Eupines (Byraxis) crassicornides Newton, 2017

Shen, Jia-Wei & Leschen, Richard A. B., 2020, Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae), Zootaxa 4777 (1), pp. 1-84 : 22

Eupines (Byraxis) crassicornides Newton, 2017

Fig. 12 ; map Fig. 50

Bryaxis crassicornis Broun, 1880: 129 .

Eupines (Byraxis) crassicornis : combination and subgeneric assignment by Raffray 1904: 207. Raffray 1911: 82, 1924: 157; Hudson 1923: 364, 1934: 182; Nomura and Leschen 2006: 254; Maddison 2010: 417.

Eupines (Byraxis) crassicornides Newton, 2017: 13 . New name for Bryaxis crassicornis Broun, 1880: 129 , preoccupied by Bryaxis crassicornis Motschulsky, 1851: 492 . Shen and Leschen 2019: 262 .

Type locality: [CL] Tairua.

Type depository: BMNH HT ♂ .

Diagnosis: Head with vertexal foveae asetose and small; antennae 10-segmented with 4-segmented antennal club; A4, A5, A6, A7 and A10 unmodified, A7 slightly asymmetrical, A8 elongate, about 3 × longer than A7, truncate at base, A9 longest, about twice as long as A8, concave at outer margin, concavity surrounded by strongly extended laminate margin, A10 conical and truncate at base.Abdominal ventrite 2 with pair of small discal protuberances, inserted with slightly curved apical setae; ventrite 6 with pair of short median struts, about one-fifth of ventral length. Protrochanters with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified.

Redescription: Body length 1.40–1.52 mm; body colour reddish brown to dark brown with lighter coloured legs. Head slightly wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes well developed, with about 33 facets; vertexal foveae asetose and small; antennal tubercles moderately developed; lacking postantennal notches; postantennal cavities large in dorsal view, about as long as antennal tubercles in dorsal view; antennae 10-segmented with 4-segmented antennal club; A4, A5, A6, A7 and A10 unmodified, A7 slightly asymmetrical, A8 elongate, about 3 × longer than A7, truncate at base, A9 longest, about twice as long as A8, concave at outer margin, concavity surrounded by strongly extended laminate margin, A10 conical and truncate at base. Pronotum as long as wide, about as wide as head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest at apical two-thirds; humeral calli and sutural stria vaguely visible. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with slightly curved apical setae; ventrite 6 with pair of short median struts, about one-fifth of ventral length. Legs with protrochanters modified with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.23 mm long, with parameres elongate and sinuate, with three pairs of short apical setae, two pairs of long subapical setae and one pair of long median setae; median lobe with two long sclerites, about as wide as parameres in dorsal view, both curved outwards at apex, left one more sinuate than right one.

Comments: This is the only species known with A8 conspicuously modified ( Figs. 12 B–D).

Distribution: North Island. ND, AK, CL.

Additional material examined: 120 ♂♂, ND: 27, Mokohinau Is., Trig I., 27/Feb/1978, G. Kuschel, sifted litter, 78/107; 12, Mokohinau Is., Fanal I., 01/Mar/1978, G. Kuschel, sifted litter and ground plants, 78/115; 1, Mokohinau Is., Burgess I., 06/Jan/1984, P.D McDonald, litter, 84/5; 2, Hen & Chickens Is., Lady Alice I., 01/Jan/1982, R. Hay, litter, 82/4; 12, Poor Knights Is., Aorangi Crater Bay and ridge to Oneho Hill, 17/Nov/1981, J.C. Watt, sifted litter 81/141; AK: 54, Noises Is., Scott I., 25/Feb/1979, J.M. Cleary, M.F. Tocker, litter 79/31; CL: 2, Mecury Is., Korapuki I., 16/Dec/1987, G. Hall, sifted litter, 89/23; 10, AMNZ, Great Barrier Is., Little Windy Hill, 220 m, 11/xii/2001 – 18/i/2002, P. Sutton, Forest edge, Malaise trap. Associated females (201 ♀♀): ND: 73, Mokohinau Is., Trig I., 27/Feb/1978, G. Kuschel, sifted litter, 78/107; 16, Mokohinau Is., Burgess I., 06/Jan/1984, P.D McDonald, litter, 84/5; 26, Mokohinau Is., Fanal I., 01/Mar/1978, G. Kuschel, sifted litter and ground plants, 78/115; 11, Hen & Chickens Is., Lady Alice I., 01/Jan/1982, R. Hay litter 82/4; 7, Poor Knights Is. Aorangi Crater Bay and ridge to Oneho Hill, 17/Nov/1981, J.C. Watt, sifted litter, 81/141; AK: 71, Noises Is., Scott I., 25/Feb/1979, J.M. Cleary, M.F. Tocker, litter 79/31; CL: 7, Mecury Is., Korapuki I., 16/Dec/1987, G. Hall, sifted litter, 89/23.

Figures

FIGURE 12. Diagnostic characters of E. (B.) crassicornides Newton, 2017. A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Same, in ventral view. E) Protrochanter. F) Ventrite 2. G) Ventrite 6. H) Aedeagus, in dorsal view. I) Same, in lateral view. J) Same, in ventral view. Scale bars: A = 1 mm, B–F = 0.2 mm, G–J = 0.1 mm.

FIGURE 50. Geographic distributions of E. (B.) crassicornides Newton, 2017, E. (B.) decens (Broun, 1893), E. (B.) dispar (Sharp, 1874), E. (B.) dugdalei sp. n., E. (B.) fraudulenta (Broun, 1886) and E. (B.) gigas sp. n. in New Zealand.