Anthobium smetanai, Shavrin, 2022

Shavrin, Alexey V., 2022, A revision of Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae Omaliinae: Anthophagini). V. Algidum, morosum and tectum groups, a new species of the fusculum group, and faunistic records, Zootaxa 5104 (3), pp. 301-346 : 333-334

Anthobium smetanai sp. n.

( Figs. 91–94 )

Type material examined: Holotype ♂: ‘ CHINA: W-Sichuan 1999 | Aba Tibet. Aut. Pref., Weizhou Co. | Quinglai Shan, Wolong Tal. 69 km | WSW Dujiangyan , 3900m, 30°54N, | 102°55E, Steine , Blüten | 15. VII., leg. M. Schülke’ <printed>, ‘Sammlung | M. Schülke | Berlin’ <green, printed>, ‘HOLOTYPE | Anthobium | smetanai p.n. | Shavrin A.V. des. 2022’ <red, printed> ( cSch).

Paratypes (5 specimens): 3 ♂♂ [two specimens dissected], 2 ♀♀: same data as the holotype (2 ♂♂, 1 ♀: cSch; 1 ♂, 1 ♀: cSh). All paratypes with additional red printed label: ‘ PARATYPE | Anthobium | smetanai sp. n. | Shavrin A.V. des. 2022’.

Description. Measurements (n=6): HL: 0.40–0.50; HW: 0.66–0.68; AL (holotype): 1.35; OL: 0.15–0.20; PL: 0.50–0.62; PW: 1.04–1.10; ESL: 1.53–1.59; EW: 1.56–1.67; AW: 1.43–1.65; MTbL (holotype): 0.70; MTrL (holotype): 0.51 (MTrL 1–4: 0.29; MTrL 5: 0.22); AedL: 0.75; TL: 3.15–3.90 (holotype).

Habitus as in Fig. 94 . Head, antennomeres 4–11, pronotum and abdomen brown; maxillary palpomeres and elytra yellow-brown; mouthparts, antennomeres 1–3 and legs yellow. Head with dense transverse microsculpture on clypeus, fine and sparse microreticulation in middle, diagonal and isodiametric in middle, basal part of head with or without irregular and very fine meshes; neck with distinct transverse microsculpture; pronotum with or without very fine transverse microreticulation in medioapical third; abdomen with dense transverse and/or isodiametric microsculpture. Head with moderately large and deep pucntation, sparser in middle and denser on infraorbital ridges; neck with fine and sparse punctation; punctation of pronotum about as that on head, sparser and finer in middle and mediobasal third (one paratype without punctures in middle); punctation of elytra about as that on pronotum, finer in middle and denser, coarser and deeper in apical portion; abdominal tergites without visible punctation.

Head 1.3–1.6 times as broad as long, with slightly elevated middle portion and deep, moderately short anteocellar foveae, reaching level of middle length of eye; postocular ridges obtuse. Apical segment of maxillary palpomere more than twice as long as preceding segment. Antennomere 2 distinctly narrower than basal antennomere, 3 about as long as and slightly narrower than 2, 4–5 distinctly shorter and broader than 3, 6–7 slightly broader than 5, 8–9 slightly shorter than 7, 9 slightly broader than 8, apical antennomere about 1.5 times as long as preapical antennomere, from apical third gradually narrowed toward subacute apex.

Pronotum about twice as broad as long, evenly narrowed both anteriad and posteriad or slightly more narrowed posteriad than anteriad; anterior angles not protruded anteriad; middle portion convex, with very indistinct, small, semicircular impression in mediobasal third in some specimens.

Elytra slightly broader than long, more than twice as long as pronotum, reaching apical margin of abdominal tergite IV.

Metatarsus 1.3 times as long as metatibia.

Male. Apical margin of abdominal tergite VIII slightly and sternite VIII widely sinuate. Aedeagus with wide basal bulb, gradually narrowed toward truncate apex; parameres moderately narrow, distinctly exceeding apex of median lobe, with two short apical and two preapical setae; internal sac long, spirally folded in basal portion ( Fig. 91 ). Lateral aspect of aedeagus as in Fig. 92 .

Female. Apical margin of abdominal tergite VIII truncate. Apical margin of abdominal sternite VIII rounded.

Comparative notes. Regarding the general shape of the body, A. smetanai sp. n. is similar to A. vitreum sp. n., from which it dffers by a paler pronotum, longer antennomeres 4–10, and a broader aedeagus.

Distribution. The species is known only from the type locality in Quinglai Shan in Xizang Autonomous Region, China ( Fig. 93 ).

Natural history. The specimens were collected at an elevation of 3900 m a.s.l. They were found under stones and occasionally on flowers.

Etymology. Patronymic, the species is named after the legendary entomologist and my colleague Aleš Smetana (1931-2021) for the pleasure of the collaborative work with him during several years, including the first three parts of the revision of East Palaearctic species of Anthobium .

Figures

FIGURE 93. Distribution of Anthobium in China and Nepal: A. cornatipenis (black circle), A. himalayense (black star), A. infractum (white square), A. morosum (white star), A. multipunctatum (white circle), A. smetanai (black square), A. umbrinum (black triangle), A. vitreum (black diamond).

FIGURES 87–92.Aedeagus of Anthobium himalayense (Figs. 87–88; paratype), A. multipunctatum (Figs. 89–90; holotype) and A. smetanai (Figs. 91–92; paratype): 87, 89, 91—parameral view, 88, 90, 92—lateral view. Scale bar: 0.1 mm.

FIGURES 94–98. Habitus of species of the Anthobium tectum group: 94—A. smetanai (holotype), 95—A. tichomirovae (holotype), 96—A. umbrinum (holotype), 97—A. vitreum (holotype), 98—A. infractum (holotype). Scale bar: 1.0 mm.