Erythrodiplax Brauer, 1868

Mauffray, William F. & Tennessen, Kenneth J., 2019, A Catalogue and Historical Study of the Odonata of Ecuador, Zootaxa 4628 (1), pp. 1-265 : 162-168

Erythrodiplax Brauer, 1868

This is the most species-rich genus (58 described species) of Anisoptera in the Neotropics. The basic reference is Borror (1942); see also Borror (1957) and Paulson (2003). At least 21 species are recorded in Ecuador and more may occur here. Adults are small to medium sized and variously colored in red, brown or black ( Figs. 95a,b,c,d ). As expected, the habitats occupied are diverse, including ponds, lakes, sloughs, marshes, brackish water, phytotelmata, and pools in slow streams. A synoptic update of the genus is warranted.

Erythrodiplax abjecta ( Rambur, 1842) . Publ. Recs. ECU: *Karsch (1891b—as E. ponderosa , description, type locality “ Ecuador ”), Ris (1911a), Campos (1922 “por Dr. Calvert”), Racenis (1954), Bridges (1993), (H06), Paulson (2003), (P11), Bota-Sierra (2014a), (H14); ES: ( Ris, 1911a); EO: Campos (1927 —as E. connata form?); GY: * Ris (1911a); PC: Ris (1911a), Paulson (2003). Also CEN, COL, VEN: (P11).

Addl. Data. CT, Sigchos: 2 ♂♂, Otonga, Esmeraldas and “Z” rivers area, in Naranjito , 1 v 1999, leg. E. Tapia, det. JJD [JJD]; 5 ♂♂, San Francisco de las Pampas, principal road near Esmeraldas River on Naranjito (-0.42, -79.0, 1450m), 1 iv 1999, leg. E. Tapia, det.? [QZAC]; 1 ♂, San Francisco de las Pampas Road between Naranjito-San Francisco de Las Pampas (-0.42, -79.0, 1500m), 14 v 1999; leg. E. Tapia, det.? [QZAC]; 3 ♀♀, San Francisco de las Pampas Otonga Reserve on B2 River above the grass (-0.42, -79.0, 1650m), 28 iv 1999, leg. L. Torres, det.? [QZAC]; 1 ♂, 4 ♀♀, San Francisco de las Pampas Otonga Reserve, Cerca del Río P-1 (-0.42, -79.0, 2200m), 11 iii 1999, leg. E. Tapia, det.? [QZAC]; GY, Guayaquil: 1 ♂, Guayaquil (-2.20, -79.90, 8m), date?, leg. & det. MJW [FSCA]; Naranjal: 1 ♀, [ Venecia ], listed as “Verecia”, (-2.17, -79.52, 23m), date?, leg. F. Campos, det. P. Calvert [FSCA]; 1 ♂, 4 ♀♀, [ San Rafael ], listed as “San Raphael” (-2.18, -79.23, 200m), date?, leg. F. Campos, det. P. Calvert [FSCA]; Yaguachi: 1 ♀, Casiguana (-2.13, -79.78, 4m), date?, leg. F. Campos, det. P. Calvert [FSCA]; IM: Cotacachi: 3 ♂♂, Santa Rosa, El Refugio de Intag Cloud Forest Lodge , south along road at large open seep (0.3773, -78.4774, 1907m), 23 viii 2018, leg. & det. WFM [FSCA, USFQ], same data but 2 ♂♂, south along road at roadside seeps (0.3693, -78.4728, 1854m), 23 viii 2018, leg & det. WFM [FSCA, USFQ]; 1 ♂, Apuela, Ruta Plaza Gutierrez, seepage into Rio Tuabunchi (0.3575, -78.4993, 1634m), 26 viii 2018, leg. & det. WFM [USFQ]; same data but 1 ♂, leg. & det. KJT [KJT]; same data but, 1 ♂, 27 viii 2018, leg. & det. KJT [KJT]; 2 ♂♂, Selve Alegre, Ruta Tolla Intag, roadside pool (0.2800, -78.5160, 1751m), 26 viii 2018, leg & det. WFM [FSCA]; same data but 1 ♂, Santa Rosa, Ruta Siempre Verde, seeps and bog (0.3672, -78.4378, 2226m), 28 viii 2018, leg. & det. WFM [FSCA]; same data but 2 ♂♂, 2 ♀♀, pool and seeps (0.3676, -78.4306, 2322m), 28 viii 2018, leg. & det. KJT [KJT]; PC, Mejía: 1 ♂, Cornejo Astorga, 9.6 km E, ditch (2012m), 16 vii 1977, leg. & det. DRP [DRP]; 3 ♂♂, Santo Domingo de los Colorados, 32 km E, waterfall & trickle, 10 x 1988, leg. & det. SWD [DRP, SWD]; 16 ♂♂, 10 km E of Tandapi, 29 km E of Tinalandia (-0.45, -78.75, 1920m), 24 x 1980, leg. & det. MJW [JJD]; Quito: 1 ♂, Maquipucuna Reserve, near Nanegalito, slow stream (1250m), 28 v 1994, leg. A. Jaramillo, det. DRP [DRP]; San Miguel Bancos: 6 ♂♂, 2 ♀♀, between Calacali and Nanegalito, about 30 km W Quito, 9 vii 1990, leg. OSF, det.? [USNM]; 2 ♂♂, Mindo, Mindo Gardens Ecolodge, vegetated cement pond (-0.0720, -78.7560, 1330m), leg. & det. WFM [FSCA, USFQ]; same data but 1 ♂, leg. & det. KJT [KJT].

