Caudicriodus woschmidti ( Ziegler, 1960 )

Drygant, Daniel & Szaniawski, Hubert, 2012, Lochkovian conodonts from Podolia, Ukraine, and their stratigraphic significance, Acta Palaeontologica Polonica 57 (4), pp. 833-861 : 846

Caudicriodus woschmidti ( Ziegler, 1960)

Figs. 9B , 10C, D.

1960 Icriodus woschmidti n. sp.; Ziegler 1960: 185, pl. 15: 16, 17, 18, 20, 21, 22.

1976 Caudicriodus woschmidti ( Ziegler 1960) ; Bultynck 1976: 21 [partim], text−fig. 1, 3: 1–2, 4: 1–4 [?non fig. 3: 3, 4: 5 =? Caudicriodus hesperius ( Klapper and Murphy, 1975) ].

?1980 Icriodus woschmidti ssp.; Schönlaub 1980b: pl. 3: 19.

2003 Caudicriodus woschmidti woschmidti ( Ziegler 1960) ; Bultynck 2003: pl. 1: 1–3.

2010 Caudicriodus woschmidti ( Ziegler 1960) ; Drygant 2010: 57, pl. 2: 3, 6–13.

For more extensive synonymy see Drygant (2010).

Material.— 20 specimens from lower part of the Khudykivtsi Formation. Sections and samples: 14/4a, 7, 8, 14 V/4 , 6 , 104 /1, 105/1.

Remarks.—Morphology of the Podolian Pa elements, especialy the outline of basal cavity and denticulation of the spindle, as well as the outline of spur and postero: lateral process ( Fig. 5 ) agree in all details with the holotype ( Ziegler 1960: pl. 15: 16a–c). The platform is slightly sigmoidal in upper view, comparatively narrow, gradually widening from the anterior end to the first transverse ridge. Spindle elongated (0.9–1.2 mm), with maximum five transverse ridges and anterior immaturate denticle. Its denticulation identical to that in Caudicriodus hesperius but the distance between transverse ridges is slightly larger.

Postero−lateral process short or a little elongated, usually weakly curved, with pointed end, bears a single ridge of denticles. Inner and outer lobes undenticulated. Basal cavity expanded behind the second or third transverse ridge. The primary cusp higher than the secondary one. The angle between the median longitudinal row and postero−lateral process bigger than 130 °. In very small juvenile specimens (with 1–3 transverse ridges) the postero−lateral process and spur are lacking ( Fig. 9B ). Variations in morphology of adult specimens from Podolia has been illustrated in Drygant (2010: pl. 2: 6–13).

The specimens assigned to Icriodus woschmidti , known from Celtiberia ( Carls 1975: pl. 2: 19, 21) and Tennessee ( Broadhead and McComb 1983: fig. 3H–J) differ from the Podolian specimens by the less expanded basal cavity ( Fig. 5 ).

Stratigraphic and geographic range.—Lower part of Caudicriodus hesperius Zone in Podolia ( Fig. 3 ), Rhenish Slate Mountains ( Germany), Iberian Chains, central Pyrenees and Guadarrama Mountains ( Spain), Sardinia, Marocco, New South Wales and Victoria ( Australia), Northwestern Canada, Appalachians.

Figures

Fig. 9. Ontogenetic development and morphological comparison of Pa elements of Podolian Caudicriodus species.

Fig. 5. A. Probable phyletic relationships within Caudicriodus, based on the evidence of the Podolian collection. B. Similar forms in other regions. D., Devonian; S., Silurian; Fm., Formation. For legend see Fig. 3.

Fig. 3. Stratigraphic range of conodonts in the Lochkovian sequence of Podolia.A. Lower part. B. Upper part (see next page). D., Devonian; S., Silurian.