Senecio macedonicus subsp. macedonicus

Calvo, Joel, Álvarez, Inés & Aedo, Carlos, 2015, Systematics of Senecio section Crociseris (Compositae, Senecioneae), Phytotaxa 211 (1), pp. 448-450 : 448-450

12a. Senecio macedonicus subsp. macedonicus

Senecio macedonicus var. glaber Regel (1941: 242) . TYPE: Greece, Thessalonica , gipfel des Chortiatis, [40º35'N 23º06'E], 8 July 1939, C.A. Regel s.n. (lectotype, designated here, G-162759!; isolectotypes G-162760!, G-162761!).

Senecio bracteatus Boissier & Orphanides in Boissier (1856: 36). Senecio macedonicus var. brachyphyllus Boissier (1875: 404) . Senecio macedonicus var. bracteatus (Boissier & Orphanides) Nyman (1879: 354) . TYPE: Greece, Peloponnese, montis Ziriae supra Trikala, [37º55'N 22º24'E], June 1852, T.G. Orphanides 109 (lectotype, designated here, G-150323 image!; isolectotypes BR-505719!, FI!, LE!, S!, UPS!, WU!).

Senecio tmoleus var. longifolius Boissier & Heldreich , nom. nud., in sched. ( McNeill et al. 2012, ICN Art. 38.1) (G-162680!, G-162679!, LE!).

Basal leaves 9–21 cm long, 2.9–6.6 cm wide (ratio basal leaf width / basal leaf length = 0.17–0.47), entire to slightly dentate (teeth 1.3–2.8 mm deep), with a petiole 5–18 cm long. Middle cauline leaves 7.2–18 cm long, 1.2–4.1 cm wide (ratio middle leaf width / middle leaf length = 0.12–0.30), sessile, sometimes attenuated into a petiole up to 8 cm long, entire to slightly dentate; upper cauline leaves 2–7 cm long, 0.6–2.8 cm wide (ratio upper leaf width / upper leaf length = 0.23–0.52), sessile, entire. Chromosome number: 2 n =40 ( Kuzmanov & Georgieva 1983: 665, Constantinidis et al. 1997: 126). Figure 18 .

Iconography: — Stojanov & Stefanoff (1948: 1172, n.º 1223); Lafranchis & Sfikas (2009: 216, 217, as photo).

Distribution and habitat: — Bulgaria, Greece, Macedonia *; grassy subalpine meadows, rocky outcrops, woods of Quercus , Pinus , Fagus , on calcareous soils, rarely siliceous; elevations of 900–1800 m ( Figure 19 ).

Phenology: — Flowering from June to August.

Discussion: — Euro+Med PlantBase (2006+, http://ww2.bgbm.org/EuroPlusMed/PTaxonDetailOccurrence.asp?NameId=7000832&PTRefFk=7000000, accessed 18 March 2014) also recorded this taxon from Macedonia, although we did not find any material from this region. Therefore this country appears with an asterisk in the distribution of this species.

Selected specimens examined. BULGARIA. Blagoevgrad: Alibotuschgebirge, in bul. N-O Mazedonien, 41º24’N, 23º36’E, 20 July 1936, A. K . Drenowski s.n. ( M) . Kyustendil: in subalpinis mt. Rila , 42º10’N, 23º15’E, July 1903, O . Bierbach s.n. ( LE) . Plovdiv: in submontanis m. Rhodope supra Staminaka, 41º59’N, 24º51’E, 10 July 1898, V . Střibrny s.n. ( B, WU); in submontanis ad Staminaka , 41º59’N, 24º51’E, July 1894, V . Střibrny s.n. ( G, K, WU) . Smolyan: in m. Rhodope prope Leskovo, 41º36’N, 25º2’E, 5 July 1896, V . Střibrny s.n. ( B, F, K, M, S); in m. Rhodope prope Leskovo , 41º36’N, 25º2’E, 4 Aug 1895, V . Střibrny s.n. ( B, L, LE, S, US WU) . Sofia: Vitosha- Sofia, 42º34’N, 23º16’E, 1937, R . Baschant s.n. ( B); m. Cepan, prope vicum Dragoman , 42º57’N, 22º59’E, 29 June 1930, N . Stojanoff & T . Georgieff s.n. ( LE, S) . GREECE. Central Greece: in m. Parnassi reg. sylvatica loco dicto Drako, 38º32’N, 22º33’E, G . Guicciardi 192 ( FI, G); Peloponnes, Achaia, Kallifoni , 39º8’N, 21º42’E, 28 June 1991, F . Krendl & W . Burri s.n. ( E) . East Macedonia and Thrace: Dramas, SE slopes of mt. Orvilos (Ali Botus), WNW of the village of Katafyton, 41º21’N, 23º38’E, 25 July 1977, A . Strid & E . Georgiadou 13319 ( C, G); Dramas, Mt. Menikion ( Boz Dagh of Serre ), 14 km from village Mikropolis , 41º12’N, 23º44’E, 5 July 1981, A . Strid et al. 18602 ( B, G) . Peloponnese: Achaia, m. Kyllenes faucibus Phlamburitza , 37º53’N, 22º26’E, 28 June 1887, T . Heldreich 939 ( BR, F, G, K, LE, M, S, UPS, WU); in monte Malevo Laconiae prope Camalus , 37º15’N, 22º34’E, July 1850, T. G . Orphanides 1139 ( G, K, UPS, WU) . Thessaly: in m. Olympo Thessaliae, 40º1’N, 22º18’E, July 1857, T. G . Orphanides s.n. ( E, WU) . West Greece: Peloponnes, Achaia, Chelmos S-Teil, Fraga, Gipfelbereich, 37º52’N, 22º0’E, 14 July 1991, F . Krendl & W . Burri s.n. ( E) .

SYSTEMATICS OF SENECIO SECTION CROCISERIS

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CALVO ET AL.

Figures

FIGURE 18. A–J. Senecio macedonicus Grisebach subsp. macedonicus—a, b. Habit (drawn from Univ. Copenhagen Excurs. 75, C).—c. Capitulum (drawn from Heldreich 939, S).—d. Involucral bract (drawn from drawn from Heldreich 939, S).—e. Supplementary bract (drawn from drawn from Heldreich 939, S).—f. Ligulate floret (drawn from drawn from Heldreich 939, S).—g. Tubular floret (drawn from Heldreich 939, S).—h. Stamens (drawn from Heldreich 939, S).—i. Achene (drawn from Greuter 15973, B).—j. Indumentum of achene (drawn from Greuter 15973, B). K–L. Senecio castagneanus Candolle (drawn from Nemetz s.n. 27 June 1897, WU)—k. Basal leaf.—l. Synflorescence.

FIGURE 19. Distribution map for Senecio macedonicus Grisebach subsp. macedonicus (▲), S. macedonicus subsp. barckhausiifolius (Boissier & Heldreich) Calvo (∆), and S. eubaeus Boissier & Heldreich (●).