Austrolophrum guichardi, Steel, 1949 b: 237

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.1

persistent identifier

https://treatment.plazi.org/id/BF684949-42A8-B58D-00DC-2DB4FD2B7C8E

treatment provided by

Felipe

scientific name

Austrolophrum guichardi
status

 

guichardi Steel, 1949b: 237 View in CoL ( Austrolophrum ; variety of cribriceps ; Type locality:

Mt. Martha, Victoria).

adelaidae Blackburn, 1892: 23 ( Amphichroum ; cited as Amphicroum ; Type locality:

S. Australia; near Adelaide).

— Bernhauer and Schubert, 1910: 70 ( Amphichroum ; catalog).

— Steel, 1949b: 235 ( Austrolophrum ; synonym of cribriceps ).

guichardi Steel, 1949 , see: cribriceps Fauvel, 1878 .

Brouniellum

[7 species; Australian Region]

Brouniellum Bernhauer, 1939f: 198 (species included: sagoloide , australe , majus , hirtellum , parcum ). TYPE SPECIES: Brouniellum hirtellum Bernhauer , fixed by original designation.

— Cameron, 1947d: 701 (characters).

— Blackwelder, 1952: 86 (type species: hirtellum ).

— Klimaszewski, Newton, and Thayer, 1996: 146 (endemic to New Zealand).

australe Broun, 1894: 426 ( Omalium ; Type locality: West Plains , Invercargill).

— Bernhauer and Schubert, 1910: 51 ( Omalium ; subgenus Omalium ; catalog).

— Bernhauer, 1939f: 200 ( Brouniellum ).

- DISTRIBUTION: New Zealand.

hilarum Broun, 1914: 90 ( Omalium ; Type locality: Erua, near Waimarino).

— Scheerpeltz, 1933: 1045 ( Omalium ; catalog).

— Kuschel, 1990: 25, 43 ( Brouniellum ; endemic; winged; habitat; New Zealand).

- DISTRIBUTION: New Zealand.

hirtellum Bernhauer, 1939f: 200 ( Brouniellum ; Type locality: Neu-Seeland: North Island: Pudding Hill , near Methven ; Auckland).

- DISTRIBUTION: New Zealand.

majus Bernhauer, 1939f: 201 ( Brouniellum ; Type locality: New Zealand).

- DISTRIBUTION: New Zealand.

parcum Bernhauer, 1939f: 201 ( Brouniellum ; Type locality: Neu-Seeland: North Island: Pudding Hill, near Methven; Howick).

- DISTRIBUTION: New Zealand.

sagoloide Sharp, 1886 a: 381 ( Omalium ; Type locality: New Zealand: Picton ).

— Broun, 1893: 1035 ( Omalium ; characters; New Zealand).

— Bernhauer and Schubert, 1910: 56 ( Omalium ; subgenus Omalium ; catalog).

— Bernhauer, 1939f: 200 ( Brouniellum ).

- DISTRIBUTION: New Zealand.

zealandicum Cameron, 1947d: 702 ( Brouniellum ; Type locality: New Zealand: Canterbury: Mistake Creek. Also cited from Stewart Island ).

- DISTRIBUTION: New Zealand.

Carcinocephalus

[4 species; Palaearctic Region]

Carcinocephalus Bernhauer, 1903c: 592 (replacement name for Astacops Bernhauer ). TYPE SPECIES: Omalium merkli Eppelsheim , fixed by R. Lucas, 1920: 164, by subsequent designation.

— Heyden, Reitter, and Weise, 1906: 132 (list of species of Europe and Caucasus).

— Bernhauer and Schubert, 1910: 61 (world catalog; 2 species).

— Winkler, 1925: 332 (catalog for Palaearctic region).

— Scheerpeltz, 1934b: 81 (characters; notes; key to species).

— Székessy, 1963: 15 (characters).

— Tóth, 1982: 68 ( Carcinocephalus ; characters; Hungary).

— Zanetti, 1987: 240 (characters; habitus; species of Italy).

— Zanetti, 1995: 7 (list of species of Italy).

— Newton, Thayer, Ashe, and Chandler, 2000: 336 (3? Nearctic species; placement of species uncertain; notes; characters in key).

Astacops Bernhauer, 1902c: 61 (species included: merkli , mehadiensis ; preoccupied by Boisduval, 1835). TYPE SPECIES: Omalium merkli Eppelsheim , fixed by objective synonymy with Carinocephalus for which R. Lucas had fixed the type species.

— Blackwelder, 1952: 64 (type species).

— Zanetti, 1987: 240 (synonym of Carcinocephalus ).

Scribaia Luze, 1906: 505 (subgenus of Omalium ; species included: blandum ). TYPE SPECIES: Omalium (Scribaia) blandum Luze , fixed by monotypy.

— Bernhauer and Schubert, 1910: 57 (subgenus of Omalium ).

— Scheerpeltz, 1934b: 81, 89 (subgenus of Carcinocephalus ; characters; notes).

— Blackwelder, 1952: 349 (type species: blanda ).

— Zanetti, 1987: 242 (subgenus of Carcinocephalus ; characters; habitus; species of Italy).

blandus Luze, 1906: 546 ( Omalium ; subgenus Scribaia ; Type locality: Italien, etruskischer Apennin, Porretta; Veltlin).

— Bernhauer and Schubert, 1910: 57 ( Omalium ; subgenus Scribaia ; catalog).

— Scheerpeltz, 1934b: 81, 89 ( Carcinocephalus ; subgenus Scribaia ; characters; notes; Italy).

— Zanetti, 1987: 242 ( Carcinocephalus ; subgenus Scribaia ; characters; Italy).

— Zanetti, 1995: 7 ( Carcinocephalus ; Italy).

- DISTRIBUTION: Italy.

bosnicus Scheerpeltz, 1934b: 86 ( Carcinocephalus ; subgenus Carcinocephalus ; Type locality: Ilidze, Bosnien, im Tale der Bosna westlich von Sarajevo).

- DISTRIBUTION: Bosnia Hercegovina.

mehadiensis Bernhauer, 1902c: 62 ( Astacops ; Type locality: Graz bei Mehadia in Südungarn).

— Bernhauer and Schubert, 1910: 61 ( Carcinocephalus ; catalog).

— Scheerpeltz, 1934b: 90 ( Carcinocephalus ; subgenus Carcinocephalus ; characters).

— Tikhomirova, 1973a: 139 ( Carcinocephalus , cited as Carinocephalus; checklist; southern Carpathians).

— Tóth, 1982: 69 ( Carcinocephalus ; characters).

- DISTRIBUTION: Romania.

merkli Eppelsheim, 1883: 305 ( Omalium , cited as Homalium ; Type locality: Kodscha- Balkan).

— Bernhauer and Schubert, 1910: 61 ( Carcinocephalus ; catalog).

— Petri, 1912: 171 ( Carcinocephalus ; Romania).

— Scheerpeltz, 1934b: 91 ( Carcinocephalus ; subgenus Scribaia ; characters; Balkan region).

— Tikhomirova, 1973a: 139 ( Carcinocephalus , cited as Carinocephalus; checklist; northern Balkans).

— Tóth, 1982: 69 ( Carcinocephalus ; characters).

- DISTRIBUTION: Romania, Balkan region.

Crymus

[2 species; Neotropical, Antarctic, and Australian Regions]

Crymus Fauvel, 1904b: 92 (species included: antarcticus ). TYPE SPECIES: Crymus antarcticus Fauvel , fixed by monotypy.

— Bernhauer and Schubert, 1911: 91 (world catalog; 1 species).

— Scheerpeltz, 1933: 1078 (world catalog supplement).

— Herman, 1970: 397 (notes).

— Steel, 1970: 240 (synonyms).

— Steel, 1970a: 24 (larval characters; notes).

— Newton, 1985: 198 (notes on austral distribution).

Arpediomimus Cameron, 1917b: 277 (replacement name for Arpediopsis Cameron ). TYPE SPECIES: Arpediopsis falklandicus Cameron , fixed by objective synonymy with Arpediopsis .

— Scheerpeltz, 1933: 1031 (world catalog supplement).

— Jeannel, 1940: 116, 117 (characters; type species: falklandicus Cameron ).

— Gourlay, 1950: 182 (notes).

— Blackwelder, 1952: 62 (type species: falklandicus ).

— Steel, 1964: 348 (characters; key to species).

— Steel, 1970: 240 (synonym of Crymus ).

— Moore and Legner, 1976: 536 (includes seashore species; notes).

Arpediopsis Cameron, 1917: 124 (species included: falklandicus ; Preoccupied by Ganglbauer, 1895). TYPE SPECIES: Arpediopsis falklandica Cameron , fixed by monotypy.

— Scheerpeltz, 1933: 1031 (synonym of Arpediomimus ).

— Gourlay, 1950: 182 (synonym of Arpediomimus ).

— Blackwelder, 1952: 62 (type species: falklandica ).

