalenae Eigenmann, 1913

Creagrutus magdalenae Eigenmann, 1913:8 . Type locality: Girardot. Holotype: FMNH 56088.

Distribution: 302 (Harold, Vari , 1994).

Creagrutus magoi Vari, Harold, 2001

Creagrutus magoi Vari, Harold, 2001:137 , fig. 55. Type locality: Venezuela. Bolivar. Rio Chaviripa, where crossed by bridge on road from Caicara to San Fernando de Atabapo (approximately 7°11’N, 66°18’W). Holotype: UF 80477 .

Distribution: 308 ( Vari, Harold, 2001 ).

Creagrutus manu Vari, Harold, 2001

Creagrutus manu Vari, Harold, 2001:140 , fig. 56. Type locality: Peru. Madre de Dios: Provincia Manu , Rio Manu basin, Parque Nacional Manu, Quebrada Agua Clara , first major quebrada on trail 1 leading from Pakitza (approximately 11°57’S, 71°17’W). Holotype: MUSM 8867.

Distribution: 318 ( Vari, Harold, 2001 ).

Creagrutus maracaiboensis (Schultz, 1944)

Creagrutops maracaiboensis Schultz, 1944:327 , figs. 45–46. Type locality: Río Negro below mouth of Río Yasa. Holotype: USNM 121531 .

Distribution: 318 (Harold, Vari , 1994).

Creagrutus mariposa Ardila Rodríguez, 2021

Creagrutus mariposa Ardila Rodríguez, 2021:5 , fig. 1, 3A, 5A, 6A, 8A, 9A. Type locality: Colombia . Quebrada La Ruitoca (En Pozo Azul), Vereda del Valle de Ruitoque , 965 mnsm, afluente del río de Oro, cuenca alta del río Lebrija , cuenca del río Magdalena, municipio de Floridablanca, Departamento de Santander. Holotype: CAR 877.

Distribution: 302 (Ardila Rodríguez, 2021).

Creagrutus martinezi Ardila Rodríguez, 2021

Creagrutus martinezi Ardila Rodríguez, 2021:12 , fig. 2C, 4B, 5D, 7B, 8D, 9D. Type locality: Colombia . Quebrada La Salitre , cuenca del río Magdalena, Municipio de San Vicente de Chucurí, Departamento de Santander. Holotype: CAR 905.

Distribution: 302 (Ardila Rodríguez, 2021).

Creagrutus maxillaris (Myers, 1927)

Creagrudite maxillaris Myers, 1927:118 . Type locality: Brazil: Sandbank on the Colombian border, Rio Negro, Cucuhy. Holotype: CAS 30419.

Distribution: 308, 314, 318 ( Vari, Harold, 2001 ).

Creagrutus melanzonus Eigenmann, 1909

Creagrutus melanzonus Eigenmann, 1909a:30 . Type locality: Crab Falls . [ Guyana]. Holotype: FMNH 52705 .

Distribution: 310, 311 ( Vari, Harold, 2001 ).

Creagrutus melasma Vari, Harold, Taphorn, 1994

Creagrutus melasma Vari, Harold, Taphorn, 1994:91 , fig. 1. Type locality: Venezuela, Estado Guarico, Parque Nacional Guatopo, Río Orituco , first bridge along road from Santa Teresa to Altagracia. Holotype: MBUCV-V-22198.

Distribution: 304, 306, 307, 309 ( Vari, Harold, 2001 ).

Creagrutus menezesi Vari, Harold, 2001

Creagrutus menezesi Vari, Harold, 2001:153 , figs. 63–64. Type locality: Brazil. Goias: Rio Tocantins basin, Ribeirao Paranoa do Meio , at road crossing 11 air km NNE of Formosa (approximately 15°25’S, 47°18’W). Holotype: MZUSP 50544 .

Distribution: 314 ( Vari, Harold, 2001 ).

Creagrutus meridionalis Vari, Harold, 2001

Creagrutus meridionalis Vari, Harold, 2001:157 , fig. 65. Type locality: Brazil. Mato Grosso: Ribeirao Chiqueirao (tributary of Rio Jangada that, in turn, drains into the Rio Cuiaba ), approximately 21 km W of Jangada (latter locality at 15°14’S, 56°29’W) on road to Barra do Bugres , Municipio de Jangada. Holotype: MZUSP 50546 .

