Verbascum tibesticum ( Quezel ) Hub.-Mor., in Bauhinia 5(1): 15. 1973.

Khamar, Hamid, Oualidi, Jalal El, Touhami, Amina Ouazzani & Civeyrel, Laure, 2023, Nomenclature and typification in Verbascum (Scrophulariaceae) from North Africa, PhytoKeys 225, pp. 115-152 : 115

Verbascum tibesticum ( Quezel) Hub.-Mor., in Bauhinia 5(1): 15. 1973.

Celsia tibestica ≡ Celsia tibestica Quézel, in Bull. Soc. Hist. Nat. Afrique N. 48: 95. 1957. Type: [Tchad]. Emi Koussi, Koudou. 2000 m, September-Novembre 1956, P. Quézel, s n. (Lectotype, designated here: AIX [AIX000033]!; isolectotype: AIX [AIX000034]!). [image of lectotype available at https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.aix000033]

Notes.

The type material of this species was collected during Quézel’s expedition to Borkou and to Tibesti in September-November 1956. In the protologue, Quézel (1957) did not designate a holotype nor did he mention the name of the herbarium where the original material was housed. We have traced three specimens at the herbarium of Muséum d’Histoire Naturelle d’Aix-en-Provence (AIX), France. The sheet AIX000033 contains a well conserved plant with mature fruit and rosette and an original label handwritten by P. Quèzel that reads "Mission Botanique de l’Institut de Recherches Sahariennes de l’Université d’Alger au Borkou et au Tibesti / Celsia tibestica nov; sp. / Emi Koussi, Koudou. 2000 m / Sept-November 1956, Dr. P. Quézel”. The sheet AIX000034 bears only a rosette and an original label handwritten by P. Quèzel reading "Mission Botanique de l’Institut de Recherches Sahariennes de l’Université d’Alger au Borkou et au Tibesti / Celsia tibestica Qz. / Koudou / Sept-November 1956, Dr. P. Quézel”. Both specimens are in accordance with the protologue, and based on Art. 9.6 of the ICN ( Turland et al. 2018) both must be treated as syntypes. The sheet bearing the barcode AIX000033 is designated here as the lectotype, because it is in better condition than the other specimen and shows all relevant characters mentioned in the protologue.