— Steel, 1964: 348 (synonym of Arpediomimus ).

— Steel, 1970: 240 (synonym of Crymus ).

antarcticus Fauvel, 1904b: 93 ( Crymus ; Type locality: Nouvelle-Georgie du Sud).

— Bernhauer and Schubert, 1911: 91 ( Crymus ; catalog).

— Holdhaus, 1932: 166 ( Crymus ; notes; South Georgia Island).

— Scheerpeltz, 1933: 1078 ( Crymus ; catalog).

— Steel, 1970: 240 ( Crymus ; characters; South Georgia).

— Steel, 1970a: 24 ( Crymus ; larval characters).

- DISTRIBUTION: South Georgia, Falkland Islands.

falklandicus Cameron, 1917: 124 ( Arpediopsis ; Type locality: Sandy Beaches in vicinity of Port Stanley, Falkland Islands).

— Scheerpeltz, 1933: 1031 ( Arpediomimus ; catalog).

— Steel, 1964: 349 ( Arpediomimus ; characters; Falkland Islands; South Georgia).

— Steel, 1970: 240 ( Crymus ; synonym of antarcticus ).

falklandicus Cameron, 1917 , see: antarcticus Fauvel, 1904 .

kronii Kiesenwetter, 1877: 161 ( Omalium ; Type locality: Auckland Inseln ).

— Fauvel, 1877: 188 ( Omalium , cited as Homalium ; cited as kronei; characters; Auckland Islands).

— Bernhauer and Schubert, 1910: 54 ( Omalium ; subgenus Omalium ; cited as kronei; catalog).

— Scheerpeltz, 1933: 1045 ( Omalium ; cited as kronei; catalog).

— Gourlay, 1950: 182 ( Arpediomimus ; cited as kronii ; collecting notes and range extension).

— Steel, 1964: 350 ( Arpediomimus ; cited as kronei; adult and larval characters; Campbell Island; Auckland Island; South Island).

— Klimaszewski, Newton, and Thayer, 1996: 146 ( Crymus ; New Zealand).

— Marris, 2000: 175 ( Crymus ; habitat; Bollons Island [Antipodes]; also known from Auckland Islands; Campbell Island; New Zealand).

- DISTRIBUTION: New Zealand, Auckland Island, Campbell Island; Antipodes Islands.

longiceps Broun, 1914: 89 ( Omalium ; Type locality: Hakapoua , Southland).

— Scheerpeltz, 1933: 1046 ( Omalium ; catalog).

— Gourlay, 1950: 182 ( Arpediomimus ; synonym of kronii , cited as kronei).

— Steel, 1964: 350 ( Arpediomimus ; synonym of kronii , cited as kronei).

longiceps Broun, 1914 , see: kronii Kiesenwetter, 1877 .

Dialycera

[5 species; Palaearctic Region]

Dialycera Ganglbauer, 1895: 739 , 743 (subgenus of Phyllodrepa ; species included: distincticornis ). TYPE SPECIES: Omalium distincticornis Baudi , fixed by monotypy.

— Luze, 1906: 576 (characters; key to species). — Bernhauer and Schubert, 1910: 49 (subgenus of Phyllodrepa ; catalog).

— Blackwelder, 1952: 122 (subgenus of Hapalaraea ; type species: distincticornis ).

— Zanetti, 1987: 202 (cited as valid genus; characters; habitus; key to species of Italy).

— Zanetti, 1995: 7 (list of species of Italy).

armena Khnzorian, 1959: 66 ( Phyllodrepa ; subgenus Dialycera ; Type locality: ArmSSR: Areni, Ekhegnadzorskii r-n).

— Tikhomirova, 1973a: 138 ( Phyllodrepa ; subgenus Dialycera ; checklist; Armenia).

— Herman, 2001: 13 ( Dialycera ).

- DISTRIBUTION: Armenia.

bimaculata Luze, 1906: 578 ( Dialycera ; Type locality: Kleinasien: Bulghar-Dagh).

— Bernhauer and Schubert, 1910: 50 ( Phyllodrepa ; subgenus Hypopycna ; catalog).

— J. Sahlberg, 1913: 58 ( Dialycera ; Caramania).

— Khnzorian, 1959: 68 ( Phyllodrepa ; subgenus Dialycera ; characters).

— Tikhomirova, 1973a: 138 ( Phyllodrepa ; subgenus Dialycera ; checklist; Asia Minor).

- DISTRIBUTION: Turkey.

distincticornis Baudi, 1870: 402 ( Omalium ; Type locality: Pedemontio; [Note: See lectotype designation by Zanetti, 1986]).

— Fauvel, 1871a: 87 [= 1872: 61] ( Omalium , cited as Homalium ; characters; France; Italy).

— Mulsant and Rey, 1880: 279 ( Hypopycna ; characters; France).

— Ganglbauer, 1895: 743 ( Phyllodrepa ; subgenus Hypopycna ; characters; Europe).

— Luze, 1906: 577 ( Dialycera ; characters; Italy; France).

— Bernhauer and Schubert, 1910: 50 ( Phyllodrepa ; subgenus Hypopycna ; catalog).

— Español, 1945: 90 ( Phyllodrepa ; subgenus Dialycera ; Spain).

— Khnzorian, 1959: 68 ( Phyllodrepa ; subgenus Dialycera ; characters).

— Zerche, 1991a: 382, 385 ( Dialycera ; aedeagus illustrated; characters discussed).

— Tóth, 1982: 51 ( Hapalaraea ; characters).

— Zanetti, 1986: 97 ( Dialycera ; lectotype designation).

— Zanetti, 1987: 202 ( Dialycera ; characters; Italy).

— Sparacio, 1995: 140 ( Dialycera ; notes).

— Zanetti, 1995: 7 ( Dialycera ; Italy).

- DISTRIBUTION: Italy, France, Spain.

subrugata Rey, 1889 : cxix ( Hypopycna ; Type locality: Hyères).

— Ganglbauer, 1895: 743 ( Phyllodrepa ; synonym of distincticornis ).

— Luze, 1906: 577 ( Dialycera ; synonym of distincticornis ).

— Bernhauer and Schubert, 1910: 50 ( Phyllodrepa ; synonym of distincticornis ).

sekerae Reitter, 1905b: 242 ( Phyllodrepa ; subgenus Dialycera ; Type locality: Italien, bei Bazzano, unweit Bologna).

— Bernhauer and Schubert, 1910: 50 ( Phyllodrepa ; subgenus Hypopycna ; catalog).

— Zanetti, 1986: 95 ( Dialycera ; synonym of distincticornis ).

— Zanetti, 1987: 202 ( Dialycera ; synonym of distincticornis ).

minuta Luze, 1906: 582 ( Dialycera ; Type locality: Galiläa: Kana).

— Bernhauer and Schubert, 1910: 50 ( Phyllodrepa ; subgenus Dialycera ; catalog).

— J. Sahlberg, 1913: 58 ( Dialycera ; Palestine).

— Khnzorian, 1959: 68 ( Phyllodrepa ; subgenus Dialycera ; characters).

- DISTRIBUTION: Israel.

sekerae Reitter, 1905 , see: distincticornis Baudi, 1870 .

striatipennis Aubé, 1850: 321 ( Omalium ; Type locality: environs de Batoum, Iméritie).

— Luze, 1906: 579 ( Dialycera ; characters; Caucasus).

— Bernhauer and Schubert, 1910: 50 ( Phyllodrepa ; subgenus Dialycera ; catalog).

— Khnzorian, 1959: 68 ( Phyllodrepa ; subgenus Dialycera ; characters).

— Tikhomirova, 1973a: 138 ( Phyllodrepa ; subgenus Dialycera ; checklist; Caucasus).

— Herman, 2001: 13 ( Dialycera ).

- DISTRIBUTION: Georgia.

Hapalaraea

[7 species; Nearctic and Palaearctic Regions]

Hapalaraea Thomson, 1858: 38 (species included: pygmaea ). TYPE SPECIES: Staphylinus pygmaeus Paykull , fixed by monotypy.

— Thomson, 1859: 50 (characters; type species: pygmaea ).

— Thomson, 1861: 200 (characters).

— Fauvel, 1871a: 83, 86 [= 1872: 57, 61] (subgroup of Omalium , cited as Homalium ).

— Fauvel, 1875a: IV [= 1875b: 206] (catalog; group of Omalium , cited as Homalium ).

— Mulsant and Rey, 1880: 280 (characters).

— Fowler, 1888: 423 (notes).

— Heyden, Reitter, and Weise, 1891: 121 (list of species of Europe and Caucasus).

— Heyden, Reitter, and Weise, 1906: 130 (subgenus of Phyllodrepa ; list of species of Europe and Caucasus).

— Bernhauer and Schubert, 1910: 49 (subgenus of Phyllodrepa ; catalog).

— Roubal, 1930: 307 (catalog; Slovakia).

— Scheerpeltz, 1933: 1042 (world catalog supplement; subgenus of Phyllodrepa ).

