Homotropus frontorius ( Thunberg 1824, Ichneumon

Klopfstein, Seraina, 2014, Revision of the Western Palaearctic Diplazontinae (Hymenoptera, Ichneumonidae), Zootaxa 3801 (1), pp. 1-143 : 66

Homotropus frontorius ( Thunberg 1824, Ichneumon ) (comb. nov.)

Homocidus subopacus Stelfox 1941 (syn. nov.)

Homotropus quadrangularis Dasch 1964a (syn. nov.)

Diagnosis. Fore wing length 4.2–5.6 mm. Antenna in females with 19 to 20 flagellomeres, apical flagellomeres with multiporous plate sensillae also present ventrally; in males with 20 to 21 flagellomeres and with narrow, long tyloids on flagellomeres 7 to 14–16. Face strongly coriaceous, punctures usually disappearing against the background sculpture. Clypeus weakly to strongly excised, with lobes somewhat angulate at outer corners. Mesopleuron coriaceous at least on lower half, punctate on lower half and along anterior margin. Mesoscutum coriaceous, especially on median part, densely punctate. Hind coxa with hind surface coriaceous and matt. Fore wing areolet closed, vein 3rs-m often unpigmented. Propodeum with only pleural carina and apical parts of longitudinal carinae present, coriaceous, rather smooth and rugose on petiolar area. Female metasoma gradually tapered posterior to third segment. First tergite with median dorsal carinae indicated at most basally, never surpassing middle of tergite, without longitudinal wrinkles, 0.85–1.3 times as long as wide; second tergite basally with few longitudinal wrinkles, 0.65–0.8 times as long as wide, second tergite 0.9–1.1 times length of first tergite; spiracle of third tergite on dorsal part, above lateral fold.

Colouration of females. Antenna black. Head and mesosoma black, with yellow on central face patch usually separated from yellow on clypeus, yellow on mouthparts, hind corner of pronotum, tegula, subtegular ridge, shoulder mark, at least upper mesepimeron, scutellum laterally and apically; usually marked with orange on hind corner of mesopleuron and on metapleuron. Legs orange, all coxae orange, femora orange, hind tibia white with a dark apex and base and sometimes subbasal marking, hind tarsus dark. Metasoma black.

Colouration of males. As in females but additionally with yellow over entire face, scape and pedicel ventrally, propleuron, mesosternum and a stripe on lower mesopleuron, fore and mid coxae, and often basal spots on tergites 3 and sometimes 4. Meso- and metapleurae usually not marked with orange.

Material examined. Holotype of Ichneumon frontorius Thunberg : Sweden, Dalarna , leg. Dr. Blom. 1♂, at UU . Holotype of Homocidus subopacus Stelfox : Ireland, Dublin, Gelansmole, 22.V.1938, 1♀, at USNM . Holotype of Homotropus quadrangularis Dasch , examined by E. Diller (personal communication).

New for Germany: München, Lochhausen, leg. E.Diller, 8.VI.1963. 1♀, at ZSM. New for Norway: Jotunheimen, Giendesheim, 1000–1500m, leg. J.E.&R.B.Benson, 7.VII.1966. 1♀, at BMNH . New for Sweden: Värmland, Munkfors kommun, Ransäter, Ransbergs herrgard. Old mixed deciduous ofrest in stream ravine, N59°47'25.59", E13°24'54.61", leg. SMTP, 18.VI.–27.VI.2005. 2♀, at NRM . New for Switzerland: Grisons, Sur , Clavenia 1987m, N46°32.005, E8°50.135, leg. S.Klopfstein, 8.VI.2007. 1♀, at NMBE.

Finland (2), Germany (2), Norway (1), Sweden (12), Switzerland (4), United Kingdom (6), USA (1).

Distribution. Holarctic.

Figures. Hind tibia ♀ ( Fig. 14 D), metasoma ♀ ( Fig. 20 B), habitus ( Fig. 31 C).

Figures

FIGURE 20. Homotropus characters mentioned in the species key. A. Homotropus melanogaster ♀, metasoma. B. Homotropus frontorius ♀, metasoma. C. Homotropus strigator ♀, propodeum. D. Homotropus pictus ♀, propodeum. E. Homotropus collinus ♀, clypeus. F. Homotropus elegans ♀, clypeus. G. Homotropus longiventris ♀, metasoma. H. Homotropus pictus ♀, metasoma. I. Homotropus venustus ♂, antenna showing tyloids. Scale bars represent 500 µm.

FIGURE 31. Habitus of Homotropus species. A. Homotropus dimidiatus. B. Homotropus elegans. C. Homotropus frontorius. D. Homotropus longiventris. E. Homotropus nigritarsus. F. Homotropus pallipes.

FIGURE 14. A. Phthorima compressa ♀, metasoma. B. Enizemum ornatum ♀, hind tibia. C. Enizemum ornatum ♂, hind tibia. D. Homotropus frontorius ♀, hind tibia. E. Enizemum ornatum ♀, first tergite. F. Enizemum nigricorne ♀, first tergite. G. Homotropus elegans ♀, first tergite. H. Homotropus sundevalli ♀, first tergite. I. Enizemum ornatum ♀, clypeus. J. Homotropus elegans ♀, clypeus. K. Fossatyloides gracilentus ♂, antenna. L. Fossatyloides gracilentus ♀, propodeum. M. Homotropus pictus ♀, propodeum. N. Homotropus crassicornis ♀, propodeum.