Distribution: 343 ( Vari, Harold, 2001 ).

Creagrutus molinus Vari, Harold, 2001

Creagrutus molinus Vari, Harold, 2001:159 , fig. 66. Type locality: Brazil. Mato Grosso: Rio Araguaia basin, riacho (small stream) tributary to Ribeirao Lajeadinho, Municipio de Alto Araguaia (approximately 17°19’S, 53°12’W). Holotype: MZUSP 41461 . [Holotype specimen was in the process of replacing its dentition in both the right dentary and premaxilla].

Distribution: 324 ( Vari, Harold, 2001 ).

Creagrutus mucipu Vari, Harold, 2001

Creagrutus mucipu Vari, Harold, 2001:162 , fig. 68. Type locality: Brazil. Goias: Municipio de Minacu, Rio Tocantins, at port of Rubao (13°44’31”S, 48°08’29”W). Holotype: MCP 19511 .

Distribution: 322a, 324 (Meza-Vargas, 2014; Vari, Harold, 2001 ; Flausino, Lima, 2019:7).

Creagrutus muelleri (Günther, 1859)

Leporinus muelleri Günther, 1859:92 . Type locality: Andes of Western Ecuador. Lectotype: BMNH 1858.7 .25.42 (82.5 mm SL). [Designated by Vari, Harold (2001:166) ].

Distribution: 313, 316 ( Vari, Harold, 2001 ).

Creagrutus nigrostigmatus Dahl, 1960

Creagrutus nigrostigmatus Dahl, 1960a:353 . Type locality: Colombia . Sucre: Tuloviejo, Cano Pechilín (approx. 9°31’N, 75°35’W). Neotype: ICN-MHN 989. [Designated by Harold, Vari (1994:10)].

Distribution: 302 ( Vari, Harold, 2001 ).

Creagrutus nigrotaeniatus Dagosta, Pastana, 2014

Creagrutus nigrotaeniatus Dagosta, Pastana, 2014:572 , fig. 1–5. Type locality: Brazil, Mato Grosso State, Sapezal Municipality, rio Juruena, Bacia do rio Tapajós , ca. 12°50’59”S, 58°55’36”W. Holotype: LIRP 8967 .

Distribution: 320 (Dagosta, Pastana, 2014).

Creagrutus occidaneus Vari, Harold, 2001

Creagrutus occidaneus Vari, Harold, 2001:167 , fig. 70. Type locality: Peru. Madre de Dios: Provincia Manu, Parque Nacional Manu, Pakitza , beach along Rio Manu close to mouth of Quebrada Pachija (approximately 11°57’S, 71°17’W). Holotype: MUSM 8869.

Distribution: 316, 318 ( Vari, Harold, 2001 ).

Creagrutus ortegai Vari, Harold, 2001

Creagrutus ortegai Vari, Harold, 2001:170 , fig. 71. Type locality: Peru. San Martin: Provincia San Martin, Banda de Shilcayo, upper portions of Quebrada Choclin, tributary to Rio Huallaga , near town of Shapaja (latter locality at 6°36’S, 76°16’W). Holotype: MUSM 7498 .

Distribution: 316 ( Vari, Harold, 2001 ).

Creagrutus ouranonastes Vari, Harold, 2001

Creagrutus ouranonastes Vari, Harold, 2001:173 , fig. 72. Type locality: Peru. Apurimac: Provincia Aymaraes, Rio Chalhuanca (approximately 14°01’S, 73°11’W). Holotype: MUSM 8872 .

Distribution: 312 ( Vari, Harold, 2001 ).

Creagrutus paraguayensis Mahnert, Géry, 1988

Creagrutus paraguayensis Mahnert, Géry, 1988:5 , figs. 4–5, Pl. 3. Type locality: Rio Azotey à Cororo, près de l’embouchure du Rio Ypané, prov. Concepcion, Paraguay. Holotype: MHNG 2386-1 .

Distribution: 343 ( Vari, Harold, 2001 ).

Creagrutus paralacus Harold, Vari, 1994

Creagrutus paralacus Harold, Vari, 1994:13 , fig. 8. Type locality: Venezuela. Tachira: Rio Catatumbo basin, Rio Táchira, 7 km N of San Antonio. Holotype: USNM 121504 .

Distribution: 303 (Harold, Vari , 1994).