— Palm, 1948: 74 (subgenus of Dropephylla ; species of Sweden).

— Tottenham, 1949: 354, 406 (type species: pygmaea Gyllenhal ; list of British species; notes).

— Blackwelder, 1952: 180 (type species: pygmaea ).

— Tottenham, 1954: 19 (characters; key to British subgenera and species).

— Hatch, 1957: 52, 76 (characters; key to species of Pacific Northwest).

— Székessy, 1963: 14 (characters).

— Lohse, 1964: 37, 40 (subgenus of Phyllodrepa ; characters).

— Pototskaia, 1967: 20 (larval characters).

— Steel, 1970a: 10 (larval characters; biological notes).

— Tikhomirova, 1973a: 138 (subgenus of Phyllodrepa ; checklist of species of USSR).

— Moore and Legner, 1975: 193 (catalog; North America north of Mexico).

— Pototskaia, 1976: 164, 165 (larval characters).

— Pope, 1977: 23 (list of British species).

— Topp, 1978: 308 (subgenus of Phyllodrepa ; larval characters).

— Burakowski, Mroczkowski, and Stefańska, 1979: 30 (catalog; Poland).

— Moore and Legner, 1979: 206 (characters; notes).

— Muona, 1979: 17 (list of Scandinavian species).

— Tóth, 1982: 50 (characters; habitus; key to species of Hungary).

— Zanetti, 1983a: 461 (characters; valid genus).

— Zanetti, 1987: 195 (characters; habitus; key to species of Italy).

— Muona and Viramo, 1986: 15 (list of species of northeastern Finland).

— Lohse and Lucht, 1989: 126 (cited as genus).

— Campbell and Davies, 1991: 88 (checklist of species; Alaska and Canada).

— Silfverberg, 1992: 21 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Boháč, 1993: 40 (list of species of Czech Republic and Slovakia).

— Zanetti, 1995: 7 (list of species of Italy).

— M. Hansen, 1996: 94 (list of Danish species).

— Downie and Arnett, 1996: 436 (characters).

— Lundgren, 1998: 38 (list of species from Florida; records for other States listed for each species).

— Newton, Thayer, Ashe, and Chandler, 2000: 336 (2 Nearctic species; notes; characters in key).

atriceps Motschulsky, 1860 , see: pygmaea Paykull, 1800 .

barri Hatch, 1957 , see: hamata Fauvel, 1878 .

hamata Fauvel, 1878d: 209 [= 1878e: 45] ( Omalium , cited as Homalium ; Type locality: Michigan, Detroit).

— Bernhauer and Schubert, 1910: 54 ( Omalium ; subgenus Omalium ; catalog).

— Blatchley, 1910: 480 ( Omalium , cited as Homalium ; characters; Indiana). — Downie and Arnett, 1996: 438 ( Omalium ; characters; USA). — Newton, Thayer, Ashe, and Chandler, 2000: 336 ( Hapalaraea ). - DISTRIBUTION: USA.

barri Hatch, 1957: 77 ( Acrolocha ; Type locality: Ida.: Juliaetta). — Steel, 1959: 82 ( Acrolocha ; appears to be same as Hapalaraea hamata ). — Newton, Thayer, Ashe, and Chandler, 2000: 336 ( Hapalaraea ; synonym of hamata ).

humerosa Fauvel, 1878d: 209 [= 1878e: 45] ( Omalium , cited as Homalium ; Type locality: Michigan, Detroit; Pennsylvanie; Missouri). — Bernhauer and Schubert, 1910: 54 ( Omalium ; subgenus Omalium ; catalog). — Notman, 1920b: 693 ( Omalium ; Florida). — Downie and Arnett, 1996: 438 ( Omalium ; characters; USA). — Lundgren, 1998: 38 ( Hapalaraea ; Florida; Michigan; North Carolina). - DISTRIBUTION: USA.

humilis Mäklin, 1853: 197 ( Omalium ; Type locality: interior pars peninsulae Kenai prope fl. Tschunuktnu). — Fauvel, 1878d: 211 [= 1878e: 47] ( Omalium , cited as Homalium ; characters; Alaska). — Bernhauer and Schubert, 1910: 53 ( Omalium ; subgenus Omalium ; catalog). — Hatch, 1957: 81 ( Omalium ; characters; British Columbia ). — Moore and Legner, 1975: 193 ( Hapalaraea ). - DISTRIBUTION: Canada, USA.

lestevoides Sawada, 1966: 257 ( Hapalaraea ; subgenus Phyllodrepa ; Type locality: Mt. Noshaq, 3800 m). - DISTRIBUTION: Afghanistan.

megarthroides Fauvel, 1878d: 208 [= 1878e: 44] ( Omalium , cited as Homalium ; Type locality: Californie). — Bernhauer and Schubert, 1910: 49 ( Phyllodrepa ; subgenus Hapalaraea ; catalog). — Hatch, 1957: 77 ( Acrolocha ; characters; British Columbia ; Idaho; Washington; Oregon). — Steel, 1959b: 82 ( Hapalaraea ). - DISTRIBUTION: Canada, USA.

pygmaea Paykull, 1800: 410 ( Staphylinus ; [preoccupied]; [Note: Bernhauer and Schubert (1910: 49) cited pygmaea Gyllenhal (1810: 223) as a species of Phyllodrepa (Hapalaraea) . Gyllenhal did not describe the species he attributed it to Paykull but he also cited pygmaeus Gravenhorst, 1806 as a subsequent reference of pygmaeus Paykull ; the Gravenhorst species cited by Gyllenhal is now in Xylodromus where it is listed as a synonym of testaceus Erichson ; pygmaeus Gravenhorst is an available name and Gravenhorst made no mention of the Paykull species]; Type locality: Westrogothia). — Gyllenhal, 1810: 223 ( Omalium ; [Note: Bernhauer and Schubert (1910: 49) cited pygmaea Gyllenhal as a valid species; Gyllenhal did not described the species as new, he attributed it to Paykull; further he treated pygmaea Gravenhorst, 1806: 206 as a subsequent reference of pygmaea Paykull ; pygmaea Gravenhorst is currently in Xylodromus ]; characters). — Stephens, 1834: 356 ( Omalium ; [Note: Stephens, 1834: 356 cited Paykull as the author of this name; Stephens, 1829: 298 cited pygmaeus with references by Gravenhorst, 1806: 206 and Paykull, 1800: 410 as the same species]; characters; England). — Lacordaire, 1835: 488 ( Omalium ; [Note: Lacordaire cited pygmaeus of Paykull, 1800, Gravenhorst, 1806 and Gyllenhal, 1810 as referring to the same species]; characters; France). — Stephens, 1839: 430 ( Omalium ; characters; England). — Erichson, 1840: 883 ( Omalium ; [Note: Erichson cited Paykull, 1800 and Gyllenhal, 1810 as referring to the same species; see pygmaeus Gravenhorst ]; characters; Germany; Sweden). — Heer, 1841: 570 ( Phloeonomus ; characters; Switzerland). — Redtenbacher, 1849: 829 ( Omalium ; characters; Austria). — Thomson, 1857a: 226 ( Omalium ; characters). — Fairmaire and Laboulbène, 1856: 645 ( Omalium ; characters; France). — Kraatz, 1857d: 1001 ( Omalium ; characters; Germany).

— Thomson, 1858: 38 ( Hapalaraea ; type species).

— Thomson, 1859: 50 (type species of Hapalaraea ).

— Thomson, 1861: 201 ( Hapalaraea ; characters; Scandinavia).

— Fauvel, 1871a: 87 [= 1872: 61] ( Omalium , cited as Homalium ; characters; France; Scandinavia; Britain; Germany; Austria; Corsica).

— Redtenbacher, 1874: 276 ( Omalium ; characters; Austria).

— Seidlitz, 1875: 238 ( Hapalaraea ; characters; Baltic region).

— Mulsant and Rey, 1880: 282 ( Hapalaraea ; characters; France).

— Fowler, 1888: 423 ( Hapalaraea ; characters; Britain).

— Seidlitz, 1891: 344 ( Hapalaraea ; characters; Europe).

— Seidlitz, 1891a: 366 ( Hapalaraea ; characters; Europe).

— Ganglbauer, 1895: 742 ( Phyllodrepa ; subgenus Hapalaraea ; characters; north and middle Europe; Corsica).

— Sainte-Claire Deville, 1906: 68 ( Omalium , cited as Homalium ; Corsica).

— Reitter, 1909: 193 ( Phyllodrepa ; subgenus Hapalaraea ; characters; Germany).

— Bernhauer and Schubert, 1910: 49 ( Phyllodrepa ; subgenus Hapalaraea ; Gyllenhal cited as author; catalog).

— Johansen, 1914: 617 ( Phyllodrepa ; subgenus Hapalaraea ; characters; Denmark).

— Portevin, 1929: 449 ( Phyllodrepa ; characters; France).

