Zieglerodina remscheidensis ( Ziegler, 1960 )

Drygant, Daniel & Szaniawski, Hubert, 2012, Lochkovian conodonts from Podolia, Ukraine, and their stratigraphic significance, Acta Palaeontologica Polonica 57 (4), pp. 833-861 : 853

Zieglerodina remscheidensis ( Ziegler, 1960)

Figs. 8N, O , 12A–G.

For synonymy see Drygant (2010) and include:

1960 Spathognathodus remscheidensis n. sp.; Ziegler 1960: 194, pl. 13: 1, 2, 4, 5, 7 [non pl. 13: 8, 10, 14].

1972 Ozarkodina steinhornensis remscheidensis ( Ziegler, 1960) ; Mashkova1972: 83, pl. 2: 19–21, 23, 24 [non pl. 2: 22].

1984 Spathognathodus eosteinhornensis Walliser ; Drygant 1984: 123, pl. 13: 5–8, 11, 12, 15 [only].

1990 Ozarkodina remscheidensis remscheidensis ( Ziegler, 1960) ; Olivieri and Serpagli 1990: pl. 4: 10 [only].

Non 2005 Ozarkodina remscheidensis ( Ziegler, 1960) ; Barrick et al. 2005: 120, pl. 1: 1 (=?), 2 (=? Z. planilingua [ Murphy and Valenzuela−Ríos, 1999]), 9, 10 (=?); pl. 2, fig. 8 (= Zieglerodina prosoplatys Mawson, Talent, Molloy, and Simpson, 2003 ).

2010 Zieglerodina remscheidensis ( Ziegler, 1960) ; Drygant 2010: 49, pl. 1: 1–10, 12.

Material.—About 460 of the Pa and about 100 of other elements (Pb−Sc) from the Dzvenyhorod, Khudykivtsi, Mytkiv, and lower part of the Chortkiv Formation. Sections and samples: 33/2, 14D/1, 14/4, 4b, 5, 5a, 5v, 6, 7, 37/1, 1a, 1b, 2, 37A/1, 2, 37B/1, 16/1, 2, 3, 4, 5, 16B/1b, 15/1a, 17/3, 4v, 12/2, 18A/5, 45/ 170 m.

Remarks.—Typical Pa elements of the species (stratigraphically later forms from the upper part of Khudykivtsi Formation) have long (0.95–1.3 mm) and high (0.2–0.33 mm) blade with straight lower margin. A denticle above the basal cavity (cusp) comparatively high. Also 2–3 anterior denticles, especially the third, higher than ohers. Basal cavity closer to the posterior end. In specimens of the stratigraphically earlier forms (Silurian/Devonian boundary beds and lower part of the Khudykivtsi Formation) height of the anterior denticles and that above the basal cavity do not differ significantly from the rest ( Fig. 6 ).

It is worthy to mention that“ Spathognathodus eosteinhornensis Walliser, 1964 ” has been considered as an ancestor species of the “ Spathognathodus remscheidensis Ziegler, 1960 ” ( Walliser 1964). Material from Podolia shows that typical specimens of “ Spathognathodus eosteinhornensis ” do not occur above the Silurian/Devonian boundary. Its uppermost occurrence has been stated in the bed with Monograptus uniformis angustidens ( Fig. 3 ). That means Zieglerodina remscheidensis (sensu stricto) do not replace the discussed species but appears independently, already in Pridoli, similarly as in Estonia ( Viira 1999).

Stratigraphic and geographic range.—From the “ Ozarkodina ” eosteinhornensis Zone (uppermost Pridoli) to lower part of Caudicriodus transiens Zone in Podolia ( Figs. 3 , 6 ), Hüinghaus Beds of Rhenish Slate Mountains, Haragan Formation of Oklahoma, Road River Formation of Canadian Cordillera, Lochkovian of Barrandian Area ( Czech Republic) and Poland, Elmside Formation of New Sout Wales ( Australia).

Figures

Fig. 8. Some discrete elements of the unassigned taxa from the Ivanye (A–M) and Khudykivtsi (N, O) formations, all in lateral view. A, B, D. Pb elements. A. Ivanye Zolote−52/515 m, ZPAL C.20/7.28. B. Zalishchyky− 1/490 m, ZPAL C.20/6.5. D. Ivanye Zolote−52/515 m, ZPAL C.20/7.16. C, E, G. Sb elements. C. Ivanye Zolote−52/515 m, ZPAL C.20/7.17. E. Zalishchyky−1/490 m, ZPAL C.20/6.13. G. Ivanye Zolote−52/515 m, ZPAL C.20/7.29. F, J. M elements; Ivanye Zolote−52/515 m. F. ZPAL C.20/7.20. J. ZPAL C.20/7.14. H, L, M. Sa elements (probably genus Pandorinellina). H. Ivanye Zolote−52/515 m, ZPAL C.20/7.19. L. Zalishchyky−1/490 m, ZPAL C.20/6.11. M. Ivanye Zolote−52/515 m, ZPAL C.20/7.24. I, K. Sc elements. I. Zalishchyky−1/490 m, ZPAL C.20/6.10. K. Ivanye Zolote−52/515 m, ZPAL C.20/7.26. N, O. Zieglerodina remscheidensis (Ziegler, 1960); Dnistrove−14/1.3 m. N. Sb element, ZPAL C.20/ 1.15. O. Sc element, ZPAL C.20/1.14.

Fig. 6. Probable phyletic relationship within Zieglerodina and Pandorinellina? based on the Podolian collection. D., Devonian; S., Silurian; Fm., Formation. For legend see Fig. 3.

Fig. 3. Stratigraphic range of conodonts in the Lochkovian sequence of Podolia.A. Lower part. B. Upper part (see next page). D., Devonian; S., Silurian.