Trichotichnus (s. str.) hamulipenis hamulipenis, Kataev & Hongbin & Wrase, 2022

Kataev, Boris M., Hongbin, Liang & Wrase, David W., 2022, New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies, Zootaxa 5159 (3), pp. 301-353 : 341

Trichotichnus (s. str.) hamulipenis hamulipenis ssp. n.

( Figs. 94 , 96 , 98–102 & 106 )

Type material. Holotype. Male , “ CHINA, Yunnan Provin., Gongshan, Dulongjiang Maku village, roadside 27.68533°N, 98.30425°E′ ′, “ 1823 m, 2004.11.2 night Liang Hongbin collector California Academy & IOZ., Chinese. Acad. Sci. ” ( IOZ).

Paratypes. China. YUNNAN: 2 males, 6 females, same data as holotype ( IOZ, CAS) ; 1 male, Gongshan, Dulongjiang, Elidang Village , beach, 28.00287°N 98.32145°E, 1640 m, 3.XI.2004, D. Kavanaugh, D.Z. Dong leg. ( CAS) ; 3 females, Gongshan Co., Dulongjiang, Bapo , along roadside, 27.73902°N 98.34975°E, 1412 m, 3.XI.2004, H.B. Liang leg. ( CAS) ; 1 male, 1 female, same data but 26.X.2004 ( IOZ) ; 4 males, 3 females, Gongshan, Dulongjiang, Miliwang , above Bapo , 27.72383°N 98.361°E, 1956 m, 31.X.2004 night, H.B. Liang leg. ( IOZ, CAS, ZIN, cWR) ; 2 male, 4 females, Gongshan, Cikai Town, 0.5 km of Kongdang , 27.88111°N 98.34063°E, 1500 m, 25.X.2004 night, D. Kavanaugh leg. ( IOZ, ZIN) ; 1 male, 1 female, same data but 7.XI.2004, D. Kavanaugh leg. ( CAS, ZIN) ; 2 males, 4 females, Baoshan Co., Nankang Yakou , 24°49.9'N 98°46.0'E, 2130 m, 4–7.XI.1998, Stop #98-120 A, D.H. Kavanaugh, C.E. Griswold, C.-L. Long, R. Li, & H.-X. He leg. (CASENT 1002500 and 1002504) ( IOZ, CAS) ; 2 males, 1 female [all teneral], Wuliangshan Mt. Range , 24°45'02"N 100°30'24"E, 2270 m, 12.VI.2011, I. Belousov, I. Kabak, A. Korolev leg. (cB&K) .

Description (5 males and 3 females measured). Habitus as in Fig. 94 . On average, body larger (length 9.0– 10.5 mm, mean 10.0 mm), elytra relatively wider and shorter (EL/EW 1.43–1.54, mean 1.47, and EL/PL 2.59–2.80, mean 2.77), pronotum in central portion very finely punctate or almost impunctate, elytral preapical sinuation ( Fig. 96 ) in most specimens slightly shallower and internal sac of aedeagus ( Fig. 100 ) without distinct curved spiny patch in apical portion near apical spine.

Distribution ( Fig. 106 ). The nominotypical subspecies is known from northwestern Yunnan ( China), where it occurs at altitudes of 1400–2130 m.

Figures

FIGURE 106. Trichotichnus hamulipenis sp. n. (nominotypical subspecies), distribution.

FIGURES 94–95. Trichotichnus hamulipenis sp. n., habitus. 94, nominotypical subspecies (above Bapo); 95, T. hamulipenis nepalensis ssp. n. (holotype). Scale bar = 1.0 mm.

FIGURES 96–97. Trichotichnus hamulipenis sp. n., elytral apex. 96, nominotypical subspecies; 97, T. hamulipenis nepalensis ssp. n. Scale bar = 0.5 mm.

FIGURES 98–105. Trichotichnus hamulipenis sp. n.: 98–102, nominotypical subspecies (98–100, holotype; 101 & 102, Cikai); 103–105, T. hamulipenis nepalensis ssp. n. (holotype). 98 & 103, terminal lamella of aedeagus; 99, 100, 104 & 105, median lobe of aedeagus; 101, gonocoxite; 102, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (98, 100, 103 & 105), lateral (99, 101 & 104) and ventral (102) views. a, apical spiny patch. Scale bars: A = 0.5 mm (98, 101–103), B = 1.0 mm (99, 100, 104 & 105).