— Scheerpeltz, 1933: 1042 ( Phyllodrepa ; subgenus Hapalaraea ; catalog).

— Palm, 1948: 74 ( Phyllodrepa ; subgenus Hapalaraea ; characters; Sweden; Denmark; Norway; Finland).

— Tottenham, 1949: 406 ( Hapalaraea ; checklist; Britain).

— Horion, 1963: 71 ( Phyllodrepa ; subgenus Hapalaraea ; Britain; Denmark; Norway; Sweden; Finland; Holland; Belgium; France; Corsica; Italy; Slovenia; Croatia; Hercegovina; Romania; Hungary; Slovakia; Czech Republic; Poland; Germany; Austria).

— Lohse, 1964: 40 ( Phyllodrepa ; subgenus Hapalaraea ; characters; central Europe).

— Smetana, 1964c: 163 ( Phyllodrepa ; subgenus Hapalaraea ; Czechoslovakia).

— Focarile, 1964: 60 ( Phyllodrepa ; Italy).

— Coiffait, 1968c: 505 ( Hapalaraea ; Corsica).

— Steel, 1970a: 10 ( Hapalaraea ; larval characters; biological notes).

— Tikhomirova, 1973a: 138 ( Phyllodrepa ; subgenus Hapalaraea ; checklist; USSR).

— Legner and Moore, 1977: 175 ( Hapalaraea ; found under bark).

— Pope, 1977: 23 ( Hapalaraea ; Britain).

— Muona, 1979: 18 ( Hapalaraea ; Finland; Russia; Sweden; Norway; Denmark).

— Burakowski, Mroczkowski, and Stefańska, 1979: 37 ( Hapalaraea ; catalog; Poland).

— Hammond, 1980: 137 ( Hapalaraea ; unconfirmed in Ireland).

— Tóth, 1980: 97 ( Phyllodrepa ; subgenus Hapalaraea ; Hungary).

— Tóth, 1982: 57 ( Hapalaraea ; characters; Hungary).

— Zanetti, 1983a: 464 ( Hapalaraea ; characters).

— Terlutter, 1984: 19, 39 ( Phyllodrepa ; notes; Germany).

— Segers, 1986: 20 ( Hapalaraea ; checklist of species; Belgium).

— Zanetti, 1987: 196 ( Hapalaraea ; characters; Italy).

— Lucht, 1987: 87 ( Phyllodrepa ; checklist; Germany; Poland; Czechoslovakia; Austria; Belgium; Netherlands; Luxembourg; Denmark).

— Lohse and Lucht, 1989: 285 ( Hapalaraea ; also cited in Phyllodrepa ; checklist; Germany; Poland; Czechoslovakia; Austria; France; Belgium; Netherlands; Luxembourg; Denmark; Sweden).

— Gusarov, 1989: 5 ( Hapalaraea ; habitat notes; Ukraine).

— Boháč, 1993: 40 ( Hapalaraea ; subgenus Hapalaraea ; Slovakia; Czech Republic).

— Silfverberg, 1992: 21 ( Hapalaraea ; Finland; Russia; Sweden; Norway; Denmark).

— Wittwer, 1993: 248 ( Hapalaraea ; Switzerland).

— Boháč, 1993: 40 ( Hapalaraea ; Czech Republic).

— Zanetti, 1995: 7 ( Hapalaraea ; Italy).

— Anderson, Nash, and O’Connor, 1997: 15 ( Hapalaraea ; Ireland).

— Owen, 1999b: 61 ( Hapalaraea ; captured underground at roots of old trees; Britain).

— Fowles, Alexander, and Key, 1999: 129 ( Hapalaraea ; saproxylic species; status as endangered or threatened species; Britain).

— Lott, 1999: 72 ( Hapalaraea ; saproxylic species; habitat; endangered status; Britain).

— Ryabukhin, 1999: 16 (type species of Hapalaraea ).

— Owen, 2000: 251 ( Hapalaraea ; pitfall trap collections from underground at roots of trees; Britain).

— Herman, 2001: 37 ( Hapalaraea ; although a junior primary homonym of Staphylinus pygmaeus Paykull , required replacement should be ingnored; prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Europe.

ulmi Motschulsky, 1857 a: 493 ( Phloeonomus ; Type locality: Fontainebleau).

— Fauvel, 1871a: 87 [= 1872: 61] ( Omalium , cited as Homalium ; synonym of pygmaeum ).

— Bernhauer and Schubert, 1910: 49 ( Phyllodrepa ; synonym of pygmaea ).

atriceps Motschulsky, 1860: 543 ( Anthobium ; Type locality: Mont. Tschatyr-Dag en Crimée).

— Hochhuth, 1862: 109 ( Anthobium ; characters; Crimea).

— Bernhauer and Schubert, 1910: 39 ( Anthobium ; catalog).

— Zanetti, 1993: 261 ( Hapalaraea ; synonym of pygmaea ).

settei Zanetti, 1983 a: 462 ( Hapalaraea ; Type locality: Haute Asco , Corsica, 1550 m).

— Zanetti, 1987: 198 ( Hapalaraea ; characters; Corsica).

- DISTRIBUTION: Corsica.

ulmi Motschulsky, 1857 , see: pygmaea Paykull, 1800 .

Hypopycna

[1 species; Palaearctic Region]

Hypopycna Mulsant and Rey, 1880: 274 (species included: rufula ; distincticornis ). TYPE SPECIES: Omalium rufulum Erichson , fixed by subsequent designation by R. Lucas, 1920: 346.

— Luze, 1906: 574 (characters).

— Heyden, Reitter, and Weise, 1906: 130 (subgenus of Phyllodrepa ; list of species of Europe and Caucasus).

— Scheerpeltz, 1933: 1043 (world catalog supplement; subgenus of Phyllodrepa ).

— Tottenham, 1949: 355, 406 (subgenus of Hapalaraea ; type species: rufula Erichson ; list of British species).

— Blackwelder, 1952: 196 (type species: rufula ).

— Lohse, 1964: 37, 41 (subgenus of Phyllodrepa ; characters).

— Steel, 1970a: 9 (valid genus; larval characters; biological notes).

— Bordoni, 1973j: 656 (subgenus of Phyllodrepa ; list of species from islands near Sicily).

— Pope, 1977: 23 (list of British species).

— Topp, 1978: 308 (cited as subgenus of Phyllodrepa ; larval characters).

— Zanetti, 1987: 199 (valid genus; characters; habitus; key to species of Italy).

— Lohse and Lucht, 1989: 126 (cited as genus; notes).

— Zanetti, 1995: 7 (list of species of Italy).

marina Ragusa, 1871 , see: rufula Erichson, 1840 .

rufula Erichson, 1840: 883 ( Omalium ; Type locality: Austria).

— Fairmaire and Laboulbène, 1856: 645 ( Omalium ; characters; France).

— Redtenbacher, 1857: 252 ( Omalium ; characters; Austria).

— Fauvel, 1871a: 86 [= 1872: 60] ( Omalium , cited as Homalium ; characters; France; Germany; Austria; Italy).

— Redtenbacher, 1874: 276 ( Omalium ; characters; Austria).

— Mulsant and Rey, 1880: 279 ( Hypopycna ; [Note: Misidentification: Bernhauer and Schubert,

1910: 49 cited distincticornis Rey as a synonym of rufula (cited as ufula); Mulsant and Rey did not describe the species as new, they attributed it to Baudi]; characters; France).

— Fauvel, 1886: 96 [= 1886a: 88] ( Omalium , cited as Homalium ; Tunisia).

— Seidlitz, 1891: 344 ( Hypopycna ; characters; Europe).

— Seidlitz, 1891a: 366 ( Hypopycna ; characters; Europe).

— Ganglbauer, 1895: 742 ( Phyllodrepa ; subgenus Hypopycna ; characters; middle Europe;

Mediterranean region).

— Fauvel, 1902b: 52 ( Omalium , cited as Homalium ; Algeria; Tunisia).

— Luze, 1906: 574 ( Hypopycna ; characters; central Europe; Mediterranean region).

— Sainte-Claire Deville, 1906: 68 ( Omalium , cited as Homalium ; Corsica).

— Reitter, 1909: 194 ( Phyllodrepa ; subgenus Hypopycna ; characters; Germany).

— Bernhauer and Schubert, 1910: 49 ( Phyllodrepa ; subgenus Hapalaraea ; cited as ufula; catalog).

— Portevin, 1929: 448 ( Phyllodrepa ; characters; France).

— Scheerpeltz, 1933: 1043 ( Phyllodrepa ; subgenus Hypopycna ; catalog).

— Blair, 1944: 20 ( Phyllodrepa ; subgenus Hypopycna ; characters; Britain).

— Horion, 1963: 73 ( Phyllodrepa ; subgenus Hypopycna ; Tunisia; France; Corsica; Sardinia;

Sicily; Italy; Dalmatia; Austria).

