Omphale incognita, Hansson, Christer & Shevtsova, Ekaterina, 2012

Hansson, Christer & Shevtsova, Ekaterina, 2012, Revision of the European species of Omphale Haliday (Hymenoptera, Chalcidoidea, Eulophidae), ZooKeys 232, pp. 1-157 : 51-55

Omphale incognita sp. n. Figures 183-197489514

Material.

Holotype female (BMNH), glued to a card, labelled "SWEDEN: Skåne, Lake Kranke, Lottagården, 55°42'N, 13°29'E, 6.vi.2006, C. Hansson & E. Shevtsova". Paratypes. 68♀ 18♂ FRANCE: 1♀ "St. Loi, 15 km W. Saint-Emiland, 14.viii.2001, J.S. Noyes" (BMNH); 1♀ "Montals, ABPHY (Gard), 11.vii.1989, G. Delvare" (RMNH). GERMANY: 5♀"no specified locality, viii.1965, from Geocrypta galii on Galium pumilum" (RMNH); 1♀ 1♂ “Baden-Württemberg, 21.vii.1972, M.J. Gijswijt" (RMNH); 1♂ "Nordrhein-Westfalen, Brochterbeck, 20.v.1971, M.J. Gijswijt" (RMNH). HUNGARY: 9♀ "Veszprem Co., Nyirad, 47°00'N, 17°27'E, 213 m, 27.vi.2010, C. Hansson & J.S. Noyes" (BMNH, CH); 1♀ "Vas Co., Bárkás Lake, 46°52'N, 16°25'E, 28.vi.2010, C. Hansson" (BMNH). NETHERLANDS: 1♀ "Delfzijl, 11.vi.1997, K. Alders" (RMNH). SWEDEN: 1♀ "Blekinge, Kristianopel, 21.vi.1998, R. Danielsson" (LUZM); 24♀ 4♂ with same label data as holotype (BMNH, CH, LUZM); 1♀ from same locality as holotype but collected 14.vii.2006 (BMNH); 1♀ “Skåne, Lake Kranke, West, 55°42'N, 13°27'E, 6.vi.2006, C. Hansson & E. Shevtsova" (BMNH); 3♀ 3♂ from same locality as previous but collected 8.viii.2006 (CH); 2♂ “Skåne, Silvåkra, 1.viii.1984, C. Hansson" (LUZM); 1♀ from same locality as previous but collected 2.vii.2005 (LUZM); 1♂ ”Skåne, Lake Kranke, Stensoffa, 55°42'N, 13°26'E, 8.viii.2006, C. Hansson & E. Shevtsova" (LUZM); 1♀ “Skåne, Vomb, 18.vii.2006, C. Hansson & E. Shevtsova" (LUZM); 1♂ ”Skåne, Vombsjöns SV strand, 21.vi.1983, C. Hansson" (LUZM); 5♀ 1♂ ”Skåne, Häckeberga swamp, 55°34'N, 13°25'E, 5.vii.2006, C. Hansson & E. Shevtsova" (BMNH, CH); 1♀ ”Skåne, Lund, 17.vii.1983, C. Hansson" (BMNH);1♀ “Skåne, Skäralid, 6-17.viii. 1994, M. Sporrong" (LUZM); 1♂ ”Närke, Örebro, 4.vii.1947, A. Jansson" (LUZM); 1♀ from same locality as previous but 14.vii.1953 (LUZM); 1♀ “Småland, Skillingaryd, 4.vii.1941, A. Jansson" (LUZM); 1♂ “Öland, Kalkstad, 56°37'N, 16°32'E, 20.vi.2007, C. Hansson" (BMNH). UNITED KINGDOM: 6♀ 1♂ "England, Norfolk, Irrimes Graves, 25.vii.1978, J.S. Noyes" (BMNH); 1♀ "England, Lincolnshire, Salmonby, 28.vii.1951, M.W.R. de V. Graham" (BMNH); 1♀ "England, Berkshire, Mapledurham, 14.vi.1975, J.S. Noyes" (BMNH); 1♂ "England, Oxfordshire, Lewknor, 13.vi.1970, M.W.R. de V. Graham" (BMNH).

Diagnosis.

Mesoscutum and scutellum usually bicoloured in golden purple or purple metallic and bluish green metallic (Fig. 184), and thus similar to Omphale phruron ; female coxae bi-coloured (Fig. 183): brown with at least apices pale (yellowish brown or pale brown), femora dark brown with apices yellowish brown; tibiae yellowish brown (occasionally dark brown). Similar to Omphale phruron but female gaster shorter, scape predominantly pale (yellowish brown), coxae and femora with apical parts pale (yellowish brown), tibiae usually completely pale, male flagellomeres 1-4 longer and with several setae apical to basal whorl (Fig. 195). Male genitalia: phallobase (Fig. 489) with volsellar setae on long extensions and with apex of setae 0.1 × the length of setae from apex of phallobase, digitus 1.6 × as long as wide; aedeagus long and slender (Fig. 489), with penis valves 3.6 × as long as wide. Male genitalia are thus very different from male genitalia in O. phruron.