Creagrutus pearsoni Mahnert, Géry, 1988

Creagrutus pearsoni Mahnert, Géry, 1988:5 . Type locality: Popoi River, Upper Beni . [ Bolivia]. Lectotype: CAS 62272. [Designated by Vari, Harold (2001:180) ].

Piabina beni Pearson, 1924:45 , Pl. 10 (fig. 5). Type locality: Popoi River, Upper Beni . [ Bolivia]. Lectotype: CAS 62272. [Designated by Vari, Harold (2001:180) ]. A secondary junior homonym of Creagrutus beni Eigenmann, 1911 . Replaced by Creagrutus pearsoni Mahnert, Géry, 1988 .

Distribution: 318, 319 ( Vari, Harold, 2001 ).

Remarks: Replacement for Piabina beni Pearson, 1924 .

Creagrutus peruanus (Steindachner, 1876)

Piabina peruana Steindachner, 1876b:596 [46]. Type locality: Monterico. [ Peru]. Lectotype: NMW 19852 (87.0 mm SL). [Designated by Vari, Harold (2001:183) ].

Creagrutus nasutus Günther, 1876:400 . Type locality: Monterico, Peru. Lectotype: BMNH 1875.10.14.26 (78.9 mm SL). [Designated by Vari, Harold (2001:183) ]. The species was described on the basis of specimens collected at the same type locality of its senior synonym, Creagrutus peruanus .

Distribution: 312 ( Vari, Harold, 2001 ).

Remarks: Priority of Piabina peruana over Creagrutus nasutus given by date of publication (former is dated December 1875, but published in 1876 and latter is dated May, 1876.

Creagrutus petilus Vari, Harold, 2001

Creagrutus petilus Vari, Harold, 2001:184 , fig. 78. Type locality: Brazil. Rondonia: Rio Marco Rondon, Pimenta Bueno (approximately 11°29’S, 61°12’W). Holotype: MNRJ 14807 .

Distribution: 321 ( Vari, Harold, 2001 ).

Creagrutus phasma Myers, 1927

Creagrutus phasma Myers, 1927:117 . Type locality: Venezuela: mouth of the Curamuni, Rio Cassiquiare. Lectotype: CAS 44210. [Designated by Vari, Harold (2001:188) ].

Distribution: 308, 314 ( Vari, Harold, 2001 ).

Creagrutus pila Vari, Harold, 2001

Creagrutus pila Vari, Harold, 2001:189 , fig. 80. Type locality: Peru. Ucayali: Provincia Padre Abad, Rio Huacamayo, km 155 on Highway (Carratera Federico Basadre) from Pucallpa to Tingo Maria. Holotype: MUSM 8874 .

Distribution: 317 ( Vari, Harold, 2001 ).

Creagrutus planquettei Géry, Renno, 1989

Creagrutus planquettei Géry, Renno, 1989:1 , fig. 1. Type locality: Crique Japigny, Arataye, affluent de l’Approuague en amont de Pierrette, Guyane. Holotype: MNHN 1989-0031 .

Distribution: 311 ( Vari, Harold, 2001 ).

Creagrutus provenzanoi Vari, Harold, 2001

Creagrutus provenzanoi Vari, Harold, 2001:195 , fig. 84. Type locality: Venezuela. Amazonas: Upper Rio Cataniapo basin, small cano above Sarama Sota. Holotype: MBUCV-V-14392.

Distribution: 308 ( Vari, Harold, 2001 ).

Creagrutus runa Vari, Harold, 2001

Creagrutus runa Vari, Harold, 2001:198 , fig. 85. Type locality: Brazil. Amazonas : Rio Negro, Parana do Jacare (approximately 0°30’S, 66°30’W). Holotype: MZUSP 29888 .

Distribution: 314 ( Vari, Harold, 2001 ).

Creagrutus saxatilis Vari, Harold, 2001

Creagrutus saxatilis Vari, Harold, 2001:200 , fig. 86. Type locality: Brazil. Goias: Niquelandia, Rio Indaial, left bank tributary of Rio Maranhao, upper Rio Tocantins basin (14°12’S, 48°37’W). Holotype: MNRJ 14544 .

Distribution: 324 ( Vari, Harold, 2001 ).

