Hemibagrus spilopterus Ng & Rainboth, 1999
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publication ID |
https://doi.org/ 10.5281/zenodo.5351788 |
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persistent identifier |
https://treatment.plazi.org/id/CF180962-FFCE-FF9C-FF40-F16AFBC98058 |
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treatment provided by |
Tatiana |
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scientific name |
Hemibagrus spilopterus Ng & Rainboth, 1999 |
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Hemibagrus spilopterus Ng & Rainboth, 1999 View in CoL
( Fig. 28 View Fig )
Hemibagrus spilopterus Ng & Rainboth, 1999: 562 View in CoL , Figs. 5 View Fig , 7a View Fig , 8a View Fig (type locality: Kandal, Bassac River at village of Prek Chey on Vietnamese border, Cambodia); Ferraris, 2007: 91; Vidthayanon, 2008: 114; Ng & Lim, 2008: 29; Rainboth et al., 2012: 72, Pl. 46, Fig. 946.
Bagrus nemurus View in CoL (non Valenciennes in Cuvier & Valenciennes, 1840) - Bleeker, 1859f: 102.
Hemibagrus nemurus View in CoL (non Valenciennes in Cuvier & Valenciennes, 1840) - Bleeker, 1864: 353; 1865a: 34; 1865b: 175; Sauvage, 1883: 154; Mo, 1991: 132; Kottelat, 1998: 100, Fig. 147; Baird et al., 1999: 91, Fig. 201; Gaanantkul et al., 2000: 34; Serov et al., 2006: 193; Vasil’eva & Vasil’ev, 2012: 201.
? Macrones luridus Tirant in Pétillot, 1911: 164 (nomen nudum); Chevey, 1929: 172
Macrones nemurus (non Valenciennes in Cuvier & Valenciennes, 1840) - Hora, 1923: 171; 1924: 468; Tirant in Pétillot, 1911: 164; Chevey, 1929: 172; 1932: 17; 1934: 201; Chevey & Le Poulain, 1940: 38.
Macrones planiceps (non Valenciennes in Cuvier & Valenciennes, 1840) - Chevey, 1932: 17; Pellegrin & Chevey, 1937: 317.
Mystus nemurus View in CoL (non Valenciennes, in Cuvier & Valenciennes, 1840) - Fowler, 1934: 95; 1937: 148, Figs. 44-47 View Fig View Fig View Fig (in part); Suvatti, 1936: 73; 1950: 297 (in part); 1981: 92; Kuronuma, 1961: 6; Mizuno & Mori, 1970: 109, Fig. 12C View Fig ; Kawamoto et al., 1972: 27, Fig. 52; Orsi, 1974: 161; Desoutter, 1975: 448; Beeckman & de Bont, 1985: 2646; Kottelat, 1985: 270; Mai & Nguy n, 1988: 49; Roberts, 1993b: 33; Serov, 1994a: 24; 1994b: 62; Lim et al., 1996: 382; Rainboth, 1996: 143, Pl. 19 Fig. 145; Nguy n, 1999: 32.
Mystus planiceps View in CoL (non Valenciennes in Cuvier & Valenciennes, 1840) - Suvatti, 1936: 75; 1950: 297 (in part); 1981: 92; Fowler, 1937: 148; 1939: 40, 58; Kuronuma, 1961: 6; Desoutter, 1975: 451; Kottelat, 1985: 270; Serov, 1994a: 24; 1994b: 63; Vasil’eva & Vasil’ev, 2012: 201.
Mystus wyckii View in CoL (non Bleeker, 1858) - Trýõng & Tr n, 1993: 213, Fig. 92.
Hemibagrus cf. nemurus - Ng & Rainboth, 1999: 558, Fig. 3 View Fig .
Hemibagrus aff. nemurus View in CoL - Kottelat, 2001b: 122, Fig. 329.
Hemibagrus sp. 1 - Nguy n, 2005: 359, Fig. 184.
Hemibagrus sp. cf. nemurus View in CoL - Rainboth et al., 2012: 72, Pl. 45, Fig. 945.
Hemibagrus planiceps View in CoL (non Valenciennes in Cuvier & Valenciennes, 1840) - Vasil’eva & Vasil’ev, 2012: 201.