— Lohse, 1964: 41 ( Phyllodrepa ; subgenus Hypopycna ; characters; central Europe).

— Steel, 1970a: 9 ( Hypopycna ; larval characters; biological notes).

— Pope, 1977: 23 ( Hypopycna ; Britain).

— Tóth, 1982: 52 ( Hapalaraea ; characters).

— Zanetti, 1987: 200 ( Hypopycna ; characters; Italy).

— Lucht, 1987: 87 ( Phyllodrepa ; checklist; Germany; Austria).

— Owen, 1993a: 187 ( Phyllodrepa ; subgenus Hypopycna ; collecting notes; England).

— Hodge and Jones, 1995: 30 ( Hypopycna ; characters; Britain).

— Sparacio, 1995: 140 ( Hypopycna ; characters; notes; Sicily).

— Zanetti, 1995: 7 ( Hypopycna ; Italy).

— Owen, 1997: 302 ( Hypopycna ; collecting notes; Britain).

— Zanetti and Sabella, 1998: 29 ( Hypopycna ; Sicily).

- DISTRIBUTION: Britain, France, Italy, Austria, Croatia, Algeria, Tunisia. marina Ragusa, 1871: 196 ( Omalium ; Type locality: Palermo).

— Fauvel, 1873b: 114 [= 1873c: 7] ( Omalium , cited as Homalium ; synonym of rufulum ).

— Ganglbauer, 1895: 742 ( Phyllodrepa ; synonym of rufula ).

— Luze, 1906: 574 ( Hypopycna ; synonym of rufula ).

— Bernhauer and Schubert, 1910: 49 ( Phyllodrepa ; synonym of rufula ). subrugata Rey, 1889 , see: Dialycera .

Ischnoderus

[15 species; Neotropical and Australian Regions]

Ischnoderus Fauvel, 1867 a: 46 [= 1868: 51] (species included: insignis ). TYPE SPECIES: Omalium insigne Fairmaire and Germain , fixed by monotypy.

— Gemminger and Harold, 1868: 671 (catalog).

— Bernhauer and Schubert, 1910: 82 (world catalog; 7 species).

— Scheerpeltz, 1933: 1070 (world catalog supplement).

— Blackwelder, 1944: 102 (checklist of species from Mexico, Central and South America , and the West Indies).

— Blackwelder, 1952: 200 (type species: insignis ).

— Coiffait and Saiz, 1968: 454 (characters; Chilean species).

Walkerellus Bernhauer, 1939f: 203 (subgenus of Ischnoderus ; species included: paradoxus ). TYPE SPECIES: Ischnoderus (Walkerellus) paradoxus Bernhauer , fixed by monotypy.

— Blackwelder, 1952: 403 (type species: paradoxus ).

arecobius Broun, 1910: 20 ( Ischnoderus ; Type locality: Waitakerei Range).

— Scheerpeltz, 1933: 1070 ( Ischnoderus ; catalog).

australicus Cameron, 1943 a: 338 ( Ischnoderus ; Type locality: W. Australia: Mundaring ). - DISTRIBUTION: Australia.

bruchi Bernhauer, 1912d: 30 ( Ischnoderus ; Type locality: Argentinien: Gob. Neuquen).

— Scheerpeltz, 1933: 1070 ( Ischnoderus ; catalog).

- DISTRIBUTION: Argentina.

cognatus Broun, 1910: 20 ( Ischnoderus ; Type locality: West Plains , Invercargill).

— Scheerpeltz, 1933: 1070 ( Ischnoderus ; catalog).

- DISTRIBUTION: New Zealand.

curtipennis Broun, 1915: 280 ( Ischnoderus ; Type locality: Pudding Hill, near Methven). — Scheerpeltz, 1933: 1070 ( Ischnoderus ; catalog).

- DISTRIBUTION: New Zealand.

fultoni Broun, 1893: 1041 ( Ischnoderus ; Type locality: Taieri).

— Bernhauer and Schubert, 1910: 82 ( Ischnoderus ; catalog).

- DISTRIBUTION: New Zealand.

genalis Broun, 1880: 118 ( Omalium ; Type locality: Whangarei Harbour).

— Broun, 1886: 943 ( Ischnoderus ).

— Bernhauer and Schubert, 1910: 82 ( Ischnoderus ; catalog).

— Kuschel, 1990: 25 ( Ischnoderus ; endemic; winged; habitat; New Zealand).

— Emberson, 1998: 32 ( Ischnoderus ; collecting notes; Chatham Islands).

- DISTRIBUTION: New Zealand, Chatham Islands.

insignis Fairmaire and Germain, 1861: 454 ( Omalium ; Type locality: Santiago).

— Fauvel, 1867a: 48 [= 1868: 53] ( Ischnoderus ; characters; Chile).

— Bernhauer and Schubert, 1910: 82 ( Ischnoderus ; catalog).

— Coiffait and Saiz, 1968: 454 ( Ischnoderus ; characters; Chile).

- DISTRIBUTION: Chile.

morosus Broun, 1893: 1041 ( Ischnoderus ; Type locality: Howick).

— Bernhauer and Schubert, 1910: 82 ( Ischnoderus ; catalog).

- DISTRIBUTION: New Zealand.

nitescens Bernhauer, 1939f: 202 ( Ischnoderus ; Type locality: Neu-Seeland: Greymouth). - DISTRIBUTION: New Zealand.

opaciceps Cameron, 1947d: 703 ( Ischnoderus ; Type locality: New Zealand: N. Auckland: Kava, Whangarei ).

- DISTRIBUTION: New Zealand.

paradoxus Bernhauer, 1939f: 203 ( Ischnoderus ; subgenus Walkerellus ; Type locality: Tasmania: Hobart).

- DISTRIBUTION: Australia.

pictulus Broun, 1893: 1040 ( Ischnoderus ; Type locality: near Howick).

— Bernhauer and Schubert, 1910: 82 ( Ischnoderus ; catalog).

- DISTRIBUTION: New Zealand.

politulus Broun, 1880: 119 ( Omalium ; Type locality: Parua).

— Broun, 1886: 943 ( Ischnoderus ).

— Bernhauer and Schubert, 1910: 82 ( Ischnoderus ; catalog).

- DISTRIBUTION: New Zealand.

tectus Broun, 1880: 117 ( Omalium ; Type locality: Parua).

— Broun, 1886: 943 ( Ischnoderus ).

— Bernhauer and Schubert, 1910: 82 ( Ischnoderus ; catalog).

— Kuschel, 1990: 25 ( Ischnoderus ; endemic; winged; habitat; New Zealand).

- DISTRIBUTION: New Zealand.

Leaskia

[2 species; Australian Region]

Leaskia Steel, 1938 a: 28 (species included: acidotiformis ). TYPE SPECIES: Leaskia acidotiformis Steel , fixed by original designation.

— Steel, 1949b: 239 (characters; illustrations; notes; key to species).

— Blackwelder, 1952: 212 (type species: acidotiformis ).

acidotiformis Steel, 1938 a: 30 ( Leaskia ; Type locality: Australia: Victoria: Wannon Valley). — Steel, 1949b: 242 ( Leaskia ; characters; illustrations; Australia).

- DISTRIBUTION: Australia.

spinipes Fauvel, 1878 g: 487 ( Amphichroum ; Type locality: Australie occidentale).

— Bernhauer and Schubert, 1910: 70 ( Amphichroum ; catalog).

— Steel, 1949b: 244 ( Leaskia ; characters; illustrations; Australia).

- DISTRIBUTION: Australia.

Macralymma

[1 species; Australian Region]

Macralymma Cameron, 1945b: 179 (species included: punctiventris ). TYPE SPECIES: Macralymma punctiventris Cameron , fixed by monotypy.

— Blackwelder, 1952: 229 (type species: punctiventris ).

— Moore and Legner, 1976: 536 (includes seashore species; notes).

punctiventre Cameron, 1945b: 179 ( Macralymma ; Type locality: New Zealand: Taieri Beach). — Emberson, 1998: 32 ( Macralymma ; collecting notes; Chatham Islands).

— Klimaszewski, Newton, and Thayer, 1996: 146 ( Macralymma ; endemic to New Zealand and Chatham Island).

- DISTRIBUTION: New Zealand, Chatham Islands.

Metaxylostiba

[3 species; Oriental Region]

Metaxylostiba Steel, 1960 a: 171 (species included: castaneipennis ). TYPE SPECIES: Phloeonomus castaneipennis Cameron , fixed by original designation and monotypy.

— Smetana, 1992c: 33, 34 (key to species; notes).

castaneipennis Cameron, 1928 g: 433 ( Phloeonomus ; Type locality: Borneo: Mt. Murud, 5000–6000 feet).

— Scheerpeltz, 1933: 1049 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

— Steel, 1960a: 171 ( Metaxylostiba ; characters; Sarawak).

— Hammond, 1984: 196 ( Metaxylostiba ; Borneo).

— Smetana, 1992c: 34 ( Metaxylostiba ; characters in key; notes; Borneo).