* Erythrodiplax amazonica Sjöstedt, 1918 . Publ. Recs. BRA, GUF, GUY, PER, TTO, VEN: ( H06 ), (P11); plus SUR: ( P16 ).

Addl. Data. OR, Orellana: 1 ♂, T. B.S., Quebrada Harpía (-0.6351, -76.1463, 221m), 11 vi 2013, leg. & det. ACRP (pers. comm., ACRP); SU, Shushufindi : 1 ♂, 3 km SW Shushufindi, stream, (-0.2136, -76.6786, 260m), 12 v 2009, leg. & det. KJT [FSCA]; 1 ♂, 1 ♀, stream, 3 km NW of Shushufindi (-0.1664, -76.6917, 290m), 13 x 2009, leg. & det. JTJ [JTJ].

Erythrodiplax andagoya Borror, 1942 . Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14); ES: Daigle et al. (1997); SD: * Borror (1942 —as “ Pichincha ”), Daigle et al. (1997 —as “ Pichincha ”). Also COL: (H06, P11, P&P); plus CEN: (P11).

Addl. Data. CT, Pangua: [JTJ]; ES, San Lorenzo: [FSCA, JJD, KJT, LSAM]; GY, Balao: [JTJ]; Guayaquil: [ANSP]; Naranjal: [FSCA]; LR, Buena Fé: [FSCA, SWD, USNM]; Quevedo: [ANSP]; MN, El Carmen: [FSCA, JJD, JTJ, KJT]; PA, Pastaza: [USNM]; PC, Quito: [FSCA, KJT, USFQ]; San Miguel Bancos: [RWG]; SD: [FSCA, JJD, KJT, LSAM, RWG, QZAC, SWD]; SU, Lago Agrio: [USNM].

Note: This often abundent species occurs mainly on the lower Pacific slope. The east slope records from the USNM (Pastaza) need to be checked as they are probably a different species.

Erythrodiplax attenuata ( Kirby, 1889) . Publ. Recs. ECU: * Borror (1942), (H06), (P11), (H14); NA: Tennessen (1996b). Also BOL, BRA, COL, PER, VEN: (H06), (P11). Plus SUR: (H06).

Addl. Data. OR, Francisco de Orellana: 1 ♂, trib. of Río Payamino, road to Loreto, 1.7 km W of jct. with Coca Road (-0.4367, -77.0017, 258m), 13 vi 1995, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. WFM [FSCA]; La Joya de los Sachas: 1 ♂, Pompeya near Puerto Pompeya Ferry Landing (-0.4400, -76.6194, 240m), 17 vii 1996, leg. & det. WFM [FSCA].

Erythrodiplax basalis ( Kirby, 1897) . Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14), Dalzochio et al. (2018); NA: * Borror (1942); OR: von Ellenrieder & Garrison (2011); PA: * Borror (1942). Also BRA, COL, GUF, GUY, PER, VEN: (H06), (P11); plus ARG BOL, PRY, URY, SUR, TTO: Paulson (2011).