Material examined. — MEKONG: LAOS: NRM 15001, 1 ex., 267.8 mm SL, Nam Ou River near Luang Prabang ; NRM 15015, 1 ex., 260.3 mm SL ; NRM 15016, 1 ex., 235.1 mm SL ; NRM 15017, 1 ex., 253.3 mm SL ; NRM 15018, 1 ex., 222.4 mm SL, Mekong River at Luang Prabang, 19°53'N 105°50'E GoogleMaps ; BMNH 1976.7.1.35–36, 2 ex., 157.5–171.4 mm SL, fish market at Vientiane ; CMK 13129, 1 ex., 169.6 mm SL, Vientiane province: Mekong River at mouth of Nam Mang and lower 100 m of Nam Mang , 18°21'48"N 103°14'16"E GoogleMaps ; CAS 92472, 4 ex., 23.8–39.4 mm SL, Attapeu province: Houay Samong, about 1 km upstream from its mouth into Xe Pian near Ban Hin Lat , Sekong watershed ; CAS 92504, 15 ex., 20.8–150.6 mm SL, Sekong watershed, small mountain stream entering right side of Se Kaman just upstream from Se Kaman 1 hydropower ; CAS 94757, 1 ex., 27.0 mm SL, Mekong drainage, Sekong watershed, Bolaven plateau, Xe Nam Noy mainstream 26 km by road from Ban Nam Tong about 1 km downstream, elevation 270 m ; CAS 94772, 3 ex., 20.0– 33.1 mm SL, Attapeu province: Sekong watershed, Xe Pian 1 km downstream from Se Pa waterfall and 5–6 km upstream from Ban Hin Lat ; CAS 96793, 1 ex., 91.0 mm SL, Se Khone .
MYANMAR: ANSP 59391 View Materials , 2 ex., 72.7–110.5 mm SL, Mekong River at Chieng Sen, 8 km S of Shan States border opposite Laos .
THAILAND: ANSP 89408 View Materials , 13 ex., 71.2–124.2 mm SL, Khemerat ; CAS 61915, 2 ex., 147.8–152.9 mm SL ; CAS 61920, 2 ex., 112.0– 112.2 mm SL ; UMMZ 186800 View Materials , 7 ex., 74.2–103.6 mm SL ; USNM 317587 View Materials , 21 ex., 77.4–111.9 mm SL ; USNM 317589 View Materials , 8 ex., 109.9–136.4 mm SL ; USNM 317607 View Materials , 1 ex., 90.3 mm SL, Ubon Ratchathani province, market at Ubon Ratchathani ; CAS 79051, 4 ex., 72.8–126.0 mm SL, Mekong river near Ban Ha Bia, 30–40 km NW of Chiang Khan on Highway 2186; CAS 96568, 4 ex., 145.4–204.8 mm SL, Mukdahan market ; CAS 96573, 5 ex., 83.9–185.3 mm SL, Mekong mainstream between Pak Ing and Jom Paing (about 4–5 km downstream) ; CAS 96582, 1 ex., 157.3 mm SL, Pak Menam Mun (mouth of Mun River ) ; CAS 96966, 3 ex., 66.5–68.9 mm SL, Huay Nam Ngam & Mekong mainstream near That Phanom ; CAS 97076, 1 ex., 68.7 mm SL, Menam Chi at Maha Chana Chai ; CMK 5043 , 4 ex., 99.1–152.8 mm SL, market at Nong Khai ; CMK 5087 , 2 ex., 124.4–139.0 mm SL ; ZRC 46012, 4 ex., 146.0–184.0 mm SL, market at That Phanom ; UMMZ 186685 View Materials , 10 ex., 45.3–56.5 mm SL, Nakorn Ratchasima province, ditches and flow, and borrow pits along drainage downstream from damsite at Pakthongchai, Korat ; UMMZ 186752 View Materials , 3 ex., 75.0– 100.9 mm SL, Khon Kaen fish market ; UMMZ 186756 View Materials , 1 ex., 84.1 mm SL, Pangneeb province, Nam Phong River, about 90 km S of Udorn (25 km W of Nam Phong village ) ; UMMZ 186769 View Materials , 3 ex., 113.6– 159.5 mm SL, market at Nakhon Phanom ; UMMZ 186779 View Materials , 5 ex., 61.0– 141.9 mm SL, Nam Kham River at base of dam holding Nong Lahan ; UMMZ 186784 View Materials , 2 ex., 168.0– 221.9 mm SL, Ubon Ratchathani province, market at Mukdahan ; UMMZ 186791 View Materials , 1 ex., 210.4 mm SL, irrigation reservoir at Ubon Ratchathani fisheries Station ; UMMZ 186814 View Materials , 1 ex., 63.6 mm SL, Ubon Ratchathani province, Mun River about 20 km downstream from Ubon Ratchathani ; UMMZ 186818 View Materials , 1 ex., 75.0 mm SL, Ubon Ratchathani province, Mun River about 5 km downstream from Ubon Ratchathani ; UMMZ 186820 View Materials , 14 ex., 52.4–156.6 mm SL, Reservoir at Maha Sarakhan . UMMZ 186827 View Materials , 2 ex., 138.8–150.5 mm SL, market at Korat ; UMMZ 186852 View Materials , 3 ex., 104.3–116.0 mm SL, Mool River of the Mekong drainage ; UMMZ 213975 View Materials , 13 ex., 67.3–182.1 mm SL, Nong Khai province, Tha Bo district, Huay Kum Kang Mo at Mekong River ; UMMZ 214368 View Materials , 3 ex., 98.1–119.0 mm SL, Ubon Ratchathani province, Mekong River at Khemerat in front of River