- DISTRIBUTION: Borneo.

hanskii Smetana, 1992c: 38 ( Metaxylostiba ; Type locality: Sarawak: 4th Division Gn. Mulu N.P.).

- DISTRIBUTION: Malaysia.

monticola Smetana, 1992c: 35 ( Metaxylostiba ; Type locality: Sabah: Borneo Sabah, Mt. Kinabalu N. P., below Laban Rata, 3150 m).

- DISTRIBUTION: Borneo.

Micralymma

[4 species; Nearctic and Palaearctic Regions] Micralymma Westwood, 1838b: 129 (species included: johnstonis ). TYPE SPECIES: Micralymma johnstonis Westwood , fixed by monotypy.

— Westwood, 1838a (August): 18 (characters; type species).

— Stephens, 1839: 422 (characters).

— Erichson, 1840: 819 (characters).

— Schiødte, 1845: 374 (characters).

— Schaum, 1852: 29 (list of species; Europe).

— Lacordaire, 1854: 134 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 625 (characters).

— Jacquelin du Val, 1857: 66 (characters).

— Redtenbacher, 1857: 241 (characters).

— Thomson, 1858: 37 (characters).

— G. Waterhouse, 1858: 31 (catalog of British species).

— Thomson, 1859: 49 (characters; type species: brevipenne ).

— Schaum, 1859: 31 (catalog; European species).

— LeConte, 1861a: 70 (characters).

— Thomson, 1861: 187 (characters).

— LeConte, 1863: 26 (list of North American species).

— Jacquelin du Val, 1868: 79 (catalog; European species).

— Fauvel, 1871a: 107 [= 1872: 81] (characters).

— Sharp, 1871d: 14 (list of British species).

— Redtenbacher, 1874: 271 (characters).

— Fauvel, 1875a:VI [= 1875b: 208] (catalog).

— Fauvel, 1878d: 222 [= 1878e: 58] (key to North American species).

— Mulsant and Rey, 1880: 4 (characters).

— Fowler, 1888: 407 (notes).

— Heyden, Reitter, and Weise, 1891: 119 (list of species of Europe and Caucasus).

— Heyden, 1893: 49 (list of species of Siberian region).

— Ganglbauer, 1895: 728 (characters).

— Heyden, Reitter, and Weise, 1906: 132 (list of species of Europe and Caucasus).

— Reitter, 1909: 188 (characters; key to German species).

— Bernhauer and Schubert, 1910: 61 (world catalog; 5 species).

— Johansen, 1914: 602 (characters).

— Leng, 1920: 94 (catalog of North American species).

— Winkler, 1925: 332 (catalog for Palaearctic region).

— Joy, 1932: 94 (key to British species).

— Scheerpeltz, 1933: 1052 (world catalog supplement).

— Kloet and Hincks, 1945: 161 (list of British species).

— Palm, 1948: 90 (characters; Swedish species).

— Tottenham, 1949: 356, 407 (type species: marina Ström = johnstonis Westwood ; list of British species; notes).

— Blackwelder, 1952: 242 (type species: johnstonis ).

— Tottenham, 1954: 28 (characters; British species).

— Steel, 1958: 138 (characters; key to species).

— Steel, 1962: 238 (key to species; notes).

— Horion, 1963: 97 (list of central European species).

— Lohse, 1964: 49 (characters; key to central European species).

— Pototskaia, 1967: 19 (larval characters).

— Kasule, 1968: 132, 134 (larval characters).

— Steel, 1970a: 17 (larval characters; biological notes).

— Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Moore and Legner, 1975: 197 (catalog; North America north of Mexico).

— Moore and Legner, 1976: 536 (includes seashore species; notes).

— Pope, 1977: 24 (list of British species).

— Topp, 1978: 309 (larval characters).

— Moore and Legner, 1979: 207 (characters; notes).

— Muona, 1979: 18 (list of Scandinavian species).

— Zanetti, 1983: 429 (characters).

— Campbell and Davies, 1991: 88 (checklist of species; Alaska and Canada).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Downie and Arnett, 1996: 436 (characters; species of Northeastern North America ).

Microcalymma Agassiz, 1847: 232 (emendation of Micralymma ). TYPE SPECIES: Micralymma johnstonis Westwood , fixed by objective synonymy with Micralymma . — Blackwelder, 1952: 243 (type species).

Microcalymma Gemminger and Harold, 1868: 663 (emendation of Micralymma ). TYPE SPECIES: Micralymma johnstonis Westwood , fixed by objective synonymy with Micralymma . — Duvivier, 1883: 197 (catalog). — Bernhauer and Schubert, 1910: 61 (synonym of Micralymma ). — Blackwelder, 1952: 243 (type species).

Micromalium Melichar, 1913: 45 (species included: caucasicum ). TYPE SPECIES: Micromalium caucasium Melichar , fixed by monotypy. — Winkler, 1925: 332 (catalog for Palaearctic region). — Scheerpeltz, 1933: 1053 (world catalog supplement). — Blackwelder, 1952: 246 (type species: caucasicum ). — Steel, 1962: 238 (synonym of Micralymma ).

brevilingue Schiødte, 1845: 377 ( Micralymma ; Type locality: insula Groenlandiae: Kikertarsoeitsiak). — Schiødte, 1846: 163 ( Micralymma ; characters; Greenland). — Fauvel, 1878d: 222 [= 1878e: 58] ( Micralymma ; characters; Greenland). — Bernhauer and Schubert, 1910: 62 ( Micralymma ; catalog). — Henriksen and Lundbeck, 1918: 496 ( Micralymma ; Greenland). — Lindroth, 1957: 256, 260, 263, 292 ( Micralymma ; notes; Siberia; tidal zone high Arctic circumpolar). — Henriksen, 1939: 43 ( Micralymma ; Greenland). — Steel, 1958: 140 ( Micralymma ; characters; Greenland to 70°N; Baffin Island). — Steel, 1962: 238 ( Micralymma ; characters). — M. Hansen and Martin, 2000: 216 ( Micralymma ; notes). - DISTRIBUTION: Greenland, Canada, USA, Russia, Islands of Arctic region.

dicksoni Mäklin, 1878: 24 [= 1878a: 300] ( Microcalymma ; Type locality: Jefremow Kamen, 72°40′ N. lat.; Krestowskoj 72°15′ N. lat.). — Mäklin, 1881: 42 ( Micralymma ; characters; Jefremov Kamen, lat. bor 72°40′; Krestowskoj, lat. bor. 72°15′). — J. Sahlberg, 1897: 367 ( Phyllodrepa ; notes; Siberia). — Poppius, 1909: 5 ( Micralymma ; Lena Valley, Siberia ). — Bernhauer and Schubert, 1910: 62 ( Micralymma ; catalog). — Van Dyck, 1921: 164 ( Micralymma ; Pribilof Islands, Alaska). — Scheerpeltz, 1933: 1053 ( Micralymma ; catalog). — Lindroth, 1957: 260, 261, 262, 292 ( Micralymma ; characters; notes; dicksoni and brevilingue are one species). — Steel, 1958: 142 ( Micralymma ; subspecies of brevilingue ; characters; Arctic coast of Siberia from Jenisei Bay to New siberian Islands; Commodore Islands; Chukotski Peninsula; Pribiloff Islands; N.W. Territory; Richards Islands). — Ryabukhin, 1999: 20 ( Micralymma ; cited as valid species; catalog; habitat; distribution; northeastern Russia).

brevipenne Gyllenhal, 1810 , see: marinum Ström, 1783 .

caucasicum Melichar, 1913: 46 ( Micromalium ; Type locality: Teberda im nordwestlichen Kaukasus). — Scheerpeltz, 1933: 1053 ( Micromalium ; catalog). — Steel, 1962: 238 ( Micralymma ; characters; Caucasus). — Tikhomirova, 1973a: 139 ( Micralymma ; checklist; northwestern Caucasus). - DISTRIBUTION: Russia.

dicksoni Mäklin, 1878 , see: brevilingue Schiødte, 1845 .

laticolle Motschulsky, 1860: 549 ( Micralymma ; Type locality: boréales de la Sibérie orientale près de la mer glaciale dans le Gouv. de Jeniseisk). — Bernhauer and Schubert, 1910: 62 ( Micralymma ; catalog). — Steel, 1958: 138 (not in Micralymma ; probably an Arpedium ). - DISTRIBUTION: Russia.

