Bronchocela, Kaup, 1827

Amarasinghe, A. A. Thasun, Kamsi, Mistar, Riyanto, Awal, Putra, Chairunas A., Hallermann, Jakob, Andayani, Noviar, Abinawanto, A. & Supriatna, Jatna, 2022, Taxonomy, distribution, and conservation status of a rare arboreal lizard Bronchocela hayeki (Müller, 1928) (Reptilia: Agamidae) from northern Sumatra Indonesia, Zootaxa 5120 (3), pp. 409-422 : 420-421

Key to the species of the genus Bronchocela (modified after Hallermann, 2005)

1.(a) Nuchal crest well-developed............................................................................ 2

(b) Nuchal crest weakly-developed.......................................................................... 7

2.(a) Nuchal crest scales shorter than ED...................................................................... 3

(b) Nuchal crest scales longer than ED....................................................................... 4

3.(a) Pale labial band absent, gular sac small and midgular scales not enlarged in males, 2–4 upper dorsal scale rows pointing upwards, 8–10 nuchal crest scales to the level of axilla, canthal edges blunt, 3 or 4 canthal scales between supranasal and anterior border of orbit, 50–76 midbody scale rows, tail colouration uniform............................ B. celebensis

(b) Pale labial band present, gular sac large and midgular scales enlarged in males, 4–7 upper dorsal scale rows pointing upwards, 10–12 nuchal crest scales to the level of axilla, canthal edges sharp, 6–11 canthal scales between supranasal and anterior border of orbit, 71–92 midbody scale rows, tail colouration banded.......................................... B. shenlong

4.(a) Mid gular scales enlarged, lateral body scales strongly keeled, 1–7 upper dorsal scale rows pointing upwards............ 5

(b) Mid gular scales not enlarged, lateral body scales feebly keeled, 1–7 upper dorsal scale rows pointing upwards.... B. orlovi

5.(a) Temporal scales keeled, no gular patch in males, 10–12 nuchal crest scales to the level of axilla, gular sac large in males, tympanum diameter 50–60% of ED...................................................................... 6

(b) Temporal scales smooth, red gular patch in males, 8–10 nuchal crest scales to the level of axilla, gular sac small in males, tympanum diameter 75–90% of ED........................................................... B. rubrigularis

6.(a) 33–59 midbody scale rows, ventrals and dorsals equal in size, 10–12 enlarged ventral scale rows, 3 rd finger shorter than 4 th finger, enlarged temporal scales present, 1 or 2 upper dorsal scale rows pointing upwards, tympanum and orbit pale in color, tail colouration banded............................................................................ B. jubata

(b) 64–75 midbody scale rows, ventrals 8–10 times larger than dorsals, 8–10 enlarged ventral scale rows, 3 rd finger longer than 4 th finger, enlarged temporal scales absent, 5–7 upper dorsal scale rows pointing upwards, tympanum and orbit dark in color, tail colouration uniform............................................................................ B. hayeki

7.(a) Three postmentals, nuchal crest scales lanceolate, less than 11 temporal scale rows between orbit and tympanum......... 8

(b) Two postmentals, nuchal crest scales crescent-shaped, 12 or 13 temporal scale rows between orbit and tympanum................................................................................................ B. vietnamensis

8.(a) No white colour patch on temporals, Supraoculars keeled, ventrals larger than dorsals in less than five times............ 9

(b) Prominent white colour patch present on temporals, Supraoculars smooth, ventrals 5–8 times larger than dorsals...................................................................................................... B. danieli

9.(a) Nuchal crest scales shorter than ED, canthal edges sharp..................................................... 10

(b) Nuchal crest scales longer than ED, canthal edges blunt........................................... B. smaragdina

10.(a) Gular sac absent in males, no upper dorsal scale rows pointing upwards......................................... 11

(b) Gular sac present in males, several upper dorsal scale rows pointing upwards.................................... 12

11.(a) 55–67 midbody scale rows.................................................................... B. burmana

(b) 67–71 midbody scale rows.................................................................... B. rayaensis

12.(a) 1 or 2 upper dorsal scale rows pointing upwards, pale labial band present............................. B. marmorata

(b) 4–10 upper dorsal scale rows pointing upwards, pale labial band absent...................... B. cristatella sensu stricto