Patrol Station S of town . UMMZ 214370 View Materials , 1 ex., 98.5 mm SL, Ubon Ratchathani province: Khong Chiam district, N shore of Mun River at Ban Dan, 2 km upstream from Mekong confluence. UMMZ 214371 View Materials , 1 ex., 185.0 mm SL, Ubon Ratchathani province, Khong Chiam district, Mun River at Ban Dan, 1.5 km upstream from Mekong confluence; UMMZ 214372 View Materials , 3 ex., 118.0– 141.3 mm SL, Ubon Ratchathani province, Mun River, 3 km downstream from bridge at Ubon Ratchathani, at mouth of small creek on N bank of the Mun ; UMMZ 214373 View Materials , 2 ex., 144.1–152.2 mm SL, Ubon Ratchathani province: Khong Chiam district, Mun River at Ban Dan, 3 km upstream from confluence with Mekong River ; UMMZ 214374 View Materials , 2 ex., 36.0– 41.8 mm SL , UMMZ 214375 View Materials , 1 ex., 42.9 mm SL ; UMMZ 214376 View Materials , 3 ex., 34.7–48.7 mm SL ; UMMZ 214377 View Materials , 1 ex., 31.9 mm SL ; UMMZ 214378 View Materials , 4 ex., 33.9–42.9 mm SL, Ubon Ratchathani province, Khong Chiam district, Mun River at Ban Dan, 3 km upstream from confluence with Mekong River ; UMMZ 214379 View Materials , 1 ex., 106.2 mm SL, Ubon Ratchathani province, stream from Bung Klang Huen to Mun River, 10 km from Mun River , 3 km E of Ubon Ratchathani ; UMMZ 214380 View Materials , 5 ex., 43.8–51.6 mm SL, Ubon Ratchathani province, Lam Dom Noi Reservoir near Ban Kham Kom ; UMMZ 214381 View Materials , 1 ex., 56.8 mm SL, Nakhon Phanom province, Mekong River at Ban Tha Kai , 20 km downstream from Mukdahan ; UMMZ 214382 View Materials , 2 ex., 137.4– 156.5 mm SL, Ubon Ratchathani province, Mun River 15 km downstream from bridge at Ubon Ratchathani ; UMMZ 214383 View Materials , 12 ex., 15.1–125.7 mm SL, Ubon Ratchathani province, Huay Kwang , 1.5 km upstream from Mun River ; UMMZ 214384 View Materials , 2 ex., 51.6–203.3 mm SL, Khon Kaen province: Nam Pong Reservoir , 3 km N of fish landing on E side of reservoir ; UMMZ 214385 View Materials , 8 ex., 13.0– 24.8 mm SL, Ubon Ratchathani province, 6 km W of Ubon on highway 23, inundated scrub vegetation near flooded creek; UMMZ 214386 View Materials , 10 ex., 100.8–150.7 mm SL, Kung Kang Moo, creek mouth into Mekong River ; UMMZ 214387 View Materials , 25 ex., 40.5–116.2 mm SL, Ubon Ratchathani province, Huay Thom –loe at Ban Bung Khee-lek, 7 km E of Khemerat , 2.5 km from Mekong River ; UMMZ 214388 View Materials , 26 ex., 39.0– 104.8 mm SL, Ubon Ratchathani province, Huay Thom-loe at Ban Bung Khee–lek, 7 km E of Khemerat , 1 km from Mekong River ; UMMZ 214389 View Materials , 20 ex., 38.6–134.5 mm SL, Nakhon Phanom province, Mekong River at Ban Tha Kai , 21 km downstream from Mukdahan ; UMMZ 214390 View Materials , 1 ex., 83.0 mm SL, Ubon Ratchathani province, Huay Kwang , S of Khong Chiam , 3 km up Mun River from Mekong–Mun River confluence ; UMMZ 214391 View Materials , 2 ex., 34.2–66.2 mm SL, Ubon Ratchathani province, Huay Phai , 7 km N of Khong Chiam , at confluence with Mekong River ; UMMZ 214392 View Materials , 13 ex., 21.3–69.5 mm SL, Nakhon Phanom province, Huay Nong Sao , 8 km SW of Nakhon Phanom , 1.5 km down footpath off main road; UMMZ 214393 View Materials , 6 ex., 90.5–215.6 mm SL, Huay Mark , 8 km N of Khong Chiam , at confluence with Mekong River , 15°22'N 105°29'E GoogleMaps ; UMMZ 214394 View Materials , 1 ex., 114.0 mm SL, Ubon Ratchathani province, Huay Kwang, at confluence with Mun River, 3 km from Mekong River ; UMMZ 214395 View Materials , 15 ex., 50.0– 134.8 mm SL, Ubon Ratchathani province: Huay Mark Tai, 1 km from Mekong River ; UMMZ 214396 View Materials , 1 ex., 183.4 mm SL, Ubon Ratchathani province: Lam Dom Noi reservoir, approx. 