Addl. Data. ES, Esmeraldas: [QZAC]; Quininde: Q); LR, Buena Fé: [USNM]; MN, El Carmen: [QZAC]; MS, Huamboya: [FSCA, QZAC]; Santiago: [SWD]; Sucua: [FSCA]; NA, Archidona: [QZAC]; Carlos Julio Arsemena Tol: [FSCA, QZAC, USNM]; El Chaco: [FSCA, UTIC, KJT, LSAM, SWD]; Tena: [DRP, FSCA, UTIC, KJT, QZAC, RWG, TWD, SWD, USNM]; OR, Francisco de Orellana: [DRP, FSCA, KJT, LSAM, QZAC, RWG, TWD, SWD, USNM]; Loreto: [USNM]; PA, Pastaza: [JTJ, KJT, USNM]; Santa Clara: [FSCA]); PC, San Miguel Bancos: [DRP]; SD: [USNM]; SU, Cascales: [FSCA, JJD, KJT, QZAC, USNM]; Lago Agrio: [FSCA, UTIC, LSAM, QZAC, RWG, SWD, USNM]; Shushufindi: [CSCA, DRP, FSCA, JTJ, KJT, LSAM, QZAC, SWD, USNM]; ZC, El Pangui: [KJT].

Erythrodiplax castanea ( Burmeister, 1839) . Publ., Recs. ECU: Fraser (1946b), Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14); ES: * Ris (1911a), Campos (1922 —“from Ris in Selys collection”), Daigle et al. (1997); NA: Borror (1942); OR: von Ellenrieder & Garrison (2011); PA: Borror (1942); ZC: Tennessen & Johnson (2008). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, TTO, VEN: (H06), (P11); plus SUR: (P16).

Addl Data. EO, Piñas: [ACRP]; Santa Rosa: [JTJ]; ES, San Lorenzo: [FSCA, JJD, KJT, LSAM, QZAC, USNM]; MS, Huamboya: [FSCA, QZAC, USNM]; OR, Francisco de Orellana: [FSCA, JJD, KJT, LSAM, NVE, QZAC,); Loreto: [FSCA, KJT, SWD]; NA, Archidona: (MAR, USNM]; Tena: [DRP, KJT, LSAM, QZAC, SWD, USFQ, USNM]; SU, Cascales: [SWD]; Lago Agrio: [SWD]; Shushufindi: [FSCA, KJT, JTJ]; ZC, Zamora: [UTIC, JTJ, KJT].

* Erythrodiplax cleopatra Ris, 1911 . Publ. Recs. CHL, PER: (H06), (P11).

Addl. Data. EO, Santa Rosa: 1 ♂, stream and associated pond, wetland about 12 km E of Arenillas, Hwy. 25 (-3.5244, -79.9831, 30m), 29 iii 2008, leg. & det. JTJ, checked by DRP [JTJ].).

Note: This is a considerable range extension to the north from previous records.

Erythrodiplax fervida (Erichson in Schomburgk, 1848) . Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14); CR: * Campos (1922 —as E. ochracea ); CB: * Campos (1922 —as E. ochracea ); GY: * Campos (1922 —as E. ochracea ). Also CEN, COL, GUF, GUY, SUR, VEN: (H06), (P11); plus TTO: (P11).

Addl. Data. EO, Piñas: 1 ♂, Buenaventura, Camino principal (-3.6520, -79.7922, 323m), 28 vi 2016, leg. & det. ACRP [ACRP]; Santa Rosa: 1 ♂, 1 cop. pair, pond, 12 km E of Arenillas, Hwy. 25 (-3.5244, -79.9831, 30m), 29 iii 2008, leg. & det. KJT [KJT]; 2 ♂♂, same data but 9 iv 2008, leg. & det. JTJ [JTJ]; MN, Chone: 1 ♂, Chone, W end of town, 9 i 1978, leg. P. Spangler & J. Anderson, det.? [USNM].

Note: According to Borror (1942), all previous references to E. ochracea ( Burmeister, 1839) in northern South America should be referred to E. fervida ; E. ochracea is a southern South American species.

Erythrodiplax funerea ( Hagen, 1861) . Publ. Recs. ECU: * Ris (1911a), Campos (1922 —“por Dr. Ris”), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997b), (H06), (P11), (H14), Dalzochio et al. (2018). Also CEN, COL: (H06), (P11).

Addl. Data. CB, Alausí: 2 ♂♂, Huigra, Río Huagra, 1 km upstream from trail (-2.2783, -78.9731, 1350m), 13 ii 1911, leg. S. Rhoads, det.? [ANSP, FSCA]; ES, San Lorenzo: 3 ♂♂, 1 ♀, Carondelet, date?, leg. F. Campos, det. MJW [ANSP]; 4 ♂♂, 2 ♀♀, Concepcion (1.05, -78.83, 25m), leg. F. Campos, det. MJW [ANSP]; GY, Guayaquil: 1 ♂, Puna Vieja (-2.97, -80.18, 30m), leg. F. Campos, det. MJW [ANSP].

Note: These are the southernmost records for this common Central American species.

