Spinaxytes tortioverpa Srisonchai, Enghoff & Panha

Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai & Panha, Somsak, 2018, A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae), ZooKeys 797, pp. 19-69 : 19

publication ID

https://dx.doi.org/10.3897/zookeys.797.29510

publication LSID

lsid:zoobank.org:pub:A5403588-F43D-40FF-81EE-2009E49823B4

persistent identifier

https://treatment.plazi.org/id/97E5F638-2A4D-4834-B93C-838548BAD087

taxon LSID

lsid:zoobank.org:act:97E5F638-2A4D-4834-B93C-838548BAD087

treatment provided by

ZooKeys by Pensoft

scientific name

Spinaxytes tortioverpa Srisonchai, Enghoff & Panha
status

sp. n.

Spinaxytes tortioverpa Srisonchai, Enghoff & Panha View in CoL sp. n. Figs 1; 4R, S; 5H; 26

Material examined.

Holotype. ♂, MALAYSIA, Perak State, Ipoh City, Gua Tempurung, limestone mountain, 4°24'58"N, 101°11'16"E, ca. 92 m a.s.l., 27 Sep. 2007, B. W. Ng and ASRU members leg. (CUMZ-pxDGT00219). Paratypes. 1 ♂, 1 ♂ gonopods missing, 1 ♀, same data as for holotype (CUMZ-pxDGT00220).

Etymology.

The name is a noun in apposition, from the Latin tortio meaning torsion and verpa for penis, refers to the distal part of gonopod (postfemoral part, solenophore and solenomere) which makes a 90 degrees torsion relative to the femoral part.

Diagnosis.

Male femora without modification; collum with 4+4 tubercles in anterior row, 1+1 tubercles in intermediate row and 2+2 tubercles in posterior row. Similar in these respects to S. hasta sp. n. and S. palmata sp. n., but differs from them by having: a completely bilobed sternal lobe between male coxae 4; postfemoral part, solenophore and solenomere angled 90 degrees with femoral part; lamina lateralis with two lobes - the lateral one spine-like, the mesal one shorter and ridge-like; solenomere very long, longer than lamina medialis.

Description.

SIZE. Length 28-30 mm (male), 30-33 mm (female); width of midbody metazona 2.0 mm (male), 2.8 mm (female). Width of rings 2 = 3 = 4 <head = 5-16, thereafter body gradually tapering towards telson.

Colour. Specimens in life with body black/brownish black. Colour in alcohol: after 10 years changed to brown; paraterga brownish white; antennae brown (except whitish distal part of antennomeres 7 and 8); head, collum, metaterga and prozona blackish brown; surface below paraterga brown/blackish brown; sterna, epiproct and legs pale brown; a few basal podomeres whitish brown.

Antennae. Reaching to body ring 8 or 9 (male) and 6 or 7 (female) when stretched dorsally.

Collum. With three transverse rows of setiferous tubercles, 4+4 tubercles in anterior row, 1+1 tubercles in intermediate row and 2+2 tubercles in posterior row; anterior margin truncate; with one inconspicuous setiferous notch at lateral margin; paraterga wing-like, long and broad, tip sharp, elevated at ca. 15°-20° in both male and female, directed caudolaterad.

Tegument. Quite dull; collum, metaterga and surface below paraterga finely microgranulate.

Metaterga. With two transverse rows of setiferous tubercles/cones and spines; metaterga 2-8 with 2+2 cones in anterior row and 2+2 spines in posterior row; metaterga 9-19 with 2+2 tubercles/cones in anterior row and 2+2 spines in posterior row; mesal spines of posterior row bigger and longer than lateral ones, gradually reduced in length and size on posterior rings.

Paraterga. Very long; directed dorsolaterad on body rings 2-17, elevated at ca. 65°-70° (male) 60°-70° (female), directed increasingly caudad on body rings 18 and 19. Ozopore visible in dorsolateral view.

Telson. Epiproct quite short; tip subtruncate; lateral setiferous tubercles inconspicuous; apical tubercles inconspicuous. Hypoproct subsemicircular; caudal margin round, with inconspicuous setiferous tubercles.

Sterna (Figure 4R). Sternal lobe between male coxae 4 completely divided into two lobes, long, spine-like; tips in situ directed ventrad; posterior surface a bit swollen, bearing 2 pores.

Legs (Figure 4S). Male femora without modification.

Gonopods (Figs 5H, 26). Coxa subequal in length to femur. Prefemoral part about 2/3 as long as femur. Femur quite enlarged distally. Postfemoral part large, broad and wide; angled 90 degrees with femur. Mesal sulcus and lateral sulcus wide. Solenophore smaller than postfemoral part: lamina lateralis apparently with two lobes demarcated from each other; lateral lobe very long, process-like, its tip in situ directed ventrad; mesal lobe short and wide, supporting solenomere: lamina medialis long, base not enlarged, tip directed mesad. Solenomere obviously longer than lamina medialis, circular in transverse section, curving down, tip directed laterad.

Distribution and habitat.

Known only from the type locality which is currently a tourist attraction (cave). We regard this species as endemic to Malaysia.

Remarks.

A photograph of a live specimen was not taken during the field survey, but our collector noticed its black or brownish black colour. All specimens were seen crawling on rock walls where they seem to blend perfectly with the substrate. No variation in morphological characters was found.

Kingdom

Animalia

Phylum

Arthropoda

Class

Diplopoda

Order

Polydesmida

Family

Paradoxosomatidae

SubFamily

Paradoxosomatinae

Tribe

Orthomorphini

Genus

Spinaxytes