2.5 km S of Ban Khan Kom on W shore; UMMZ 214397 View Materials , 44 ex., 19.9–131.7 mm SL, Khon Kaen province: Nam Pong Reservoir , 2.1 km N of damsite on E shore ; UMMZ 214398 View Materials , 4 ex., 55.2–98.9 mm SL, Khon Kaen province: Nam Pong Reservoir , 4.8 km N of damsite on E shore ; UMMZ 214938 View Materials , 4 ex., 55.2–98.9 mm SL, Khon Kaen province: Nam Pong Reservoir , 4.8 km N of damsite on E shore ; UMMZ 214399 View Materials , 8 ex., 38.2–105.1 mm SL , UMMZ 214400 View Materials , 8 ex., 33.7–92.5 mm SL , UMMZ 214401 View Materials , 12 ex., 61.0– 129.8 mm SL, Nong Khai province, Tha Bo district, Mekong River at Ban Tha Sa-det ; UMMZ 214402 View Materials , 3 ex., 50.0– 72.6 mm SL, Nakhon Phanom province, roadside ditch near Ban Thong, 6 km S of That Phanom ; UMMZ 214403 View Materials , 3 ex., 66.6–77.9 mm SL, Ubon Ratchathani province, Mun River , 1 km upstream of Ubon Ratchathani ; UMMZ 214404 View Materials , 3 ex., 56.0– 93.7 mm SL, Ubon Ratchathani province, Mun River , 3 km downstream from Ubon Ratchathani ; UMMZ 214406 View Materials , 11 ex., 37.6–55.3 mm SL, Chiang Mai province, Fang River , 23 km S of Fang on Vuang Mai Road –Highway 107; UMMZ 214407 View Materials , 1 ex., 33.5 mm SL, Ubon Ratchathani province, Huay Hin Taek, just across Mun River from Ban Dan, 1.5 km up Huay Hin Taek from Mun River ; UMMZ 214408 View Materials , 5 ex., 16.8–23.4 mm SL, Ubon Ratchathani province, Khong Chiam district, Huay Hin Taek, just across Mun River from Ban Dan , 0.75 km up Huay Hin Taek from Mun River ; UMMZ 214409 View Materials , 1 ex., 137.8 mm SL, Ubon Ratchathani province, Mun River , just below Ubon Ratchathani ; UMMZ 214412 View Materials , 1 ex., 191.3 mm SL, Ubon Ratchathani province, Khong Chiam district, Mun River at Ban Dan, 3 km upstream from Mekong confluence; UMMZ 214487 View Materials , 47 ex., 40.7–138.1 mm SL, Ubon Ratchathani province: Khong Chiam district, Huay Kwang, 0.3 km from Mun River ; UMMZ 224551 View Materials , 4 ex., 53.2–125.8 mm SL, Ubon Ratchathani province, Lam Dam Noi Reservoir at Ban Kham Kom ; UMMZ 226867 View Materials , 5 ex., 72.6–113.1 mm SL , UMMZ 226868 View Materials , 20 ex., 81.5–121.5 mm SL, Nong Khai province, Tha Bo district, Mekong River backwater at Ban Tha Sadet, 17°52'36"N 102°35'30"E GoogleMaps ; USNM 109580 View Materials , 1 ex., 76.8 mm SL, Mekong at Chieng Sen Kao ; USNM 109582 View Materials , 7 ex., 45.2–71.7 mm SL, N Thailand ; USNM 305707 View Materials , 2 ex., 101.4–133.1 mm SL, Ubon Ratchathani province, Warin Chamrap market ; USNM 317576 View Materials , 1 ex., 135.7 mm SL, Thailand , 15°14'N 104°52'E GoogleMaps ; USNM 317588 View Materials , 1 ex., 162.4 mm SL, Ubon Ratchathani province, Victory Rapids (Kaeng Ja wa) on Mun river about 5 km W of Mekong River ; USNM 317590 View Materials , 17 ex., 96.5–118.0 mm SL, Khon Kaen province, Huai Phra Ku , about 7 km E of Khon Khaen on highway 209; USNM 317591 View Materials , 2 ex., 124.9–186.8 mm SL, Roi Et province, Lam Chi , 1.5 km below highway 23 bridge 4 km W of Selaphum ; USNM 317592 View Materials , 13 ex., 101.4–142.0 mm SL, Ubon Ratchathani province, Mun River at Bung Wai about 7 km W of Ubon ; USNM 317593 View Materials , 6 ex., 142.8– 182.4 mm SL, Thailand, Khon Kaen province, Mekong River drainage, 15°45'N 102°37'E GoogleMaps ; MCZ 60936, 1 ex., 216.0 mm SL, Khon Kaen province, Lake Kaeng Lawa , 16°8'N 102°42'E GoogleMaps ; USNM 317594 View Materials , 32 ex., 84.0– 127.5 mm SL, Nakhon Phanom province, market at Nakhon Phanom ; USNM 317595 View Materials , 3 ex., 163.8–200.1 mm SL, Nong Khai province, Mekong River near Ban Kong Nang ; USNM 317608 View Materials , 1 ex., 115.5 mm SL, Nakhon Ratchasima province, flood plain pool of Lam Nam Mun at Amphoe Phimai ; USNM 317609 View Materials , 1 ex., 145.8 mm SL, Ubon Ratchathani province, purchased from fisherman on Mae Nam Mun , just E of Phibun Mangsahan ; ZRC 40896, 3 ex., 59.2–101.2 mm SL, Nakhon Ratchasima province, drained irrigation ponds or padi fields along Highway 24, fed by Klong Kabau, 14°44'34.8"N 102°12'31.7"E GoogleMaps ; ZRC 46157, 1 ex., 157.0 mm SL, market at Phibun Mangsahan , 46 km E of Ubon Ratchathani ; ZRC 39345, 4 ex., 95.2–135.3 mm SL, Yasothon province, Amphoe Muang Yasothon .
