Calosota panamaensis, Gary A. P., Gibson, 2010

Gary A. P., Gibson, 2010, Calosota Curtis (Hymenoptera, Chalcidoidea, Eupelmidae) - review of the New World and European fauna including revision of species from the West Indies and Central and North America, ZooKeys 55, pp. 1-75 : 45-46

Calosota panamaensis sp. n. Figs 2140

Etymology.

Based on the country of origin of the only known specimen.

Type material.

HOLOTYPE ♀ (AEIC). Panama, nr Hato Del Volcan, 4700 feet, Jul. 1982, B. Gill; CNC Photo 2009-37; Holotype Calosota panamaensis Gibson.

Description.

FEMALE (Fig. 40). HOLOTYPE: length about 3.5 mm (mesonotum arched). Color. Head with frontovertex, parascrobal region and paraclypeal region dark brown with obscure reddish-violaceous sheen under some angles of light, scrobes and interantennal region to ventral margin of toruli dark or variably extensively bright violaceous-purple under some angles of light, and most of clypeal region and transverse band below toruli bluish-green. Maxillary and labial palpi dark. Antenna with scape brownish-yellow over almost basal quarter, otherwise dark brown with bluish-green lusters under some angles of light, pedicel with similar bluish-green luster, but flagellum dark brown. Tegula yellow. Mesonotum (Fig. 21) bright green with variable reddish-coppery reflections under different angles of light (frenal area concealed by dorsellum). Acropleuron greenish anteriorly similar to mesonotum, but grading to bluish-violaceous posteriorly. Legs (Fig. 40) with front leg extensively light brownish-yellow with knee and apex of tibia more distinctly yellow, but tibia subbasally and dorsally except for apex dark brown with greenish luster under some angles of light; middle leg yellowish beyond coxa except dorsal surface of tibia somewhat darker brownish-yellow at least subbasally; hind leg mostly yellowish beyond coxa, but femur more brownish, particularly ventrally, over about basal half and dorsal surface of tibia slightly brownish subapically. Fore wing hyaline; setae uniformly brown. Gaster (Fig. 40) mostly brown dorsally, but apex of penultimate tergum, syntergum more obscurely, and terga laterally bluish-green to violaceous.

Structure. Head in dorsal view about 1.9 × as wide as long, with IOD about 0.4 × head width; IOD: MPOD: OOL: POL: LOL = 55: 13: 8: 16: 10; in frontal view about 1.25 × as wide as high, with about middle of torulus in line with lower orbits; malar space about 0.47 × eye height. Head with frontovertex and parascrobal region meshlike coriaceous to about level of interantennal region, the sculpture more obscurely tapered ventrally between smooth and shiny scrobes and interantennal region dorsally, though interantennal region becoming increasingly more conspicuously coriaceous ventrally; clypeal region microcoriaceous and paraclypeal region mostly obliquely coriaceous-alutaceous, though paraclypeal region and lower parascrobal region smooth and shiny along inner orbit. Head with brownish setae except for bare scrobal depression. Antenna (Fig. 40) with flagellum clavate; length of flagellum + pedicel about 1.5 × as long as width of head; scape: pedicel: fu1-fu8: clava = 61(12): 23(10): 11(7), 16(8), 17(9), 18(9), 18(9), 15(9), 15(10), 15(13): 52(21). Mesoscutum (Fig. 21) strongly reticulate, the reticulations obviously larger medially than on lateral lobes but even largest reticulations very slightly concave rather than distinctly flat-bottomed, and with inconspicuous brownish setae; notaulus extending from spiracle as curved furrow on inclined anterior surface, without distinct anteroadmedian line and neither line extending posteriorly on dorsal surface; parapsidal line a more distinct microsculptured region. Axilla elongate-slender, separated by about 4 × own width. Scutellum (Fig. 21) low convex, about 1.25 × as long as wide; meshlike reticulate with concave reticulations similar in size to that on lateral lobes; with inconspicuous brownish setae. Mesopleuron with exposed, setose lower mesepimeron (cf. Fig. 53); acropleuron finely sculptured, obliquely coriaceous-alutaceous anterior to oblique microsculptured region and very finely longitudinally coriaceous-alutaceous to coriaceous-aciculate posteriorly. Fore wing with cc: mv: stv: pmv = 36: 34: 10: 20; basal cell entirely setose; cubital area extensively bare but setose anteriorly along mediocubital fold and closed by setae along posterior margin over about apical half; disc uniformly setose except for slightly developed bare band at juncture of basal and mediocubital folds. Metacoxa setose along dorsal and ventral margins and over about basal half of outer surface. Propodeum with callus setose to posterior margin; bare anteriorly between spiracle and foramen. Gaster (Fig. 40) about 2.3 × as long as mesosoma; with brownish setae dorsally but more whitish setae laterally; posterior margin of penultimate tergum not extending to level of cerci, the precercal portion equal to about one-third distance between cerci; syntergum with medial length measured to apex of penultimate tergum about 1.6 × transcercal width and about 0.9 × as long as penultimate tergum.

MALE. Unknown.

Biology.

Unknown.

Distribution.

Known only from Panama (Map 5).

Recognition.

Among the four species having a coriaceous frontovertex and extensively smooth and shiny scrobal depression (see under Calosota albipalpus ), Calosota panamaensis is most similar to Calosota speculifrons (see further under latter species). It is quite possible that females collected in the future may have more extensively and conspicuously brown legs than the holotype. The short precercal portion of the syntergum of the holotype does not appear to be an artifact of preservation because it is as strongly and uniformly sculptured as the remainder of the syntergum.