Cerocoma (Cerocoma) schaefferi ( Linnaeus, 1758 )

Turco, Federica & Bologna, Marco A., 2011, Systematic revision of the genus Cerocoma Geoffroy, 1762 (Coleoptera: Meloidae: Cerocomini) 2853, Zootaxa 2853 (1), pp. 1-71 : 24-25

Cerocoma (Cerocoma) schaefferi ( Linnaeus, 1758)

Figs 2S , 3S , 4S , 5F , 6S

Meloe schaefferi Linnaeus, 1758: 420 .

Cerocoma schaefferi, Fabricius, 1781: 331 ; Reitter, 1913: 193.

Cerocoma viridis Fourcroy, 1785: 163 .

Cerocoma schraderi Baudi, 1878a: 1041–1042 (nec Kraatz 1863).

Cerocoma schaefferi var. affinis Baudi, 1878b: 358 .

Cerocoma schaefferi var. obscuripes Reitter, 1885: 14 (after Motscholsky).

Cerocoma schaefferi var. viridula Reitter, 1885: 14 .

Cerocoma schaefferi var. orensis Reitter, 1913: 193 .

Cerocoma (Cerocoma) schaefferi, Kaszab, 1951: 264–265 , 268, 272; Dvořák, 1989: 6, 10; Bologna, 1991: 149.

Cerocoma (Cerocoma) schaefferi orensis, Kaszab, 1951: 264 , 268, 272.

Cerocoma (Cerocoma) schaefferi viridula, Kaszab, 1951: 265 , 268, 272.

Type locality. “ Germania ” ( Linnaeus 1758).

Type specimens. No types of the nominate form have been examined. As explained for C. prochaskana , we accept the unpublished selection of Reitter’s types, probably made by Kaszab, and we designate them lectotype and paralectotypes. The lectotype of C. schaefferi viridula has been studied, as well as lectotype and paralectotype of C. schaefferi orensis .

Lectotype ♂: Rhilo Dagh. (white, handwritten) // Holotypus 1885 / Cerocoma Schäfferi / v. viridula ♂ / Reitter (white with a red frame, printed and handwritten) // LECTOTYPE ♂ / Cerocoma schaefferi / v. viridula Reitter, 1885 / Turco & Bologna des. 2009 (red, printed) ( HNHM). Right last palpomere is missing.

Lectotype ♂: Sierra de Queija / Prov. Orense / A. KRICHELDORFF (white, printed) // Holotypus 1919 / Cerocoma Schäfferi / v. orensis ♂ / Reitter (white with a red frame, printed and handwritten) // LECTOTYPE ♂ / Cerocoma schaefferi / v. orensis Reitter, 1913 / Turco & Bologna des. 2009 (red, printed) ( HNHM). Kaszab wrote on the label “1919” instead of “1913”, the correct date of description.

1 paralectotype ♀: Sierra de Queija / Prov. Orense / A. KRICHELDORFF (white, printed) // Allotypus 1919 / Cerocoma Schäfferi / v. orensis ♀ / Reitter (white with a red frame, printed and handwritten) // PARALECTO- TYPE ♀ / Cerocoma schaefferi / v. orensis Reitter, 1913 / Turco & Bologna des. 2009 (red, printed) ( HNHM). Kaszab wrote on the label “1919” instead of “1913”, the correct date of description. Right metatibia and metatarsomeres are missing.

Description. Male. Body metallic green with sparse and long yellowish setae, denser on head and thorax; legs dark yellow with base of mesofemora and the entire metafemora dark; antennae and mouthparts, including maxillary palpi, yellow.

Head transverse with slightly protruding eyes and frontal calli raised over the head, bearing a tuft of yellowish setae on the external side ( Fig. 5F ). Maxillary palpi wide with palpomeres II–III flattened and curved (particularly the III); IV elongate and slightly bent ( Fig. 3S ). Antennae modified with antennomere I bearing a very narrow and long expansion on the external side and dorsal keel very high and flattened, with a slightly curved dorsal edge; III– VI ventrally expanded; III widely expanded and with a deep incision on its ventral edge; IX swollen and rounded ( Fig. 2S ).

Pronotum slightly transverse. Protibiae slightly expanded dorsally in a short keel ( Fig. 4S ). Protarsomeres slightly dorso-ventrally flattened.

Gonostyli, in lateral view, straight, with apical lobes directed forward; apical lobes, in dorsal view, slightly swollen and parallel. Apex of aedeagus rounded; aedeagal hooks equal in size. Sclerotised hooks of endophallus small and very close to each other, equal in size, both pointing outwards ( Fig. 6S ).

Female. Not distinctly modified. Refer to key for diagnostic characters.

Taxonomy. The four available infraspecific names described by Baudi di Selve (1878b) and Reitter (1885; 1913) were considered in the range of the species variability without subspecific value. They represent forms with dark setation distributed in part of the Iberian Peninsula (f. orensis), or with dark legs and setation, common from S Italy, Balkans and S Russia (f. affinis, f. viridula). Preliminary molecular studies (García-París, unpublished), support these conclusions. The infraspecific form obscuripes was referred by Kaszab (1951) to C. dahli , but we did not confirm this placement because the holotype was not examined.

Distribution. Portugal, Spain, France, S Italy (and possibly extinct in S Tyrol), Belgium, Netherlands, Switzerland, Germany, Latvia, Czech Republic, Slovakia, Austria, Hungary, Slovenia, Croatia, Bosnia-Herzegovina, Serbia, Montenegro, Macedonia, Bulgaria, Greece, Romania, Belarus, Ukraine, S Russia, Turkey, Armenia, Azerbaijan, Syria, Kazakhstan. Possibly extinct in Latvia. Records from Israel-Palestine and Iran need confirmation.

Figures

FIGURE 2. Male right antenna, lateral view: A, C. schreberi; B, C. martae; C, C. ephesica; D, C. prevezaensis; E, C. festiva; F, C. graeca; G, C. malatyensis; H, C. albopilosa; I, C. longiseta; J, C. barthelemyi; K, C. vahli; L, C. scovitzi; M, C. latreillei; N, C. adamovichiana; O, C. confusa; P, C. azurea; Q, C. turcica; R, C. muehlfeldi; S, C. schaefferi; T, C. prochaskana; U, C. dahli; V,C. bernhaueri; W, C. simplicicornis; X, C. gloriosa; Y, C. rapillyi; Z, C. marginiventris; AA, C. macedonica; AB, C. kunzei; AC, C. bodemeyeri. Scale bar = 1 mm.

FIGURE 3. Male right maxillary palp, dorsal view: for lettering refer to caption to Fig. 1. Scale bars = 1 mm (1: C–E, G–J, L– R, Y, AA–AC; 2: B, F, K, S–T, X, Z; 3: A, U–W).

FIGURE 4. Male right protibia, lateral view: for lettering refer to caption to Fig. 1. Scale bar = 1 mm.

FIGURE 5. Male head, dorsal view: A, C. schreberi; B, C. scovitzi; C, C. vahli; D, C. adamovichiana; E, C. kunzei; F, C. schaefferi. Scale bar = 1 mm.

FIGURE 6. Male external genitalia (from left to right: tegmen, dorsal and lateral views; aedeagus, lateral view): for lettering refer to caption to Fig. 1. Scale bars = 1 mm (1: A–E, H, L–M, R; 2: F–G, I–K, N–P, S–T, X–Y, AA–AC; 3: Q, U–W, Z).