Chimaerolelaps Desjardins, 2007

Desjardins, Christopher A., 2007, Phylogenetics and classification of the world genera of Diparinae (Hymenoptera: Pteromalidae), Zootaxa 1647 (1), pp. 1-88 : 33-34

Chimaerolelaps Desjardins , New Genus

( Fig. 62 )

Type Species: Chimaerolelaps villosa Desjardins , New Species

Diagnosis: Chimaerolelaps is unique among diparines in 2 ways. First, it has 4 pairs of scutellar bristles, while all other diparines have at most 2 pairs. Second, it has an elongate anellus which is longer than broad ( Fig. 62 ), while all other diparines have an anellus that is broader than long ( Figs. 59, 60 ). Chimaerolelaps may superficially resemble Lelaps or Netomocera . Chimaerolelaps has a clypeal margin which is protruding and straight, while Lelaps has a median clypeal tooth ( Fig. 16 , mct). Chimaerolelaps also has a symmetrical flagellum, while the flagellum in Netomocera is asymmetrical.

Description: Female. Head: Occipital margin rounded; occipital carina present; upper face without strong, transversely carinate sculpture; eyes not posteriorly extended beyond occipital margin; inner eye margins uniformly convex; eyes bare; scrobe present and scrobal channel parallel-sided dorsal to toruli; dorsal margin of scrobe rounded; toruli not on shelf, junction of upper and lower face rounded; antennae symmetrically clavate; antennal formula 11173; anellus longer than broad; pedicel, first funicular segment, second funicular segment subequal in length; claval apex without thick tuft of micropilosity; apical clypeal margin protruding and straight; malar groove present; single pair of strong, dark bristles on vertex present. Mesos- oma: dorsum of mesosoma with strong, dark bristles (1 pair median scutal, 1 pair lateral scutal, 3 pairs anterior scutellar, 1 pair posterior scutellar); pronotum short, collar-like; notauli not arched and meeting posterior scutal margin at lateral edge of scutoscutellar suture; lateral lobes of scutum similar in color to remainder of scutum; posterior scutal margin without setose groove; scutellum large, convex, and sharply sloped posteriorly; axillae convex and not reduced; posterior notal wing process present, pointed; frenal sulcus present; metanotum present as narrow, sculptured band; propodeum at least 1.5X longer than high; propodeum with dorsoventrally flattened spine near anterior margin; plicae absent; suture between postspiracular area and metapleuron diagonal; propodeal foramen circular, open in one plane; longitudinal invagination of pronotum, mesepimeron and metapleuron absent; prepectus elongate, reaching tegula; tegula normal, flap-like; axillary wing sclerite not visible; acropleuron normal, not expanded; mesopleuron with smooth and sculptured regions posteriorly; metacoxa posteriorly convex, with transverse striations; metacoxa posteriorly without thick vertical brush of setae; 2 metatibial spurs, <1.5X width of tibia at point of insertion. Metasoma: Petiole cylindrical, with single pair of setae, at least 2X as long as broad; GT1 expanded, covering at least half of metasoma length; GT1 rounded lateral to petiole insertion; cercal setae elongate; cercal brush present. Male: Unknown.

Discussion: Chimaerolelaps is positioned phylogenetically as sister-group to (or near in some analyses) the Dipara clade, although the only synapomorphy for this clade is the presence of petiolar setae in the female.

Etymology: chimaero-, to represent the odd assemblage of ancestral and derived features present in this taxon, and -lelaps, to ally the genus with Diparinae . The genus is identified in some collections with a manuscript name. As the manuscript name was somewhat misleading with regards to the phylogenetic position of the group, it is not used here.

Number of Species: 1 described species.

Distribution: Costa Rica.

Hosts: Unknown.

Key to Species: none.

Figures

FIGURES 59–64. Lateral habitus views: 59, Neapterolelaps viridescens; 60, N. mitteri; 61, Cerodipara sabensis; 62, Chimaerolelaps villosa; 63, Conophorisca littoriticus; 64, C. grisselli.

FIGURES 11–16. Conophorisca grisselli: 11, mesosoma (lateral view); 12, metasoma (lateral view); 13, cercus. Lelaps sp. A: 14, antenna; 15, clava; 16, clypeus, mct = median clypeal tooth.