Paramblypterus rohani ( HECKEL , 1861)

Štamberg, Stanislav, 2013, Knowledge Of The Carboniferous And Permian Actinopterygian Fishes Of The Bohemian Massif - 100 Years After Antonín Frič, Acta Musei Nationalis Pragae Series B 69 (3 - 4), pp. 159-182 : 172-173

Paramblypterus rohani ( HECKEL, 1861)

Pl. 1, fig. 1

1861 Palaeoniscus Rohani HECK. ; Heckel and Kner, p. 51, figs 1-3.

1861 Palaeoniscus luridus HECK. ; Heckel and Kner, p. 54, fig. 4.

1861 Palaeoniscus obliquus HECK. ; Heckel and Kner, p.56, fig. 5.

1861 Palaeoniscus caudatus HECK. ; Heckel and Kner, p. 58, fig. 6.

1894 Amblypterus Rohani (HECKEL).; Fritsch, p. 104; figs 297–301; pl. 123.

1895 Amblypterus caudatus (HECK.) .; Fritsch, p. 111.

1895 Amblypterus luridus (HECKEL) .; Fritsch, p. 109; fig. 302.

1895 Amblypterus obliquus (HECKEL) ; Fritsch, pp. 109, 111; figs 304, 305.

1942 Amblypterus rohoni HECKEL ; Lemke and Weiler, pp. 7-16, figs 2-12.

1967 Janassa lacustris sp. n.; Zídek, p. 203; pl. 1, figs 1, 2.

1969 Paramblypterus rohani HECKEL (1861) ; Heyler, p. 72; fig. 22; pl. 14, figs 1, 2.

1971 Paramblypterus rohani ; HEYLER, figs 1–18, 20–22, 24, pls. 1–4.

1975 Paramblypterus rohani ( HECKEL, 1861) ; Štamberg, p. 305; figs 1, 2a; pls. 1–2.

1976 Paramblypterus rohani ( HECKEL, 1861) ; Štamberg, p. 12; figs 1–17; pls. 1, 2, fig. 1; pls. 3–7.

1976 Paramblypterus caudatus HECKEL, 1861 ; Štamberg, p. 37; figs 22–24; pl. 2, fig. 2; pls. 10–12.

1976 Paramblypterus luridus HECKEL, 1861 ; Štamberg, p. 44; pl. 2, fig. 3.

1976 Paramblypterus obliquus HECKEL, 1861 ; Štamberg, p. 45.

1993 Paramblypterus gelberti (GOLDFUSS, 1847) ; Štamberg, p. 84, figs 1, 2; pl. 1.

2006 Paramblypterus rohani ( HECKEL 1861) ; Štamberg, pp. 225, 227.

2007

2007

2008

2008

2008 Paramblypterus rohani ( HECKEL, 1861) ; Štamberg, pp. 6–8; figs 1–5.

Paramblypterus caudatus ( HECKEL, 1861) ; Štamberg, p. 8, fig. 1.

Paramblypterus rohani ( HECKEL, 1861) ; Štamberg and Zajíc, p. 157, fig. 235.

Paramblypterus caudatus ( HECKEL, 1861) ; Štamberg and Zajíc, p. 157, fig. 236.

Paramblypterus gelberti (HECKEL, 1847) ; Štamberg and Zajíc, p 158, fig. 238.

D i a g n o s i s: Fusiform body gibbous in front of the dorsal fin, not exceeding 35 cm total length. Posterior margin of the scales is serrated, area of serrated scales above and below lateral sensory line and number of dents on their posterior margin decline to the anal fin. Square postrostral, convex anteriorly. Oblong frontals, two times longer than wide, with strongly curved interfrontal suture, and one or two process on lateral side of frontals. Parietals approximately square, two times shorter than the length of the frontals. Triangular dermosphenotic. Distinct sculpture is formed by ridges on the bones of the skull roof. 6 to 7 suborbital bones. Spiracular bone present. Sclerotic ring present. Preoperculum conspicuously bent anteriorly along posterior and dorsal borders of the deep maxillary plate. Scale count

29

41

9 22 37

L e c t o t y p e ( Štamberg 1976): Specimen M 849 designated by Štamberg (1976) from the material figured by Heckel ( Heckel and Kner 1861) is deposited in the National Museum, Prague.

T y p e h o r i z o n a n d l o c a l i t y: Košťálov; Rudník Horizon, Vrchlabí Formation, Asselian, Krkonoše Piedmont Basin.

O c c u r r e n c e: Čistá, Košťálov, Košťálov (“Kovář´s Mill”), Košťálov (“Behind the Tavern”), Kovářsko, Kundratice (“Doly”), Kundratice (“Gorge”), Příkrý (“Honkův creek”), Semily (“Behind the factory”), Rybnice (“Hrádecký creek”), Otovice (“Černý brook”), Veselá (“Veselka brook”), Horní Kalná (“Dump of the Adam mine”).

R e m a r k s: Paramblypterus rohani ( HECKEL, 1861) is the most common actinopterygian fish of the Rudník Horizon (Vrchlabí Formation, Asselian, Early Permian) of the Krkonoše Piedmont Basin. Isolated bones of this species are also occasionally found in the younger Kalná Horizon (Prosečné Formation, Sakmarian, Early Permian) of the Krkonoše Piedmont Basin and in the Otovice Horizon (Broumov Formation, Asselian) of the IntraSudetic Basin. Paramblypterus rohani is also common in the locality Surmoulin (Surmoulin Formation, Early Permian) of the Autun Basin in the French Massif Central ( Heyler 1971).

Paramblypterus rohani was initially described under the name Palaeoniscus rohani together with P. caudatus , P. luridus and P. obliquus in the paper of Heckel and Kner (1861). Fritsch (1894, 1895) assigned all of these species to the genus Amblypterus . He redescribed these species in more detail, provided figures, and recognized a resemblance of the above-mentioned species to Amblypterus duvernoy ( Agassiz, 1833) . However, Fritsch (1894) considered two specimens figured by Agassiz (1833) as Amblypterus duvernoy to be two different species. Agassiz had failed to provide a detailed description of the head bones, scales and fins, and for that reason, Fritsch (1894, 1895) introduced the species Amblypterus rohani , A. obliquus , A. caudatus , A. luridus together with A. vratislaviensis as a variety of A. duvernoy . He redescribed only A. reussii separately. After new studies of numerous comparative material, and the high variation of the interfrontal suture, lateral process on the frontal, and for the high deformation of the body of some specimens, I consider the species A. rohani , A. luridus , A. obliquus and A. caudatus to be only one species P. rohani . Also, the specimen described by Štamberg (1993), and Štamberg and Zajíc (2008) as Paramblypterus gelberti actually belongs to P. rohani . The new taxon described by Zídek (1967) as a tooth of a holocephalian fish Janassa lacustris represents only a cleithrum of P. rohani .

Great similarity was noted several times between P. rohani and P. duvernoy (see Fritsch 1894, Woodward 1891, Štamberg 1976, Dietze 2000). The shape of the body, position of the fins and the anatomy of the head of both species are very close or identical. Lastly, Dietze (1999, 2000) did cast light on the relationship between both species. It is necessary after my last studies to confirm the presence of the sclerotic ring in the orbit of P. rohani , whereas it is missing ( Dietze 2000) on P. duvernoy . The sclerotic ring is a delicate structure and it probably also occurred in P. duvernoy . I expect that both species will be united in the future.