Erythrodiplax fusca ( Rambur, 1842) . Recs. ECU: * Calvert (1906 —as E. connata , “form d ”), Campos (1922 —as E. connata fusca ), Fraser (1946b—as E. connata fusca ), Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14); BO: Borror (1942); CB: Ris (1911a), Navás (1934); ES: Daigle et al. (1997), Paulson (2003); LR: Borror (1942); NA: Borror (1942), Paulson (2003); OR: von Ellenrieder & Garrison (2011); PC: Paulson (2003); SD: Daigle et al. (1997 —as “ Pichincha ”), Dalzochio et al. (2018). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, URY, VEN: (H06), (P11); plus TTO: (P11).

Addl. Data. BO, San Miguel: [USNM]; CN, La Troncal: [KJT, SWD]; CB, Alausí: [FSCA]; Pallatanga: [ANSP]; CT, Pangua: [JTJ]; Sigchos: [QZAC]; EO, Piñas: [FSCA]; Santa Rosa: [JTJ]; ES, Esmeraldas: [DRP]; San Lorenzo: [ANSP, FSCA, KJT, USNM]; GY, Duran: [ANSP, FSCA]; Naranjal: [FSCA]; Yaguachi: [ANSP, FSCA, USNM]; Imbabura, Ibarra: [FSCA, KJT, QZAC, USNM]; LJ, Catamayo: [JTJ]; Loja: [FSCA, UTIC]; Macará: [ANSP, FSCA]; LR, Buena Fé: [DRP, SWD, USNM]; Quevedo: [ANSP, FSCA, USNM]; Ventanas: [JTJ]; MN, El Carmen: [QZAC]; Jipijapa: [FSCA]; Portoviejo: [JTJ]; MS, Gualaquiza: [ANSP, FSCA]; Santiago: [FSCA, KJT, SWD]; NA, Archidona: [FSCA]; El Chaco: [UTIC]; Tena: [ANSP, DRP, FSCA, KJT, LSAM, SWD, USNM]; OR, Francisco de Orellana: [FSCA, UTIC, KJT, QZAC, USFQ, USNM]; Loreto: (MAR); PA, Santa Clara: [KJT]; Pastaza: [FSCA]; PC, Mejía: [SWD]; San Miguel Bancos: [DRP, FSCA, KJT, NVE, RWG, USNM, USFQ]; Quito: [DRP]; SD: [CSCA, DRP, FSCA, UTIC, JJD, KJT, QZAC, SWD, USNM]; SU, Cascales: [KJT, SWD, USNM]; Lago Agrio: [FSCA, UTIC, QZAC, SWD, USNM]; Shushufindi: [DRP, FSCA, SWD, USNM]; ZC, Yantzaza: [KJT, USNM]; Zamora: [FSCA, UTIC, JTJ, KJT, LSAM, QZAC, USNM].

Note: All former records of E. connata ( Burmeister, 1839) and E. connata connata from Ecuador are of E. fusca according to Paulson (2003). Also the various references to forms “a–d” of connata could be either E. fusca or even E. abjecta . Since these specimens were not available to study nor contribute additional provincial or cantonal records to either species, their data are not included in this study. The abdomen of west slope specimens is not pruinose and specimens are often referred to as “red form” whereas those from the eastern slopes are pruinose.

Erythrodiplax ines Ris, 1911 . Publ. Recs. ECU: Ris (1916 —as E. connata var ines ), Schmidt (1942), Fraser (1946b), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997b), (H06), (P11), (H14); LJ: Tennessen & Johnson (2008); NA: Borror (1942), Tennessen (1996b), Daigle (1996); PA: Borror (1942); SD: *Ris (1911a—Holotype & Allotype: [SD:] “Santa Inez”[ZMUH]), Ris (1916, 1918), Campos (1925), Borror (1942), Bridges (1993), Tennessen (2006). Also BOL, COL, PER: (H06), (P11).

Addl. Data. CB, Alausí: [ANSP, FSCA]; Cumandá: [FSCA]; Pallatanga: [DRP, FSCA]; LJ, Catamayo: [JTJ]; Loja: [FSCA, JTJ, KJT]; MS, Huamboya: [FSCA]; Limón Indanza: [FSCA, KJT, QZAC]; Morona: [FSCA, KJT, QZAC, USNM]; NA, Archidona: [FSCA, KJT, QZAC, LSAM, RWG, SWD, TWD, USNM]; El Chaco: [FSCA, KJT, SWD]; Quijos: [FSCA, JJD, KJT, QZAC, TWD, USNM]; OR, Loreto: [FSCA, JTJ, MAR); SU, Gonzalo Pizarro: [DRP, QZAC, SWD]; PA, Mera: [SWD, USNM]; TU, Baños: [FSCA, JTJ, RWG, QZAC, USNM]; ZC, Yantzaza: [USNM]; Zamora: [FSCA, UTIC, USNM].

