Pardosa fuscula (Thorell, 1875)

Marusik, Yuri M., 2023, A new species related to Pardosa atrata (Araneae, Lycosidae) from Armenia makes the distribution range of the atrata group disjunctive, ZooKeys 1180, pp. 225-235 : 225

Pardosa fuscula (Thorell, 1875)

Figs 3D, G , 4C, D , 5G-I , 6

Lycosa fuscula Thorell, 1875: 501 (♀).

Pardosa fuscula : Dondale and Redner 1987: 2, figs 1, 2, 17-19 (♂♀).

Pardosa fuscula : Dondale and Redner 1990: 171, figs 219-223 (♂♀).

Pardosa fuscula : Paquin and Dupérré 2003: 162, figs 1799-1802 (♂♀).

Pardosa atrata For complete list of taxonomic references, see WSC (2023).

Material examined.

Canada, Yukon Territory: 1♀ (IBPN), Kluane Lake, Cultus Bay, 61°11'N, 138°20'W, Rat Lake , pebbly NW bank, 23.07.1993 (Y.M. Marusik) ; 3♀ (IBPN), Kluane Lake, Christmas Bay , 61°03'N, 138°21'W, 22.07.1993 (Y.M. Marusik) ; 1♀ (IBPN), environs of Carmacks , 62°11'N, 136°22'W, around small lake, willow- Carex vegetation, 17.03.1993 (Y.M. Marusik) ; 1♀ (IBPN), environs of Carmacks , 62°11'N, 136°22'W, oligotrophic bog near small lake, 17.07.1993 (Y.M. Marusik) . USA: 9♂ 9♀ (IBPN), Washington State , Chelan Co., Fish Lake , 588 m, 48°N, 121°W, sphagnum bog, 19.05.1996 (Y.M. Marusik) .

Comments.

Although P. fuscula has been treated in eight taxonomic papers, proper figures showing details of the copulatory organs, including the embolic division, were given only by Dondale and Redner (1987, 1990). It is worth mentioning that Banks (1910) considered P. fuscula to be a junior synonym of P. modica (Blackwall, 1846).

Diagnosis.

The male of this species differs from those of the other Pardosa atrata -group species in having a conical terminal apophysis (Ta) (vs tip of terminal apophysis rounded; cf. Fig. 3E-G ). The female of P. fuscula can be distinguished by the lack of round lateral extensions of the septal base (Bm) (vs present) and also in having relatively longer receptacles (cf. Fig. 5 C, F, I ).

Description.

See Dondale and Redner (1987, 1990).

Type locality.

Strawberry Harbour (55.149°N, 59.023°W), Labrador, Canada.

Distribution.

This species is restricted to the Nearctic (Fig. 6 ) and occurs from westernmost Alaska to easternmost Canada, from the Mackenzie River mouth (ca 69°N) south to northern New Mexico (ca 36°N).

Figures

Figure 3. Embolic division (A-D) and tegular apophysis (E-G) of Pardosa armeniaca sp. nov. (A, B, E), P. atrata (C, F from Polar Ural) and P. fuscula (D, G from Washington State). A, C-G ventral B anterior. Abbreviations: Co - conductor, Em - embolus, Te - terminal apophysis. Scale bars: 0.2 mm.

Figure 4. Epigyne of Pardosa armeniaca sp. nov., P. atrata, and P. fuscula A-C P. armeniaca sp. nov. D-F P. atrata G-I P. fuscula A, B, D, E, G, H ventral C, F, I dorsal D from Utsjoki, Finland E from Yamalo Nenets Okrug, Russia G, H from Washington State. Abbreviations: Bm - rounded margin of septal base, Re - receptacle, Sb - septal base, Ss - septal stem, Sw - septal wing. Scale bars: 0.2 mm.

Figure 5. Embolic division and epigyne in Pardosa atrata and P. fuscula A, B P. atrata from Oulu C, D P. fuscula from Washington State A, C embolic division, ventral B, D epigyne, ventral. Abbreviations: Co - conductor, Em - embolus, Te - terminal apophysis. Scale bars: 0.1 mm.

Figure 6. Distribution map with records of Pardosa armeniaca sp. nov. (red square), P. atrata (blue dot), P. fuscula (grey diamond), the type locality of P. narymica (blue triangle), doubtful records of P. atrata (question mark), and Chinese records of P. atrata that may belong to other species (pink dots).