Allodia shimai Magnussen

Magnussen, Trude, Soli, Geir E. E. & Kjaerandsen, Jostein, 2019, Allodia Winnertz from the Himalayas, with nine species new to science (Diptera, Mycetophilidae), ZooKeys 820, pp. 119-138 : 125

Allodia shimai Magnussen sp. n. Fig. 2C

Diagnosis.

The dorsal lobe of the gonostylus is heavily sclerotised, dark and with numerous strong setae. The median lobe is distinctly square-shaped with four strong bristles at the edge. Allodia shimai is a dark species.

Type locality.

NEPAL: Province no. 1 (Kosi Zone), Sankhuwasabha District, Thudam, 3500-3800 m a.s.l.

Type specimens.

Holotype: male. 3 printed labels: (E. NEPAL) Thudam (3500-3800m) 27°45'N, 87°31'E --- 27°46'N, 87°33'E / June 27, 1972 H. Shima leg. Kyushu Univ. Col. / TSZD-JKJ-104961. (KUEC, pinned with genitalia in separate microvial). Paratypes: NEPAL: Same data as for holotype except 24 Jun 1972, TSZD-JKJ-104963, male (KUEC); Tanga La, 27°40'N, 087°36'E, 4000-4600 m a.s.l, 6 Jul 1972, Leg. H. Shima, TSZD-JKJ-104962, male (TMU).

Description.

Body length 2.5-4.4 mm; wing length 2.7-3.9 mm. Colouration. Head and clypeus dark brown. Mouthparts and palpomeres dark yellow. Antennae brown, with scape, pedicel and basis of first flagellomere yellow. Scutum dark brown. Lateral sclerites dark brown, except for yellow antepronotum. Wings clear without markings. Halteres whitish yellow. Legs yellow. Abdomen dark brown. Terminalia yellow. Head. Three ocelli present, median ocellus smaller than laterals, lateral ocelli touching eye margin. Head covered with fine trichia, making a silvery appearance, several larger setae in row above eye margin. Antennae approx. twice as long as thorax. Scape with several setae dorsally, pedicel with several setae on distal third. Flagellomeres cylindrical, densely clothed with fine trichia. First flagellomere almost twice as long as scape. Thorax. Antepronotum with two long and two shorter setae. Scutum covered with uniform small, pale setae; strong prealar and postalar setae. Few medioposterior discal bristles present. Scutellum with two strong bristles. Laterotergite with four strong setae. Other lateral sclerites bare. Legs. All tibiae with short setae arranged in rows. Mid tibia with five long anterodorsal and numerous short posterodorsal bristles. Hind tibia with seven anterodorsal and four posterodorsal bristles. Wings. Sc short, ending in R. Length of rm more than double the length of stem of posterior fork. Base of anterior fork opposite base of posterior fork. R1 with setae, R5 without. Male terminalia. Tergite IX medially divided, each part rounded, covered with minute trichia, with one strong apical bristle several small setae. Hypandrial lobe heavily sclerotised and elongated. Gonostylus with dorsal lobe prominent, broad, evenly sloping towards slightly prolonged posteroventral corner; outer surface with numerous setae, denser towards basis. Median lobe of gonostylus short, almost squared, with four strong setae along posterior margin and one smaller seta on ventral edge. Ventral lobe of gonostylus club-shaped, pointing ventrally, with some setae apically. Basal part of gonostylus rounded, with many pronounced, thick, short setae. Internal part of gonostylus with long, caudally projecting process.

Etymology.

Named after Hiroshi Shima, the collector of the type specimens.

Remarks.

The dorsal lobe of the gonostylus is somewhat similar to A. scalprata , but much more prominent and larger. Additionally, in A. shimai the internal part of the gonostylus possess a caudally projecting process, which is lacking in A. scalprata , and the median lobe is prominently sclerotised and unlike that in other Allodia species.