Description.

Female. Length of body 0.9-1.5 mm. Antenna with scape yellowish brown with dorsal margin dark brown, pedicel and flagellum dark brown; pedicel + flagellum 1.8 × as long as distance between eyes; first flagellomere 1.1 × as long and 1.3 × as wide as second flagellomere (Fig. 191); flagellomeres 2-4 ventrally with one set of long setae attached subbasally and reaching beyond apex of flagellomere attached to; clava 1-segmented. Face golden purple (Fig. 186), with weak striae (Fig. 192); clypeus golden with purple metallic tinges, smooth, semicircular, 1.8 × as wide as high; gena golden purplish; lower frons golden purple, with weak reticulation, subtorular and interscrobal areas green metallic and smooth; antennal scrobes join on frontal suture; frontal suture V-shaped; upper frons golden with purple and green metallic tinges, with very weak reticulation; vertex golden purple, with engraved weak reticulation (Fig. 193). Occipital margin rounded (Fig. 193).

Mesoscutum with anterior ½ golden purple and posterior ½ bluish green metallic (Fig. 184), occasionally with entire midlobe golden purple, with engraved reticulation (Fig. 190), midlobe with one pair of setae (posterior pair); notauli as indistinct impressions in posterior ½. Scutellum with anterior ½– ⅔ golden purple and posterior ⅓ –½ bluish green metallic (Fig. 184), to completely golden purple, with engraved reticulation (Fig. 190); 1.2 × as long as wide, with anterior margin almost straight. Axillae golden purple (Fig. 184). Dorsellum golden purple (Fig. 184), concave with weak reticulation (Fig. 190), 0.3 × as long as wide, and 0.4 × as long as length of median propodeum. Entire lateral mesosoma golden with green metallic tinges (Fig. 183); transepimeral sulcus strongly curved forwards. Propodeum golden green with median part purplish (Fig. 184), smooth (Fig. 190); propodeal callus with two setae. Coxae yellowish brown with bases brown to brown with apices yellowish brown (Fig. 183); femora dark brown with apex yellowish brown; tibiae yellowish brown, occasionally dark brown; foretarsus dark brown, mid- and hind tarsi yellowish brown becoming gradually darker towards apex; midleg with first tarsomere 0.3 × as long as length of tarsus. Forewing transparent, veins yellowish brown, setae dark brown (Fig. 188); speculum closed; admarginal setae 7-11, arising from marginal vein; radial cell bare; postmarginal vein 1.0 × as long as stigmal vein, stigmal vein slender. Hind wing tran-sparent, apex pointed (Fig. 188). Forewing WIP (Fig. 189) with apical ½ magenta, basal ½ with wide bands in blue, yellow and magenta.

Petiole dark brown. Gaster with first tergite bluish green metallic, remaining tergites dark brown with golden green tinges, ovate and 1.1 –1.3× as long as length of mesosoma; 7th tergite 0.07 × as long as length of gaster.

Male. Length of body 0.9-1.3 mm. Features as in female except as follows. Antenna dark brown metallic; pedicel + flagellum 2.5 × as long as distance between eyes; flagellomeres 1-4 with verticillate setae and with setae reaching beyond apex of flagellomere attached to, with scattered setae apical to basal whorl (Fig. 195); clava 1-segmented. Face golden red with green metallic tinges, to golden green, strigose; clypeus golden red with green metallic tinges to golden green, smooth, trapezoid to semicircular, 1.6 × as wide as high; gena golden purple; lower frons golden red, golden green, or bluish green metallic (Fig. 187); antennal scrobes join frontal suture separately (Fig. 196); upper frons golden red, golden purple, or bluish green metallic, with weak reticulation; vertex dark brown with purple metallic tinges.

Mesoscutum golden red or golden purple with green metallic tinges (Fig. 185). Scutellum with anterior ⅔ golden purple, posterior ⅓ green metallic (Fig. 185), to completely golden purple or purple metallic; 1.3 × as long as wide. Some Swedish speci- mens with mesoscutum and scutellum greenish blue metallic. Dorsellum golden to golden green (Fig. 185), slightly concave and smooth (Fig. 194), 0.4 × as long as wide, and 0.5 × as long as length of median propodeum. Lateral pronotum golden; prepectus, mesepisternum and mespeimeron dark brown metallic. Legs with coxae and femora dark brown metallic; tibiae pale brown to dark brown; tarsi yellowish brown to pale brown; midleg with first tarsomere 0.3 × as long as length of tarsus. Forewing admarginal setae 6-7, arising from marginal vein; stigmal vein slightly enlarged to slender.

Petiole dark brown. Gaster with first tergite golden green, remaining tergites dark brown with golden tinges, 1.0 × as long as length of mesosoma. Phallobase and aedeagus as in Fig. 489.

Host.

Geocrypta galii ( Diptera : Cecidomyiidae ) on Galium spp.

Distribution.

France, Germany, Hungary, Netherlands, Sweden, United Kingdom (Fig. 514).

Etymology.

From the Latin incognita = unknown.