CAMBODIA: UMMZ 232611 View Materials , holotype, 147.5 mm SL, Kandal: Bassac River at village of Prek Chey on Vietnamese border, 10°57'N 105°6'E GoogleMaps ; CAS 93949, 1 paratype, 203.2 mm SL, Stung Siem Reap near Angkor Wat ; CMK 15121, 1 paratype, 168.2 mm SL ; CAS 209760 View Materials , 1 paratype, 171.0 mm SL ; ZRC 43324, 10 paratypes, 171.7–241.4 mm SL, Phnom Penh , Chompuvon market, approx. 5 km SW of city ; ZRC 43322, 1 paratype, 209.9 mm SL, Phnom Penh, Chipa Ampe market, across Tonle Sap River NE of city ; ZRC 43323, 1 paratype, 309.1 mm SL, Phnom Penh , central market ; UMMZ 181163 View Materials , 2 paratypes, 98.8–109.0 mm SL, Kandal: Prek Andhor, tributary from E of Tonlé Sap River , 15 km NW of Phnom Penh ; UMMZ 232644 View Materials , 1 paratype, 126.5 mm SL, Kandal: Prek Bak Nam at fishing lot 9, just upstream of Phum Chong Sao , 11°2'N 105°8'E GoogleMaps ; UMMZ 232699 View Materials , 3 paratypes, 206.4–230.0 mm SL, Kompong Chhnang , morning market, 12°15'N 104°40'E GoogleMaps ; UMMZ 232731 View Materials , 2 paratypes, 93.0–127.0 mm SL, Tonlé Sap at exit to Great Lake , 4 km NW of Chhnok Trou, at Kompong Thom fishing lot 2 ; CAS 93268, 5 ex., 36.1–43.5 mm SL, Ratanakiri province, O Chalerm, approx. 5–10 km E of Lomphat ; CAS 94779, 47 ex., 10.5–104.6 mm SL, Ratanakiri province, O Champha, 4–8 km upstream from its mouth into Tonlé San near Te Veng ; CAS 96642, 5 ex., 43.3–129.4 mm SL, Ratanakiri province, O Kamang Chong, 20–25 km E of Lomphat ; MNHN A1071 View Materials , 3 ex., 197.8–273.4 mm SL, Cambodia . MNHN 1937–34 View Materials , 1 ex., 138.7 mm SL, Angkor ; MNHN 1974–27 View Materials , 24 ex., 29.1–144.2 mm SL, Tonlé Sap km 9 ; UMMZ 181204 View Materials , 1 ex., 188.2 mm SL, Mekong River N of Phnom Penh ; UMMZ 214411 View Materials , 1 ex., 98.7 mm SL, Mekong River and tributary from km 1393 to 1596 ; UMMZ 232130 View Materials , 1 ex., 137.9 mm SL, Kandal: Tonle Sap River 35 km upstream from Phnom Penh (11°52'N 104°47'E) GoogleMaps ; UMMZ 232634 View Materials , 1 ex., 115.2 mm SL, Kandal: Prek Mong Ya, just upstream from confluence with Bassac River at fishing lot 10 (10°57'N 105°7'E) GoogleMaps ; UMMZ 234410 View Materials , 1 ex., 139.4 mm SL, Kandal: Prek Ta Pov , 13 km S of Phnom Penh.