marinum Ström, 1783: 65 ( Staphylinus ; Type locality: Norwegen). — Fauvel, 1871a: 108 [= 1872: 82] ( Micralymma ; characters; France; Scandinavia; Denmark; Britain). — Seidlitz, 1875: 237 ( Micralymma ; characters; Baltic region). — Seidlitz, 1891: 343 ( Micralymma ; characters; Europe). — Seidlitz, 1891a: 366 ( Micralymma ; characters; Europe). — Ganglbauer, 1895: 729 ( Micralymma ; characters; France; Britain; Denmark; Scandinavia). — Poppius, 1905: 11 ( Micralymma ; Shetland Islands). — Sainte-Claire Deville, 1907: 40 ( Micralymma ; characters; France). — Reitter, 1909: 188 ( Micralymma ; characters; Germany). — Bernhauer and Schubert, 1910: 62 ( Micralymma ; catalog). — Johansen, 1914: 602 ( Micralymma ; characters; Denmark). — Henriksen and Lundbeck, 1918: 496 ( Micralymma ; Greenland). — Portevin, 1929: 443 ( Micralymma ; characters; France). — Scheerpeltz, 1933: 1053 ( Micralymma ; catalog). — West, 1937: 45 ( Micralymma ; Faroe Islands). — Henriksen, 1939: 43 ( Micralymma ; Greenland). — Paulian, 1941: 147 ( Micralymma ; cited as Micralymna; larval characters). — Palm, 1948: 90 ( Micralymma ; characters; Sweden; Norway; Finland). — Lindroth, 1957: 231, 244, 245, 260, 263 ( Micralymma ; notes on distribution and dispersal). — Steel, 1958: 140 ( Micralymma ; characters; correction of citation of original description; France; Britain; Shetlands; Faroes; western and northern Scandinavia; Bear Island; Iceland; Greenland; Maine; Newfoundland). — Larsson and Gígja, 1959: 66 ( Micralymma ; biological notes; Iceland). — Steel, 1962: 238 ( Micralymma ; characters). — Horion, 1963: 97 ( Micralymma ; Amphi-Atlantic; France; Belgium; Holland; Britain; Norway; Russia; Shetlands; Faroes; Greenland; Newfoundland; Germany). — Lohse, 1964: 49 ( Micralymma ; characters; central Europe). — Pototskaia, 1967: 19 ( Micralymma ; larval characters). — Steel, 1970a: 17 ( Micralymma ; larval characters; biological notes). — Pope, 1977: 24 ( Micralymma ; Britain). — Hammond, 1980: 139 ( Micralymma ; Ireland). — Frank, 1982: 9 ( Micralymma ; list of parasites). — Elliott, King, and Fordy, 1983 ( Micralymma ; habitat; natural history). — Thayer, 1985b ( Micralymma ; natural history notes; Newfoundland; Quebec; Maine; Massachusetts; New Hampshire). — Elliott and King, 1986 ( Micralymma ; adaptations for intertidal living). — Segers, 1986: 21 ( Micralymma ; checklist of species; Belgium). — Lucht, 1987: 88 ( Micralymma ; checklist; Germany; Benelux; Sweden). — Hicks, 1990: 304 ( Micralymma ; habitat preferences; Newfoundland). — Downie and Arnett, 1996: 436 ( Micralymma ; characters; Canada; USA). - DISTRIBUTION: Russia, Europe, Iceland, Greenland, Canada, USA.

brevipenne Gyllenhal, 1810: 234 ( Omalium ; Type locality: Sweden). — Zetterstedt, 1828: 56 ( Omalium ; characters; Lapland). — Zetterstedt, 1838: 54 ( Omalium ; characters; habitat; Lapland). — Erichson, 1840: 820 ( Micralymma ; characters; Sweden; England). — Schiødte, 1845: 376 ( Micralymma ; characters; Sweden; Norway; Scotland). — Schiødte, 1846: 162 ( Micralymma ; characters; Norway; Sweden; Scotland). — Küster, 1854: 22 ( Micralymma ; characters; Europe).

— Fairmaire and Laboulbène, 1856: 625 ( Micralymma ; characters; France).

— Laboulbène, 1858 ( Micralymma ; anatomy of adults and immatures).

— Staudinger, 1857: 286 ( Micralymma ; notes; Iceland).

— Thomson, 1859: 49 (type species of Micralymma ).

— Thomson, 1861: 188 ( Micralymma ; characters; Scandinavia).

— Fauvel, 1871a: 108 [= 1872: 82] ( Micralymma ; synonym of marinum ).

— Redtenbacher, 1874: 271 ( Micralymma ; characters).

— Mulsant and Rey, 1880: 6 ( Micralymma ; characters; France).

— Fowler, 1888: 407 ( Micralymma ; characters; Britain).

— Ganglbauer, 1895: 729 ( Micralymma ; synonym of marinum ).

— Bernhauer and Schubert, 1910: 62 ( Micralymma ; synonym of marinum ).

— Scheerpeltz, 1933: 1053 ( Micralymma ; synonym of marinum ).

— Steel, 1958: 140 ( Micralymma ; synonym of marinum ).

— Pope, 1977: 24 ( Micralymma ; synonym of marina ).

johnstonis Westwood, 1838b: 130 ( Micralymma ; includes larval and pupal characters; Type locality: Berwick-upon-Tweed).

— Westwood, 1838: 165 ( Micralymma ; habitat).

— Westwood, 1838a (August): 19 ( Micralymma ; characters; type species).

— Stephens, 1839: 427 ( Micralymma ; characters; England).

— Erichson, 1840: 820 ( Micralymma ; synonym of brevipenne ).

— Schiødte, 1846: 162 ( Micralymma ; synonym of brevipenne ).

— Küster, 1854: 22 ( Micralymma ; characters; Europe).

— Hardy, 1851: 53 ( Micralymma ; synonym of brevipenne ).

— Fairmaire and Laboulbène, 1856: 625 ( Micralymma ; synonym of brevipenne ).

— Mulsant and Rey, 1880: 6 ( Micralymma ; synonym of brevipenne ).

— Ganglbauer, 1895: 729 ( Micralymma ; synonym of marinum ).

— Bernhauer and Schubert, 1910: 62 ( Micralymma ; synonym of marinum ).

— Steel, 1958: 140 ( Micralymma ; synonym of marinum ).

stimpsonii LeConte, 1863 a: 57 ( Micralymma ; Type locality: coast of Maine).

— Fauvel, 1878d: 222 [= 1878e: 58] ( Micralymma ; characters; Maine).

— Bernhauer and Schubert, 1910: 62 ( Micralymma ; catalog).

— Scheerpeltz, 1933: 1053 ( Micralymma ; catalog).

— Steel, 1958: 140 ( Micralymma ; synonym of marinum ).

— Thayer, 1985b: 49 ( Micralymma ; synonym of marinum ).

stimpsonii LeConte, 1863 , see: marinum Ström, 1783 .

Nesomalium

[4 species; Antarctic Region]

Nesomalium Steel, 1964: 353 (species included: campbellense , pacificum , imitator , insulare ).

TYPE SPECIES: Nesomalium campbellensis Steel View in CoL , fixed by original designation.

— Steel, 1970a: 25 (larval characters).

— Moore and Legner, 1976: 536 (includes seashore species; notes).

— Klimaszewski, Newton, and Thayer, 1996: 146 (endemic to New Zealand and nearby islands). campbellense Steel, 1964: 354 View in CoL ( Nesomalium View in CoL ; also larval characters; Type locality: Campbell

Island: Moubray Hill, 200 m).

- DISTRIBUTION: Campbell Island.

imitator Steel, 1964: 358 View in CoL ( Nesomalium View in CoL ; Type locality: Campbell Island: Lookout Bay).

- DISTRIBUTION: Campbell Island.

insulare Kiesenwetter, 1877: 163 View in CoL ( Omalium View in CoL ; Type locality: Auckland Islands ).

— Fauvel, 1877: 189 ( Omalium View in CoL , cited as Homalium View in CoL ; characters; Auckland Islands).

— Bernhauer and Schubert, 1910: 54 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Gourlay, 1950: 183 ( Omalium View in CoL ; republication of original description).

— Steel, 1964: 357 ( Nesomalium View in CoL ; characters; Auckland Islands).

pacificum Kiesenwetter, 1877: 163 View in CoL ( Omalium View in CoL ; Type locality: Auckland Islands ).

— Fauvel, 1877: 190 ( Omalium View in CoL , cited as Homalium View in CoL ; characters; Auckland Islands).

— Fauvel, 1878g: 484 ( Omalium View in CoL , cited as Homalium View in CoL ; characters).

— Bernhauer and Schubert, 1910: 55 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Gourlay, 1950: 183 ( Omalium View in CoL ; subgenus Omaliomimus View in CoL ?; republication of original description). — Steel, 1964: 357 ( Nesomalium View in CoL ; characters; Auckland Islands; Enderby, Ocean, Ewing, and Rose Island).

- DISTRIBUTION: Auckland Islands.

zealandicum Cameron, 1948c: 723 View in CoL ( Omalium View in CoL ; Type locality: New Zealand: Auckland [Island]: Musgrave Pen; Carnley Harbour ).

— Gourlay, 1950: 184 ( Omalium View in CoL ; correction of Type locality; synonym of albipenne View in CoL ).