Note: According to Heckman (2006), usually found at elevations about 1000m; this statement seems to hold true; in addition, this species occurs on both sides of the Andes.

Erythrodiplax kimminsi Borror, 1942 . Publ. Recs. ECU: Fraser (1946b), Paulson (1983), Tsuda (1986, 1991), (H06), Bridges (1993), (P11), (H14); GY: * Campos (1922 —as E. erichsoni ); CB: * Campos (1922 —as E. erichsoni ); LR: Borror (1942 —description); OR: von Ellenrieder & Garrison (2011). Also BRA, CEN, COL, PER, VEN: (H06), (P11).

Addl. Data. EO: Santa Rosa: [JTJ, KJT]; GY, Balao: [JTJ]; LR, Buena Fé: [DRP, FSCA, SWD, USNM]; Quevedo: [ANSP, FSCA]; NA, Tena: [DRP, FSCA, JJD, KJT, SWD, TWD, USNM]; OR, Francisco de Orellana: [FSCA, KJT, LSAM, QZAC, RWG, SWD, TWD, USNM]; Joya de los Sachas: [FSCA]; Loreto: [JTJ, KJT]; PA, Santa Clara: [RWG]; SD: [USNM]; SU, Lago Agrio: [FSCA, KJT, QZAC, SWD, USNM]; Shushufindi: [DRP, FSCA, SWD].

* Erythrodiplax lativittata Borror, 1942 Publ. Recs. ARG, BRA, COL, PER, VEN: (H06), (P11).

Addl. Data. SU, Shushufindi: 1 ♀, Limoncocha, Río Napo (-0.4000, -76.6167, 269 m), 2 viii 1974, leg. Boyce A Drummond III, det. WFM [FSCA].

* Erythrodiplax laurentia Borror, 1942 . Publ. Recs. BRA, GUF, GUY, SUR, VEN: (H06), (P11).

Addl. Data. SU, Cascales : labeled “ Napo Prov. ”, 31 km W Lago Agrío and 4 km S, 20 ix 1975, leg. A. Langley & J. Cohen, det. RWG with note “in geographic range of E. kimminsi , but morphology of laurentia ” [USNM].

Erythrodiplax melanica Borror, 1942 . Publ. Recs. ECU: (H14). Also BRA, PER: (P16).

Addl. Data. OR, Francisco de Orellana: Shiripuno Lodge , 27 km SE Río Shiripuno bridge, (-1.1047, -76.7317, 235m), date not provided, WAH; SU: Cuyabeno Reserve, Sendero Palma Roja , 1.4 km SE of Cuyabeno Lodge (- 0.0151, -76.1698), date not provided, data provided by WAH.

Erythrodiplax melanorubra Borror, 1942 . Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14), Dalzochio et al. (2018); BO: * Borror (1942); MS: * Borror (1942 —as “ Azuay ”), Paulson (2003); NA: * Borror (1942), Tennessen (1996b), Paulson (2003); PA: * Borror (1942); SD: * Borror (1942 —as “ Pichincha ”); TU: * Borror (1942). Also ARG, BOL, BRA, PRY, PER, VEN: (H06), (P11); plus COL: (H06); and CHL, GUF: (P11).

Addl. Data. CT, Sigchos: [FSCA, QZAC]; MS, Huamboya: [FSCA]; Limón Indanza: [FSCA, KJT]; Morona: [FSCA, KJT]; Santiago: [KJT]; Taisha: [DRP]; NA, Archidona: [FSCA, KJT, TWD, USNM]; Carlos Julio Arosemena Tola: [FSCA]; Tena: [DRP, KJT, QZAC, SWD, TWD, USNM]; OR, Loreto: [JJD, KJT, LSAM, QZAC, SWD, TWD, USNM]; PA, Mera: [FSCA, KJT, USNM]; Pastaza: [USNM]; Santa Clara: [FSCA, USNM]; PC, Quito: [QZAC]; San Miguel Bancos: [QZAC]; SD: [QZAC, SWD]; SU, Cascales: [USNM]; Gonzalo Pizarro: [QZAC, USNM]; Lago Agrio: [USNM]; Shushufindi: [KJT]; TU, Baños: [FSCA, JTJ, SWD]; ZC, Yantzaza: [FSCA]; Zamora: [FSCA, KJT].