VIETNAM: MCZ 25589, 1 ex., 114.9 mm SL, Thu Dau Mot ; MNHN A2393 View Materials , 1 ex., 268.6 mm SL ; MNHN A2394 View Materials , 4 ex., 137.4–247.9 mm SL ; MNHN B283 View Materials , 3 ex., 198.1–232.4 mm SL ; MNHN B285 View Materials , 3 ex., 165.0– 263.5 mm SL, Cochinchina ; MNHN B281 View Materials , 2 ex., 86.1–106.8 mm SL, Saigon ; NRM 31079, 1 ex., 256.1 mm SL, Song Langna (probably near Saigon – Da Lat road) ; UMMZ 214410 View Materials , 1 ex., 187.1 mm SL, fish market at Vinh Long ; UMMZ 224622 View Materials , 1 ex., 38.5 mm SL, Chau Doc, Mekong-Bassac River , 3 km downstream from Chau Doc ; UMMZ 224645 View Materials , 1 ex., 104.5 mm SL, Chau Doc fish market ; UMMZ 224830 View Materials , 2 ex., 47.1–76.4 mm SL, Chau Doc, S end of Vinh Tuong Island, Bassac River .
CHAO PHRAYA: THAILAND: ANSP 59375 View Materials , 2 ex., 71.2–77.2 mm SL ; ANSP 59377 View Materials , 2 ex., 67.3–74.6 mm SL ; ANSP 59379 View Materials , 1 ex., 55.7 mm SL ; ANSP 59380 View Materials , 3 ex.; 125.0– 157.7 mm SL ; ANSP 59383 View Materials , 1 ex., 118.5 mm SL ; ANSP 59384 View Materials , 3 ex., 75.9–86.1 mm SL ; ANSP 59387 View Materials , 2 ex., 110.7–113.5 mm SL, Chieng Mai on Mae Nam Ping; ANSP 59389 View Materials , 2 ex., 100.4–127.1 mm SL, Bangkok , 48 km up the Mae Nam Chao Phraya ; ANSP 59467 View Materials , 1 ex., 34.9 mm SL, Silom Canal. , Bangkok ; ANSP 88047 View Materials , 1 ex., 96.2 mm SL, Me Poon ; ANSP 89483 View Materials , 1 ex., 87.5 mm SL, Phitsanulok ; ANSP 89533 View Materials , 13 ex., 90.0– 140.3 mm SL, Bangkok ; ANSP 141235 View Materials , 8 ex., 47.6–70.1 mm SL, Tachin ; BMNH 1898.11 .8.110, 1 ex., 224.0 mm SL, Ayutthaya . CAS 92860, 1 ex., 64.0 mm SL, Mae Nam Ping, about 41 km N of Chiang Mai ; CMK 4834 , 1 ex., 102.3 mm SL, Lampong province, Mae Nam Yom drainage, Huai Mae Phlung from Ban Pong to 17 km upstream, 18°42'N 99°58'E GoogleMaps ; CMK 4874 , 3 ex., 115.0– 159.6 mm SL, Ayutthaya province , market at Ayutthaya; CMK 5018 , 3 ex., 96.8–116.1 mm SL, market at Sukhothai ; CMK 5281 , 1 ex., 157.2 mm SL ; ZRC 46158, 1 ex., 202.7 mm SL, market at Nakhon Sawan ; FMNH 50847 View Materials , 1 ex., 89.1 mm SL, Nakhon Sawan province , Pak Nam Po; FMNH 50848 View Materials , 3 ex., 169.2–174.6 mm SL, Kamphaeng Phet province , Wong Pratart farm; MCZ 35534, 2 ex., 122.1–140.6 mm SL, Mae Nam Ping at Chieng Mai , 18°48'N 98°59'E; MNHN 1554 View Materials , 4 ex., 95.0– 136.4 mm SL, Me Nam River ; MNHN 1567 View Materials , 4 ex., 87.0– 127.3 mm SL, Bangkok ; UMMZ 186696 View Materials , 1 ex., 138.9 mm SL, Nakhon Sawan province , Chao Phraya River , floodwaters ca 20 km W of Nakhon Sawan ; UMMZ 186733 View Materials (81), 62.8–173.0 mm SL , UMMZ 195382 View Materials , 1 ex., 74.9 mm SL, Maharaj province, Chao Phraya River, floodwaters, 17.5 km N of Ayutthaya, Koke Tong Canal drainage ; UMMZ 186740 View Materials , 7 ex., 64.3–105.7 mm SL, Maharaj province, Koke Tong Canal , 17.5 km N of Ayutthaya ; USNM 103197 View Materials , 1 ex., 222.1 mm SL: Mae Nam Chao Phraya , Ayutthaya ; USNM 109566 View Materials , 14 ex., 33.9–106.2 mm SL, Mae Nam Ping at Chieng Mai ; USNM 109567 View Materials , 1 ex., 207.8 mm SL, Huey O, branch of Man Nam Nan ; USNM 109568 View Materials , 1 ex., 121.1 mm SL, Mae Nam Chem, tributary of Man Nam Ping ; UMMZ 214405 View Materials , 3 ex., 83.8–112.0 mm SL, creek running under Highway 1, 23 km S of Chiang Mai, Mae Nam Ping drainage ; USNM 118449 View Materials , 1 ex., 134.7 mm SL, Chieng Mai ; ZRC 29583–29596 View Materials , 14 ex., 177.6–214.2 mm SL , ZRC 29659–29665 View Materials , 7 ex., 132.3–158.0 mm SL ; ZRC 40305, 144.3 mm SL, Bangkok .
