Glyphidops (Glyphidops) steyskali, Sepúlveda, Tatiana A., Wolff, Marta I. & De Carvalho, Claudio J. B., 2014

Sepúlveda, Tatiana A., Wolff, Marta I. & De Carvalho, Claudio J. B., 2014, Revision of the New World genus Glyphidops Enderlein (Diptera: Neriidae), Zootaxa 3785 (2), pp. 139-174 : 154-156

7. Glyphidops (Glyphidops) steyskali , new species

( Figs. 7 A–I, 15C)

Diagnosis. Dark-brown with bluish reflections. Frontal vitta yellow with blackish “Y” shaped stripe from antennal base to ocellar tubercle. Genal seta straight and slender. First flagellomere elliptic with arista apical; length twice or more width. Legs dark brown with apex paler to yellowish. Surstylus spatulate.

Male. Body length 7.2 mm. Wing length 7.2 mm and width 5.9 mm. Blackish-brown with violetbluishreflections; head and thorax partly yellow ( Fig. 7 A).

Head. Slightly elongate in lateral view; dorsally almost as long as wide; 1.5 mm length and 1.3 mm width. Frontal vitta yellow with blackish “Y” shaped stripe from antennal base to ocellar tubercle ( Fig. 7 C); fronto-orbital plate brown, sometimes paler near posterior fronto-orbital seta. Postocellar seta slightly convergent; length three times transverse diameter of ocellar tubercle; inner vertical seta straight and 80% length of postocellar seta; outer vertical seta curved, divergent and same length as inner vertical seta. Genal seta same length as postocellar seta. Postgena relatively wide, discretely narrowing posteriorly. Occiput brown; height, 3.2 times length ( Fig. 7 D).

Antennal base brown. Antennae length 60% length of head. Scape blackish-brown; 0.2 mm long. Pedicel same color as scape; 0.2 mm long; inner process of pedicel elongate, finger-like with pointed apex. First flagellomere lanceolate with slightly rounded apex; 0.6 mm long; length, 2.3 times width. Arista apical, with dense white pubescence.

Thorax. Pleuron with bluish reflections. Anterior notopleural seta hair-like. Posterior notopleural seta, same length as scutellar seta but stronger. Supra-alar seta same length as postalar seta but slender. Dorsocentral seta subequal to scutellar seta. Scutellar median yellow stripe sometimes inconspicuous or absent; scutellar seta straight, 70% length of scutellum. Proepisternum with three anteroventral brown setulae. Katepisternum more densely pruinescent than the rest of pleuron; katepisternal seta half length of posterior notopleural seta. Katatergite length 1.7 times height. Metepimeron yellow.

Legs. Mid coxa with one lateral seta. Femora velvety dark brown, distally paler to yellowish; mid femur anteriorly with one median seta.

Wing ( Fig. 7 E). Basicosta without seta. Halter yellow with knob darkened.

Abdomen. Same color as thorax, dorsally paler. Syntergite 7+8 shiny yellowish-brown. Epandrium brown. Anterior lobe of surstylus subshiny yellowish-brown, spatulate and inflated ( Figs. 7 F–G).

Variation. Body length 5.2–8.2 mm. Wing length, 4.5–7.2 mm and width 1.3–1.8 mm. Head. Fronto-orbital plate paler to yellowish-brown. Occiput height twice length. Thorax. Katatergite yellow in one male from Peru. Legs. Mid femur anteriorly with one to four median setae. Wing. Halter entirely yellow. Abdomen. Anterior lobe of surstylus considerably spatulate and inflated, also variable in color.

Female. Similar to male, differs as follows: body length 6.6–7.6 mm ( Fig. 7 B). Wing length 5.6–6.2 mm and width 1.8– 1.5 mm. First flagellomere length, up to three times width. Katerpisternal seta absent. Oviscape subshiny brown and yellow in one specimen from Colombia; length 2.0–2.5 times maximum width ( Figs. 7 H–I).

Etymology. The specific epithet steyskali is in homage to the dipterist George C. Steyskal, for his contributions to knowledge of acalyptrate. George C. Steyskal had previously segregated specimens of this species as new, from material deposited in the USNM, although he never described it.

Material examined. Type material. Holotype: Glyphidops (Glyphidops) steyskali , new species (♂). Ecuador, Puerto Orellana: Rio Tiputini (0°38.2´S, 76°8.9´W), 12–26.viii.1999, W. Mathis, A. Baptista & M. Kotrba [ USNM] (without right fore leg). Paratypes: 1 ♀, Colombia, Huila, PNN Cueva de los Guacharos, Cedros 1°37´S, 76°6´W, 1950m, Malaise, 20.iv–5.v.2003, C. Cortés [IAvH]; 3 ♂, Meta, PNN Sumapaz, Jardin Botánico, 3°48´N, 73°56´W, 730m, Malaise, 4–24.i.2002, H. Vargas [IAvH]; 1 ♀, Colombia, Caqueta, Sucre nr.km 35, 24.i.1969, Duckworth & Dietz [ USNM]; 1 ♂ 2 ♀, Colombia, Vaupés, Estación Biológica Moshiro-Itajura (Caparú) Centro Ambiental, 1°4´S, 69°31´W, 60m, Malaise, 20.i–1.ii.2003, M. Hharkey & D. Arias [IAvH]; 1 ♂, Amazonas, PNN Amacayacu, Matamata, 3°41´S, 70°15´W, 150m, Malaise, D. Chota [IAvH]; 1 ♂ 1 ♀, ♂, Ecuador, Puerto Orellana: Rio Tiputini (0°38.2´S, 76°8.9´W), 12–26.viii.1999, W. Mathis, A. Baptista & M. Kotrba [ USNM]; 2 ♂ 2 ♀, Ecuador, Zam-Chin. Prov. Cumbaratza, 12.vi.1977, P. J. Spangler & D. R. Givens [ USNM]; 7 ♂ 2 ♀, Ecuador, Puerto Orellana: Rio Tiputini (0°38.2´S, 76°8.9´W), 12-26.viii.1999, W. Mathis, A. Baptista & M. Kotrba [ USNM]; 1 ♂, Peru, Loreto, San Antonio, 14.viii.1965, J. C. Hitchcock Jr. [ USNM]; 1 ♂, Peru, Huanuco, Cochicote, 12.ix.1965, J. C. Hitchcock Jr. [ USNM]; 1 ♂, Peru, Madre de Dios: Manu, Rio Manu, Patkitza (5 km E), Aguajal, 19.ix.1988, A. Freidberg [ USNM]; 1 ♂ 5 ♀, Brazil, Amazonas, São Gabriel Cachoeira, 3–6.iv.2008, B. Ronchi-Teles/ Acampamento da torre da LBA, McPhil, Concentrado Bia [ DZUP]; 2 ♂, Brazil, Amazonas, Manaus, RFA Ducke, VSR fruta, 18–20.ix.2010, Cordeiro, Guedes, Grisales & Haseyama [ DZUP].

Distribution. Colombia, Ecuador, Brazil (Amazonas), Peru ( Fig. 15 C).

Figures

FIGURE 7. Glyphidops (Glyphidops) steyskali, new species; A. Male, habitus; B. Female, habitus; C. Head, dorsal view; D. Head, lateral view; E. Wing; F. Epandrium, dorsal view; G. Epandrium, lateral view; H. Oviscape, dorsal view; I. Oviscape, lateral view. Scales: 1 mm.

FIGURE 15. Distribution maps for Glyphidops. A. G. (G.) obscurus Hennig; B. G. (G.) ruselatus, new species; C. G. (G.) steyskali, new species; D. G. (G.) vittatus (Cresson).