Creagrutus seductus Vari, Harold, 2001

Creagrutus seductus Vari, Harold, 2001:202 , fig. 87. Type locality: Brazil. Mato Grosso: Upper Rio Araguaia basin, Corrego Fundo, Municipio de Barra do Garcas (approximately 15°53’S, 52°15’W). Holotype: MZUSP 51026 .

Distribution: 307, 314 ( Vari, Harold, 2001 ).

Creagrutus taphorni Vari, Harold, 2001

Creagrutus taphorni Vari, Harold, 2001:205 , fig. 88. Type locality: Venezuela. Guarico: Rio Orituco, Parque Nacional Guatopo, first bridge along road from Santa Teresa to Altagracia. Holotype: MBUCV-V-29288.

Distribution: 307 ( Vari, Harold, 2001 ).

Creagrutus tuyuka Vari, Lima, 2003

Creagrutus tuyuka Vari, Lima, 2003:583 , fig. 1. Type locality: Brazil, Amazonas, Rio Tiquié , comunidade de Cachoeira Comprida, beach below the cachoeira (0°15’44”N, 70°01’05.2”W). Holotype: MZUSP 65679 .

Distribution: 314 ( Vari, Harold, 2001 ).

Creagrutus ungulus Vari, Harold, 2001

Creagrutus ungulus Vari, Harold, 2001:209 , fig. 89. Type locality: Peru. Madre de Dios: Provincia Manu, Quebrada Soga , tributary of Rio Alto Madre de Dios, 1 km upstream from Erika (opposite Salvacion; approximately 12°53’S, 71°12’W). Holotype: MUSM 8878 .

Distribution: 318 ( Vari, Harold, 2001 ).

Creagrutus varii Ribeiro, Benine, Figueiredo, 2004

Creagrutus varii Ribeiro, Benine, Figueiredo, 2004:598 , figs. 1–10. Type locality: Brazil: Goiás, Município de Catalão, Rio São Marcos basin (tributary of Rio Paranaíba basin), Córrego da Prata (17°47’26”S, 47°35’48”W). Holotype: LIRP 4342 .

Distribution: 344 (Ribeiro et al., 2004).

Creagrutus veruina Vari, Harold, 2001

Creagrutus veruina Vari, Harold, 2001:213 , fig. 91. Type locality: Venezuela. Amazonas: Rio Cataniapo, 200 m above Las Pavas. Holotype: MBUCV-V-29072.

Distribution: 308 ( Vari, Harold, 2001 ).

Creagrutus vexillapinnus Vari, Harold, 2001

Creagrutus vexillapinnus Vari, Harold, 2001:215 , fig. 92. Type locality: Brazil. Amazonas : Rio Negro, Cachoeira de Sao Gabriel (0°08’S, 67°05’W). Holotype: MZUSP 29894 .

Distribution: 308, 314 ( Vari, Harold, 2001 ).

Creagrutus xiphos Vari, Harold, 2001

Creagrutus xiphos Vari, Harold, 2001:218 , fig. 93. Type locality: Venezuela. Bolivar: Sand bank along Rio Mato (7°02’N, 65°13’W). Holotype: ANSP 165815 .

Distribution: 308 ( Vari, Harold, 2001 ).

Creagrutus yanatili Harold, Salcedo, 2010

Creagrutus yanatili Harold, Salcedo, 2010:378 , fig. 1. Type locality: Peru, Departamento de Cuzco, Provincia de La Convención, Río Urubamba (Ucayali Dr.), vicinity of Chaullay , 13°00’48”S, 72°38’31”W). MUSM 24082 .

Distribution: 312 (Harold, Salcedo, 2010).

Creagrutus yudja Netto-Ferreira, Moreira, 2018

Creagrutus yudja Netto-Ferreira, Moreira, 2018:251 , fig. 1. Type locality: Brazil, Pará , Altamira , Praia do Levi , Rio Iriri, Rio Xingu basin, 3°50’37”S, 52°44’03”W. Holotype: MNRJ 48662 .

Distribution: 322 (Netto-Ferreira, Moreira, 2018).

Creagrutus zephyrus Vari, Harold, 2001

Creagrutus zephyrus Vari, Harold, 2001:220 , fig. 94. Type locality: Venezuela. Amazonas: Rio Casiquiare , playa and backwater about 2 km downstream from mouth of Rio Pamoni (2°48’N, 65°57’W). Holotype: ANSP 161238 .

Distribution: 307, 314 ( Vari, Harold, 2001 ).