MAE KHLONG: THAILAND: CAS 79049, 3 ex., 102.8–131.1 mm SL, Huay Sangkalia , 7 km N of Sangklaburi on road to Chedi Sam Ong ; UMMZ 186715 View Materials , 5 ex., 108.2–121.5 mm SL, Rajburi province , Mae Khlong River, purchased in market at Rajburi ; UMMZ 186846 View Materials , 1 ex., 104.9 mm SL, Mae Nam Mae Khlong at Ban Pong, 2 km downstream ;
BANGPAKONG: THAILAND: CAS 92870, 1 ex., 63.7 mm SL, Bangpakong drainage, open channel in large swamp approx. 9 km SSW of Prachinburi town .
Diagnosis. — Hemibagrus spilopterus is distinguished from all members of the H. nemurus species group in having the interorbital space gently-curved (vs. flat). This is more clearly seen when the head is viewed dorsally. It further differs from H. capitulum in having the adpressed dorsal fin (excluding the filamentous extensions of the fin rays if present) not reaching (vs. reaching to or beyond) the adiposefin origin, from H. filamentus in having a larger dorsal to adipose distance (10.8–17.6% SL vs. 7.7–10.9), and from H. fortis in having the anteriormost branch of the dorsal-fin rays longer than other branches (vs. branches of dorsal-fin rays of almost equal length) giving the dorsal fin a distinctly jagged (vs. rounded) distal margin ( Fig. 20 View Fig ) and a shorter, deeper adipose fin (maximum height of adipose fin 1.7–3.2 times in the length of its base vs. 2.8–4.9). It is further distinguished from H. hoevenii in having the premaxillary tooth band not (vs. partially) exposed when the mouth is closed, rounded (vs. tapering) caudal-fin lobes, absence (vs. presence) of a broad and conspicuous dark margin around the caudal fin, a rounded (vs. triangular) anal fin, broader membranes of the dorsal fin which give it a rounded (vs. triangular) appearance, and filamentous extensions on the dorsal fin, if present, not reaching beyond middle of adipose-fin base (vs. very long filamentous extensions of the first two dorsal-fin rays, when present, that reach beyond the posterior base of the adipose fin), and from H. nemurus in having a larger eye in specimens larger than approx. 150 mm SL (12–18% HL vs. 9–11).
Description. — Biometric and meristic data as in Table 16. General description as for genus. Head depressed and broad, body moderately compressed; head width 56–73% HL. Interorbital space gently curved. Eye diameter 12–18% HL. Premaxillary tooth band not exposed when mouth is closed. Dorsal profile rising evenly but not steeply from tip of snout to origin of dorsal fin, then sloping gently ventrally from there to end of caudal peduncle. Ventral profile horizontal to origin of anal fin, then sloping dorsally to end of caudal peduncle. Adipose-fin base short, approximately as long as anal-fin base; maximum height 4.7–7.4% SL (1.7–3.2 times in length of adipose-fin base). Adipose-fin origin separated from base of last dorsal-fin ray by distance of 10.8–17.6% SL. Dorsal spine stout, with 7–11 serrations along posterior edge. Anteriormost branch of each dorsal-fin ray longer than other branches, imparting jagged distal margin to fin; membranes broad, imparting rounded appearance.Adpressed dorsal fin (excluding filamentous extensions of rays) not reaching adipose-fin origin; filamentous extensions not reaching beyond middle of adipose-fin base. Pectoral spine stout, with 15–26 large serrations along posterior edge. Anal fin rounded, origin slightly posterior to adipose origin. Caudal fin forked; upper and lower lobes rounded. Maxillary barbels reaching barbels reaching at least vertical through base of last anal-fin ray (209–299% HL). Maximum recorded size 500 mm SL ( Serov et al., 2006).
Colour. — Dorsal surface of head and body uniform brownish-grey; ventral surface of head and body dirty white; adipose fin brownish-grey, posterior half with distinct black spot; fin-rays and inter-radial membranes of all fins grey; dorsal surface of barbels brownish-grey, ventral surface dirty white.
Distribution. — Hemibagrus spilopterus is known from the river drainages in Indochina: Mekong [upstream to Chieng Sen], Dong Nai, Ban Pakong, Chao Phraya and Mae Khlong ( Fig. 26 View Fig ).
