Clinonana, Osborn, 1938

Jones, Joshua R. & Deitz, Lewis L., 2009, Phylogeny and systematics of the leafhopper subfamily Ledrinae (Hemiptera: Cicadellidae) 2186, Zootaxa 2186 (1), pp. 1-120 : 67

Genus Clinonana reinstated to Scarinae : Gyponini

Clinonana Osborn, 1938: 13 . Type species: C. magna Osborn, 1938: 13 , pl. I, fig. 2. Clinonana was originally described in Gyponinae Stål, 1870 (= Scarinae Amyot and Serville, 1843 ), but Kramer (1966) considered it to belong to Ledrinae based on the formula of the macrosetal formula of the metathoracic femur (2+1) and its “general habitus.” While other similarities exist between this genus and many ledrines—including the position of the ocelli on the crown, the punctate texture of the dorsum, the large size, green color, and most notably, the laterally produced carinate extensions off the pronotum (reminiscent of Tituria and Neotituria )— these characteristics can all be attributed to plesiomorphy, ecological convergence, or both. A close examination of the metathoracic femur of C. mirabilis reveals numerous smaller auxiliary setae subapical to the apical macrosetae of which Kramer makes no mention (Pl. 13B), which are completely uncharacteristic of ledrines (except Xedreota ), and other characters such as the shape of the face and presence of the complete appendix on the forewing are all characteristic of Scarinae . The phylogenetic analyses herein (see “Phylogenetic results and discussion” section under Results and Discussion, above) placed Clinonana with Gypona Germar. Material examined: C. mirabilis : 1 male, 3 females, 1 (abdomen missing), Guiana, NCSU, JRJ_Led1_031–032, 034–035, 037, 2 males, 2 females, Guiana, MNHN, JRJ_Led1_029–030, 033, 036. Pl. 5J, 13B, 15A, 18D.