Hydraena particeps

Perkins, Philip D., 2011, New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae), Zootaxa 3074, pp. 1-198 : 45

Particeps Complex

Perkins (1980: 36)

Nine species were placed in the particeps Subgroup by Perkins (1980). That grouping now appears to be composite, and will require additional study to more accurately adduce relationships. As a first step, H. orcula is removed and placed in the Orcula Complex.