Habitat and biology. — Hemibagrus spilopterus is found in a wide variety of lotic and lentic habitats. The species feeds predominantly on small fishes and crustaceans (Beeckman & de Bont, 1985; Rainboth, 1996). In the Cambodian Mekong, it moves into the flooded forests to spawn, with the young being first seen in August; the fish migrate back towards the rivers in November–December ( Rainboth, 1996).
Remarks. — Ng & Rainboth (1999) considered the material identified as H. nemurus from central Indochina to belong to a distinct species, although they did not formally name it. They pointed out that the name Macrones luridus Tirant in Pétillot, 1911, possibly referring to this species, is a nomen nudum and published only once in a list of fishes (republished in Chevey, 1929) sent by Tirant to the Musée Guimet d’Histoire Naturelle, Lyon. In the same study, they described H. spilopterus , distinguishing it from congeners by the presence (vs. absence) of a distinct black spot on the adipose fin. Our investigation shows that the material identified by Ng & Rainboth as H. cf. nemurus and H. spilopterus are conspecific and that this material is distinct from H. nemurus s. str. and other Sundaic species in the H. nemurus species group. The only difference between the H. cf. nemurus of Ng & Rainboth and H. spilopterus is the intensity of the black spot on the adipose fin, with no other significant differences in morphology of the material being found in our study. As noted by Ng & Rainboth, a black spot is also present on the adipose fin in some individuals of H. capitulum (particularly those from central Sumatra), although they considered the spot in H. spilopterus to be more rounded and intensely coloured.. Given the variability displayed among individuals of this character, we conclude that it is not a useful diagnostic character.
The species identified as Hemibagrus sp. 1 by Nguy n (2005) is likely to be H. spilopterus . This species, for which material from the Mekong River drainage in northern Vietnam (Song Nam Rom) was examined, was described as being most similar to H. wyckii . It was described as differing from H. wyckii in having very long maxillary barbel reaching to the anal fin, short nasal barbel not reaching the orbital margin, adipose fin originating at vertical through anal-fin origin, length of adipose-fin and anal-fin bases equal and about two-fifths of the distance between the dorsal- and adiposefin bases. Although this information is scanty and could presumably refer to H. spilopterus or H. wyckioides (two other congeners likely to occur in the area), the illustration of this species (Nguy n, 2005: Fig. 184) is more similar to H. spilopterus on the account of the relatively short adipose-fin base. However, H. spilopterus has not (yet) been encountered in the Nam Ou drainage (of which the Song Nam Rom is a part of; MK pers. obs.). Despite this, we re-identify the Hemibagrus sp. 1 of Nguy n (2005) as H. spilopterus based on the information available and the illustration of this species.
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Genus |
Hemibagrus spilopterus Ng & Rainboth, 1999
| Ng, Heok Hee & Kottelat, Maurice 2013 |
Hemibagrus sp.
| Rainboth, W 2012: 72 |
Hemibagrus aff. nemurus
| Kottelat, M 2001: 122 |
Mystus planiceps
| Serov, D 1994: 24 |
| Serov, D 1994: 63 |
| Kottelat, M 1985: 270 |
| Desoutter, M 1975: 451 |
| Kuronuma, K 1961: 6 |
| Suvatti, N 1950: 297 |
| Fowler, H 1939: 40 |
| Fowler, H 1937: 148 |
| Suvatti, N 1936: 75 |
Mystus nemurus
| Lim, P & Lek, S 1996: 382 |
| Rainboth, W 1996: 143 |
| Serov, D 1994: 24 |
| Serov, D 1994: 62 |
| Roberts, T 1993: 33 |
| Kottelat, M 1985: 270 |
| Desoutter, M 1975: 448 |
| Orsi, J 1974: 161 |
| Kawamoto, N 1972: 27 |
| Kuronuma, K 1961: 6 |
| Suvatti, N 1950: 297 |
| Fowler, H 1937: 148 |
| Suvatti, N 1936: 73 |
| Fowler, H 1934: 95 |
Macrones planiceps
| Pellegrin, J 1937: 317 |
| Chevey, P 1932: 17 |
Macrones nemurus
| Chevey, P 1934: 201 |
| Chevey, P 1932: 17 |
| Chevey, P 1929: 172 |
| Hora, S 1924: 468 |
| Hora, S 1923: 171 |
| Petillot, L 1911: 164 |
Macrones luridus
| Chevey, P 1929: 172 |
| Petillot, L 1911: 164 |
Hemibagrus nemurus
| Serov, D 2006: 193 |
| Gaanantkul, K 2000: 34 |
| Baird, I. G & Inthaphaisy, P & Kisouvannalath, B 1999: 91 |
| Mo, T 1991: 132 |
| Sauvage, H 1883: 154 |
| Bleeker, P 1865: 34 |
| Bleeker, P 1865: 175 |
| Bleeker, P 1864: 353 |
Bagrus nemurus
| Bleeker, P 1859: 102 |