Erythrodiplax paraguayensis (Förster, 1904) . Publ. Recs. ECU: (P11), (H14), Dalzochio et al. (2018); NA:*Daigle (1996). Also ARG, BOL, BRA, GUY, PRY, SUR, VEN: (H06), (P11); plus COL: (P&P).

Addl. Data. OR, Francisco de Orellana: 1 ♂, Río Tiputini slough, km marker 31 at Yasuni Road , Yasuni Pre- serve, 17 xi 1997, leg. & det. JJD; verified ix 2011 -JJD [JJD].

Erythrodiplax parvimaculata Borror, 1942 . Publ. Recs. ECU: (H06), (P11), (H14); SU: * Donnelly (1998). Also BOL, VEN: (H06), (P11); plus BRA: (H06); and COL Palacino et al. (2017).

Addl. Data. OR, La Joya de los Sachas: 1 ♂, ditch, Pompaya, 0.5 km N of Río Napo ferry, 19 vii 1996, leg. & det. KJT [KJT]; Loreto: 2 ♂♂, Pano, 12 ix 1990, leg. OSF, det.? [USNM]; SU, Shushufindi: La Selva Lodge, forest (-0.4983, -76.3733, 200m), 25 xi 1997, leg. & det. TWD [SWD]; 2 ♀♀, same data but 12 x 1988, leg. & det. SWD [SWD]; 1 ♂, same data but 13 x 1988 [SWD]; 1 ♂, Shushufindi , 3 km SW, stream (-0.2136, -76.6786), 12 v 2009, leg. & det. KJT [FSCA].

Erythrodiplax tenuis Borror, 1942 . Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), (P11), (H14); NA: * Borror (1942 —description), Bridges (1993), Tennessen (1996b), OR: Tennessen (1998), von Ellenrieder & Garrison (2011). Also PER, BRA (P11).

Addl. Data. NA, Carlos Julio Arosemena Tola: 9 ♂♂, 9 ♀♀, [Ila], 52 km N of Puyo on Road to Tena (-1.17, -77.85, 527m), 26 x 1980, leg. & det. MJW [FSCA]; El Chaco: 1 ♀, stream, 53 km N of Chaco, 18 viii 1980, leg. & det. SWD [SWD]; Tena: 1 ♂, J.S.B.S., Pond #18, 19 xi 1991, leg. E. McCallie, det.? [USNM]; 1 ♀, Río Chinchipino, 27 km E of Puerto Napo (-1.0728, -77.5864, 385m), 15 vi 1995, leg. & det. KJT [KJT]; 3 ♂♂, same data but 13 xi 1997, leg. & det. JJD [JJD]; 2 ♂♂, sedge marsh, 1.9 km E of 2nd stream E of Río Sinde, E of Puerto Napo on J.S.B.S. Road, 23 vii 1996, leg. & det. SWD [SWD]; OR, Francisco de Orellana: 1 ♀, trib. of Río Pay- amino, road to Loreto, 1.7 km W of jct. with Coca Road (-0.4367, -77.0017, 258m), 13 vi 1995, leg. & det. KJT [KJT]; 1 ♂, Puerto Francisco de Orellana, 8.2 km W on road to Loreto from Coca-Sachas Road, pond (-0.4578, -77.0381, 288m), 18 xi 1997, leg. & det. WFM [FSCA]; 2 ♂♂, 2 ♀♀, pond, P.N.Y., km marker 29.5, 17 xi 1997, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, Anangu SE across Río Napo from Providencia Lago Anangu, “on Río Napo ” (-0.5213, -76.4361, 300m), 9 v 1975, leg. T. Emmel, det. MJW [FSCA]; PA, Pastaza: 4 ♂♂, 1 ♀, pond 3 km N of Comunidad Chuwitayo, S of Puyo, 9 xi 1997, leg. & det. KJT [KJT]; 7 ♂♂, 3 ♀♀, Cuasha, approx. 4 km N on Hwy. 45 (94 km N of Sucua), cut grass pond (-1.83, -77.82, 884m), 9 x 1997, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 2 ♂♂, pond on road between Macas and Puyo (-1.8633, -77.8150, 900m), 9 xi 1997, leg. & det. TWD [SWD]; Santa Clara : adults, 39 km N of Puyo, 5 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 1 ♀, Puyo Road , 19.3 km S of Río Napo, near Zatzayaco (455m), 21 vii 1996, leg. & det. KJT [KJT]; SU, Lago Agrio : adults, 2 km N of Lago Agrio, 26 ix 1975, leg. A. Langley, det.? [USNM]; adults, same data but 27 viii 1975 [USNM]; Shushufindi: 1 ♀, Limoncocha, Río Napo (-0.4000, -76.6167, 269m), 4 x 1974, leg. B Drummond III, det. MJW [FSCA]; ZC, El Pangui , 2 ♂♂, near stream about 15 km S of El Pangui, 6 xi 1997, leg. & det. KJT [KJT].

