Homotropus signatus ( Gravenhorst 1829, Bassus

Klopfstein, Seraina, 2014, Revision of the Western Palaearctic Diplazontinae (Hymenoptera, Ichneumonidae), Zootaxa 3801 (1), pp. 1-143 : 72-73

Homotropus signatus ( Gravenhorst 1829, Bassus )

Tryphon compressiventris Cresson 1868

Homotropus hygrobius Thomson 1890

Homotropus bifoveolatus Kriechbaumer 1894

Zootrephes montanus Davis 1895

Diagnosis. Fore wing length 3.5–5.1 mm. Antenna in females with 19 flagellomeres, apical flagellomeres with multiporous plate sensilla also present ventrally; in males with 19 to 21 flagellomeres and with narrow, long tyloids on flagellomeres 7/8 to 14/15. Face strongly coriaceous, punctures disappearing against the background sculpture. Clypeus strongly excised at least in females, with lobes somewhat angulate at outer corners. Mesopleuron coriaceous at least on lower half, weakly punctate on lower half and along anterior margin. Mesoscutum finely coriaceous, with weak and sparse punctures. Hind coxa with hind surface coriaceous and matt. Fore wing areolet open or closed, if closed then vein 3rs-m usually unpigmented. Propodeum with pleural and lateral longitudinal carinae present, and usually also petiolar area surrounded by carinae or at least indicated by strong rugae (sometimes almost completely carinate), coriaceous and strongly rugose on hind surface. Female metasoma strongly compressed posterior to third segment. First tergite with median dorsal carinae often surpassing middle of tergite, with some longitudinal wrinkles laterally, 1.0–1.5 times as long as wide; second tergite basally with few short longitudinal wrinkles, 0.65–0.8 times as long as wide in females, 0.8–1.2 in males, second tergite 0.8–1.0 times as long as first tergite; spiracle of third tergite on dorsal or lateral part, above or behind lateral fold.

Colouration of females. Antenna bright orange or brown. Head and mesosoma black, with yellow on central face patch which is separated from yellow on clypeus, yellow on mouthparts, hind corner of pronotum, tegula, subtegular ridge, small to large shoulder mark, mesepimeron; scutellum black. Legs orange, all coxae either orange or black, fore and mid coxae often largely yellow apically, femora orange, hind tibia usually entirely orange, apex and hind tarsus usually not dark. Metasoma black, tergites 2 to 4 or 5 all or mainly orange.

Colouration of males. As in females but additionally with yellow over entire face, scape and pedicel ventrally, antenna usually orange below; mesosoma usually without additional yellow coloration, fore and mid coxae entirely yellow.

Material examined. 11 syntypes of Homotropus hygrobius Thomson : Sweden, Skåne. 5♂, 6♀, at ZIL. Paralectotype of Homotropus bifoveolatus Kriechbaumer : Spain, Mallorca,leg. Maragues. 1♂, at ZSM.

New for Greece: Col de Metsovo/Metsovo/Pindos, leg. J.Aubert, 6.V.1955. 2♂, 1♀, at MZL.

Finland (2), Germany (3), Greece (3), Hungary (16), Iceland (14), Netherlands (1), Russia (3), Spain (1), Sweden (>100), Switzerland (30), Turkey (1), United Kingdom (50).

Distribution. Holarctic.

Figures. Face ♀ ( Fig. 7 H), propodeum ( Fig. 9 D), tip of metasoma and ovipositor sheaths ♀ ( Fig. 15 I), epicnemical carina ( Fig. 16A ), habitus ( Fig. 32 B).

Figures

FIGURE 7. Macro- and microsculpture of diplazontine metasomas and faces. A – D: tergites 1 – 3 of the metasoma. A. Diplazon scutatorius, smooth and shining between the strong and dense punctures. B. Diplazon tetragonus, with partly rugulous sculputure basally on tergite 2, and coriaceous sculpture on tergite 3, both tergites bearing weak punctures on a smooth background behind the strong transverse impressions. C. Tymmophorus obscuripes, with rugose sculpture on tergites 1 and 2, becoming more coriaceous on tergite 3. D. Woldstedtius holarcticus, with coriaceous tergites and striae basally on tergite 2. E – H: faces showing sculpture and shape of clypeus. E. Sussaba flavipes, with a face that is coriaceous only on the middle elevation. F. Woldstedtius holarcticus. G. Diplazon scutatorius. H. Homotropus signatus. F. – H. all with strongly coriaceous and matt sculpture. Scale bars represent 500 µm.

FIGURE 9. Propodeum of diplazontine species. A. propodeum of Diplazon scutatorius with areas and carinae as termed by Townes (1965) marked. ba = basal area, pa = petiolar area (fused with areola, see 9 C), la 1 = first lateral area, la 2 = second lateral area, pla 12 = fused first and second pleural areas, pla 3 = third pleural area, mp = mesopleuron, 1 = transverse carina, 2 = median longitudinal carina, 3 = lateral longitudinal carina, 4 = pleural carina. B. same as A, but viewed laterally. C. Propodeum of Tymmophorus obscuripes with full carination and even an indication of the areola between the basal and petiolar areas. D. Propodeum of Homotropus signatus with carinae partly reduced but with the posterior ends of median and lateral longitudinal carinae present and transverse carina indicated by rugae. E. Propodeum of Woldstedtius holarcticus with carinae completely reduced. Scale bars represent 500 µm.

FIGURE 32. Habitus of Diplazontinae species. A. Homotropus pictus. B. Homotropus signatus. C. Homotropus sundevalli. D. Homotropus vitreus. E. Phthorima compressa. F. Phthorima picta.

FIGURE 15. A. Syrphophilus tricinctorius ♀, metasoma. B. Homotropus crassicornis ♀, metasoma. C. Bioblapsis polita ♀, propodeum and first tergite. D. Homotropus crassicornis ♀, propodeum and first tergite. E. Bioblapsis polita ♀, metasoma. F. Bioblapsis polita ♀, antenna. G. Woldstedtius citropectoralis ♀, apical metasomal segments and ovipositor sheaths. H. Diplazon neoalpinus ♀, apical metasomal segments and ovipositor sheaths. I. Homotropus signatus ♀, apical metasomal segments and ovipositor sheaths.

FIGURE 16. A. Homotropus signatus ♀, epicnemical carina. B. Syrphoctonus tarsatorius ♀, epicnemical carina. C. Homotropus elegans ♀, clypeus. D. Syrphoctonus tarsatorius ♀, clypeus. E. Enizemum ornatum ♀, clypeus.