Vindilisina subtribe nov.

Bourgoin, Thierry & Wang, Menglin, 2020, Parahiraciini (Hemiptera, Fulgoromorpha, Issidae): one new genus, two new species and three new subtribes, ZooKeys 997, pp. 69-94 : 69

Vindilisina subtribe nov. Figs 49 , 50

Type genus.

Vindilis Stål, 1870.

Diagnosis.

The subtribe is identified and separated according to the following list of characters:

Head capsule with vertex and frons in a slightly convex margin in lateral view. Compound eyes elongated, almost twice as long as wide in lateral view.

Hindwing trilobate, strongly notched at CuA2-CuP; Sc-R-MP-CuA and CuP-Pcu-A1 lobes almost the same wide.

Posterior margin distinctly notched at A1 2.

A2 lobe surpassing half-length of CuP-Pcu-A1 lobe, apically rounded and about 1/4 thinner than the anterior one, and with margins sub-parallel.

Sc-R-MP-CuA lobe not covered with a set of numerous transverse veins, a few incomplete ones in CuP-Pcu-A1 lobe.

CuA and CuP merging well before the anterior notch.

Pcu and A1 merging in basal half of forewing on some distance.

A2 present, single.

Note.

The genera Vindilis and Nisoprincessa are transferred in this new subtribe, which is currently distributed only in Philippines (Palawan) ( Gnezdilov 2017). The strong CuA-CuP notch looks similar to the other Parahiraciini , but both genera lack the reticulate venation of the Parahiraciina subtribe nov. or Scantiniina subtribe nov. In return, with exception of Hemisphaeriini , they share with Scantiniina subtribe nov. and other Hemisphaeriinae taxa the distinct plesiomorphic trilobate conformation of the hindwing. With Sarimini Wang, Zhang & Bourgoin, 2016 and Kodaianellini Wang, Zhang & Bourgoin, 2016, they exhibit the basal apomorphic merging of veins Pcu and A1. Without molecular data, its position in the phylogeny of Hemisphaeriinae remains uncertain and thus is currently left as a subtribe in the Parahiraciini .

Figures

Figures 47 - 52. Hindwing pattern and venation. 47, 48 Scantiniina subtrib. nov.: Scantinius bruchoides (Walker, 1858) (modified from Gnezdilov 2012, fig. 3) 49, 50 Vindilisina subtrib. nov.: Vindilis fornicata Stal, 1870 (modified from Gnezdilov 2018, fig. 3) 51, 52 Parahiraciina Cheng & Yang, 1991 subtrib. nov.: Tetricodes tamdaoensis Vanslembrouck & Constant, 2018 (modified from Vanslembrouck and Constant 2018, fig. 4).