Erythrodiplax umbrata (Linnaeus, 1758) . Publ. Recs. ECU: * Calvert (1906), Ris (1911a), Paulson (1983), Tsuda (1991), (H06), (P11), (H14), Dalzochio et al (2018); CR: Campos (1922); CB: Campos (1922); EO: Campos (1927); GY: Calvert (1906), Campos (1922, 1932), Borror (1942); LR: * Calvert (1906), Campos (1922); NA: Borror (1942). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, URY, VEN: (H06), (P11); plus TTO: (P11).

Addl. Data. ES, Esmeraldas: [DRP, QZAC]; San Lorenzo: [ANSP, FSCA]; GY, Daule: [USNM]; Duran: [ANSP]; Guayaquil: [ANSP, FSCA, USNM]; Naranjal: [FSCA]; Playas: [FSCA]; LJ, Macará: [ANSP]; LR, Babahoyo: [USNM]; Buena Fé: [FSCA]; Montalvo: [JJD]; MN, Chone: [USNM]; El Carmen: [QZAC]; Sucre: [USNM]; NA, Tena: [DRP, FSCA, UTIC, KJT, LSAM, USNM]; OR, Francisco de Orellana: [FSCA, KJT, QZAC, USFQ]; Loreto: [KJT]; SD: [USNM]; SU, Cascales: [USNM]; Lago Agrio: [UTIC, QZAC]; Shushufindi: [DRP, FSCA, UTIC, USNM]; ZC, Zamora: [USNM].

Erythrodiplax unimaculata (De Geer, 1773) . Publ. Recs. ECU: Tsuda (1986), (H06), (P11), (H14); PA: * Karsch (1890 —as Micrathyria hemimelaena Holotype ♂: [PA], “Sarayacu”), Ris (1911), Borror (1942). Also BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, SUR, VEN: (H06), (P11); plus TTO: (P11).

Addl. Data. NA, El Chaco, 1 ♂, stream 39 km NE Chaco, 18 viii 1980, leg. & det. SWD [RWG]; Tena: 1 ♂, pool, Río Sinde, 5 km E of Puerto Napo (-1.0517, -77.7544, 450m), 13 vi 1996, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but 12 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♂, 4 ♀♀, Reserva Etnica Waorani, Onkone Gare Camp , along road (-0.63 -76.6), 30 i 1994, leg. P. Araujo, det.? [USNM]; 1 ♂, San Francisco de Orel- lana [downriver about 20 minutes on the south shore], [Arcoiris Jungle Lodge], San Carlos on Río Napo (-0.4550, -76.0042, 300m), 8 v 1975, leg. T. Emmel det. MJW [FSCA]; 1 ♀, same data but leg. B. Drummond III, det. MJW [FSCA]; 4 ♀♀, Primavera, forest trail from Río Napo to Laguna Taracoa, 27 vii 1977, leg. & det. DRP [DRP]; 3 ♂♂, 2 ♀♀, [La Nueva Esperanza] [across Río Napo from] near Primavera “Limoncocha”, Río Napo & Lake Tara- coa (-0.45, -76.77, 300m), 5 ♂♂, 4 ♀♀, same data but 26 viii 1980, leg. & det. SWD [JJD, KJT, SWD]; 3 ♂♂, 10 ♀♀, same data but 4 xi 1980, leg. & det. MJW [FSCA]; 1 cop. pair, same data but 26 xi 1980 [FSCA]; 3 ♀♀, same but 9 vii 1983, leg. T. & P. Gamtenbein, det. RWG [CSCA]; 2 ♂♂, 1 ♀, Yuturi, 25 ii 1990, leg. & det.? [QZAC]; 1 ♂, Payamino, km 13.7, 30 xii 1983, leg. & det.? [QZAC]; 1 ♂, P.N.Y., ditches around research station, 11 vi 1995, leg. & det. SWD [SWD]; 1 ♂, P.N.Y., 2.3 km E of Yasuni Biological Research Station, small stream (-0.67, -76.37, 183m), 9 vi 1995, leg. & det. WFM [FSCA]; 3 ♂♂, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (-0.60, - 76.47W, 229m), 12 vi 1995, leg. & det. KJT [KJT]; 3 ♀♀, 12 vi 1995, leg. & det. WFM; 1 ♂, 1 ♀, same data but 17 vii 1996 [FSCA, QZAC, USNM]; 1 ♂, 1 ♀, same data but 17 xi 1997, leg. & det. TWD [SWD]; 1 ♂, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (-0.4367, -77.0017, 258m), 13 vi 1995, leg. & det. WFM [FSCA]; La Joya de los Sachas: 1 ♂, 1 ♀, Pompeya near Puerto Pompeya Ferry Landing (-0.4400, -76.6194, 240m), 17 vii 1996, leg. & det. WFM [QZAC]; 1 ♂, 1 ♀, 0.5 km W of Puerto Pompeya Ferry Landing roadside ditches (-0.4444, -76.6263, 238m), 19 vii 1996, leg. & det. WFM [FSCA]; PC, Quito: 1 ♂, Guayllabamba (2000m), 9 i 1993, leg. & det.? [QZAC]; San Miguel Bancos: 1 ♂, Puerto Quito Endega (750m), xii 1983, leg. & det.? [QZAC]; SU, Cascales: 1 ♂, [El Dorado de Cascales], 58 km W of Lago Agrío on Hwy. 45 at “El Paraiso”, stream/marsh (-0.08, -77.20, 366m), 8 vi 1995; leg. & det. WFM [FSCA]; Lago Agrio: 2 ♂♂, 1 ♀, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]; 2 ♀♀, stream, E edge of Lago Agrio, 18 viii 1980, leg. & det. SWD [SWD]; 1 ♂, pond 3 km W of Lago Agrío on Hwy. 45 (-0.97, -76.83, 275m), 8 vi 1995, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [LSAM]; Shushufindi : 4 ♂♂, 1 ♀, Limoncocha , Río Napo (-0.4000, -76.6167, 269m), 4–6 ix 1972, leg. T. Emmel, det. MJW & SWD [FSCA]; 1 ♂, 1 ♀, same data but 21 iii 1974 [FSCA]; 6 ♂♂, 2 ♀♀, same data but 11–14 v 1975 [FSCA]; 5 ♀♀, same data but 5–24 xi 1973, leg. B. Drummond III, det. MJW & SWD [FSCA]; 1 ♂, same data but 11 i 1974 [FSCA]; 1 ♂, same data but 17 iii 1974 [FSCA]; 1 ♂, 2 ♀♀, same data but 1–31 viii 1974 [FSCA]; 7 ♂♂, 6 ♀♀, same data but 10–15 v 1975 [FSCA]; 1 ♀, same data but 21 xii 1979, leg. C. Burne, det. SWD [FSCA]; 3 ♂♂, 5 ♀♀, same data but 23–28 viii 1980, leg. & det. SWD [SWD]; 21 ♂♂, 24 ♀♀, same data but 31 x –24 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, 1 ♀, same data but lake edge, 5 vii 1971, leg. D. Pearson, det. DRP [DRP; 1 ♂, same data but 2 x 1971 [DRP]; 1 ♂, same data but 11 xii 1971 [DRP]; 1 ♀, same data but 13 i 1972 [DRP]; 1 ♂, same data but 20 ii 1972 [DRP]; 1 ♀, same data but 24 ii 1972 [DRP]; 6 ♀♀, same data but road through partially cleared forest, 12 viii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but forest trail, 10 viii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but 28 i 1972 [DRP]; 1 ♂, same data but 1 iii 1972; 3 ♀♀, same data but clearing, 11/ 12 viii 1972, leg. E. Stiles, det. DRP [DRP]; 11 ♀♀, same data but 23–26 vii 1977, leg. & det. DRP [DRP]; 3 ♂♂, same data but swamp, 26 vii 1977, leg. & det. DRP & S. Hills [DRP]; 2 ♂♂, 4 ♀♀, [Limoncocha] [about 40 km E] or 100 km E of Coca [on Río Napo], Lago Garzacocha La Selva Jungle Lodge (-0.4819, -76.3733, 235m), 12 x 1988, leg. & det. SWD [FSCA, USNM]; 1 ♂, same data but 13 iv 1992, leg. & det. E. Smith [SWD]; 1 ♂, 1.2 km W on Hwy. 22, Río Shushufindi (-0.1853, -76.6617, 255m), 16 vii 1996, leg. & det. WFM [LSAM].

Figures

FIGURE 95a. Erythrodiplax fusca ♂, ECUADOR, Orellana Province, pond, road SW of Loreto, 2 xi 2013; photo K. J. Ten- nessen.