— Steel, 1964: 348,357 ( Nesomalium View in CoL ; Gourlay’s citation as synonym of Omalium albipenne View in CoL is erroneous; synonym of pacificum View in CoL ).

zealandicum Cameron, 1948 View in CoL , see: pacificum Kiesenwetter, 1877 View in CoL .

Nipponophloeostiba View in CoL

[1 species; Palaearctic Region]

Nipponophloeostiba Watanabe, 1962 View in CoL a: 77 (species included: verrucifera View in CoL ). TYPE SPECIES: Nipponophloeostiba verrucifera Watanabe View in CoL , fixed by original designation and monotypy. — Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Shibata, 1976: 106 (checklist of species of Japan).

— Watanabe, 1990: 106 (characters).

verrucifera Watanabe, 1962 View in CoL a: 78 ( Nipponophloeostiba View in CoL ; Type locality: Miyake Is.: near Tairoike).

— Watanabe and Shibata, 1972: 60 ( Nipponophloeostiba View in CoL ; Yaku-shima Island).

— Watanabe, 1990: 109 ( Nipponophloeostiba View in CoL ; characters; Japan).

— Kishimoto, 1995: 76 ( Nipponophloeostiba View in CoL ; Ryukyu Islands).

- DISTRIBUTION: Japan.

Noumalia View in CoL

[1 species; Oceanic Region]

Noumalia Steel, 1960: 5 View in CoL (species included: adusta ). TYPE SPECIES: Noumalia adusta Steel , fixed by original designation and monotypy.

adusta Steel, 1960: 6 ( Noumalia View in CoL ; Type locality: New Caledonia: Mt. Ignambi ).

- DISTRIBUTION: New Caledonia.

Omaliomimus View in CoL

[10 species; Australian and Antarctic Regions]

Omaliomimus Jeannel, 1940: 115 View in CoL , 117 (species included: litoreus View in CoL , setipes View in CoL , conicus View in CoL , variipenne View in CoL ). TYPE SPECIES: Omalium litoreum Broun , by original designation.

— Blackwelder, 1952: 273 (type species: litoreus View in CoL ).

— Steel, 1964: 344 (characters; key to species of Campbell Island).

— Steel, 1970a: 24 (larval characters).

— Moore and Legner, 1976: 536 (includes seashore species; notes).

— Thayer, 1996: 158 (notes).

— Klimaszewski, Newton, and Thayer, 1996: 146 (endemic to New Zealand and nearby islands). actobius Broun, 1893: 1035 View in CoL ( Omalium View in CoL ; Type locality: Sandspit, Port Chalmers).

— Bernhauer and Schubert, 1910: 51 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Newton and Thayer, 1995: 293 ( Omaliomimus View in CoL ).

— Thayer, 1996: 159 ( Omaliomimus View in CoL ).

- DISTRIBUTION: New Zealand.

albipennis Kiesenwetter, 1877: 162 View in CoL ( Omalium View in CoL ; Type locality: Auckland Island ).

— Fauvel, 1877: 188 ( Omalium View in CoL , cited as Homalium View in CoL ; characters; Auckland Island).

— Gourlay, 1950: 184 ( Omalium View in CoL ; republication of original description).

— Steel, 1964: 347 ( Omaliomimus View in CoL ; characters; Campbell Island).

— Watson, 1967: 34 ( Omaliomimus View in CoL ; notes; mention of existence of larva; habitat; Macquarie Island).

— P. Greenslade, 1990: 48 (Omaliominus; wingless species; Macquarie Island).

— Klimaszewski and Watt, 1997: 169 ( Omaliomimus View in CoL ; characters; collecting notes; New Zealand). - DISTRIBUTION: Auckland Island, Macquarie Island, Campbell Island, New Zealand. variipennis Lea, 1920: 30 View in CoL ( Omalium View in CoL , cited as Homalium View in CoL ; Type locality: Macquarie Island). — Tillyard, 1920: 19 ( Omalium View in CoL , cited as Homalium View in CoL ; [Note: Cited as new species but attributed to Lea in appendix of same article]; illustrations of adult; larval characters and adults; Macquarie Island).

— Scheerpeltz, 1933: 1048 ( Omalium View in CoL ; catalog).

— Jeannel, 1940: 117 ( Omaliomimus View in CoL ).

— Steel, 1964: 347 ( Omaliomimus View in CoL ; synonym of albipenne View in CoL ).

— Watson, 1967: 34 ( Omaliomimus View in CoL ; synonym of albipenne View in CoL ).

— P. Greenslade, 1990: 48 (Omaliominus; synonym of albipenne View in CoL ).

flavipennis Cameron, 1948c: 723 View in CoL ( Omaliomimus View in CoL ; Type locality: New Zealand: Auckland: Port Ross ).

— Gourlay, 1950: 183 ( Omalium View in CoL ; provisional synonym of pacificum View in CoL ).

— Steel, 1964: 347 ( Omaliomimus View in CoL ; synonym of albipenne View in CoL ; Gourlay’s citation as synonym of pacificum View in CoL is erroneous).

— Watson, 1967: 34 ( Omaliomimus View in CoL ; synonym of albipenne View in CoL ).

— P. Greenslade, 1990: 48 (Omaliominus; synonym of albipenne View in CoL ).

carinigerus Broun, 1893: 1036 ( Omalium View in CoL ; Type locality: Port Chalmers ) .

— Bernhauer and Schubert, 1910: 52 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Thayer, 1996: 159 ( Omaliomimus View in CoL ).

- DISTRIBUTION: New Zealand.

chalmeri Broun, 1893: 1037 View in CoL ( Omalium View in CoL ; Type locality: Dunedin).

— Bernhauer and Schubert, 1910: 52 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Thayer, 1996: 159 ( Omaliomimus View in CoL ).

- DISTRIBUTION: New Zealand.

conicus Fauvel, 1878 View in CoL g: 484 ( Omalium View in CoL , cited as Homalium View in CoL ; Type locality: Nouvelle-Zélande). — Bernhauer and Schubert, 1910: 52 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Jeannel, 1940: 117 ( Omaliomimus View in CoL ).

- DISTRIBUTION: New Zealand.

flavipennis Cameron, 1948 View in CoL , see: albipenne Kiesenwetter, 1877 View in CoL .

laetipennis Broun, 1910: 19 View in CoL ( Omalium View in CoL ; Type locality: Riverton Beach).

— Scheerpeltz, 1933: 1045 ( Omalium View in CoL ; catalog).

— Thayer, 1996: 159 ( Omaliomimus View in CoL ).

- DISTRIBUTION: New Zealand.

litoreus Broun, 1886: 942 View in CoL ( Omalium View in CoL , cited as Homalium View in CoL ; Type locality: Mokohinau).

— Bernhauer and Schubert, 1910: 54 ( Omalium View in CoL ; subgenus Omalium View in CoL ; cited as littoreum View in CoL ; catalog). — Jeannel, 1940: 115, 117 ( Omaliomimus View in CoL ).

— Kuschel, 1990: 25 ( Omaliomimus View in CoL ; endemic; winged; habitat; New Zealand).

- DISTRIBUTION: New Zealand.

robustus Broun, 1911: 96 View in CoL ( Omalium View in CoL ; [preoccupied]; Type locality: Pitt Island ).

— Scheerpeltz, 1933: 1047 ( Omalium View in CoL ; catalog).

— Emberson, 1998: 33 ( Omaliomimus View in CoL ; collecting notes; Chatham Islands).

— Herman, 2001: 37 ( Omaliomimus View in CoL ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Chatham Islands (N.Z).

setipes Broun, 1909: 230 View in CoL ( Omalium View in CoL ; Type locality: Invercargill).

— Scheerpeltz, 1933: 1047 ( Omalium View in CoL ; catalog).

— Jeannel, 1940: 117 ( Omaliomimus View in CoL ).

- DISTRIBUTION: New Zealand.

variipenne Lea, 1920 View in CoL , see: albipenne Kiesenwetter, 1877 View in CoL .

venator Broun, 1909 View in CoL a: 98 ( Omalium View in CoL ; Type locality: Campbell Island ).

— Scheerpeltz, 1933: 1048 ( Omalium View in CoL ; catalog).

— Steel, 1964: 345 ( Omaliomimus View in CoL ; characters; Campbell Island; Auckland Island; Macquarie Island).

— Watson, 1967: 35 ( Omaliomimus View in CoL ; notes; habitat; Macquarie Island).

— P. Greenslade, 1990: 48 (Omaliominus; wingless species; Macquarie Island).

— Marris, 2000: 175 ( Omaliomimus View in CoL ; habitat; Bollons Island [Antipodes]; also known from Auckland Islands; Campbell Island; Macquarie Island; Snares Islands; New Zealand).

- DISTRIBUTION: Campbell Island, Auckland Island, Macquarie Island, Snares Islands, Antipodes Islands, New Zealand.

Omaliopsis View in CoL

[8 species; Ethiopian, Madagascan, Nearctic , and Neotropical Regions]

Omaliopsis Jeannel, 1940: 116 View in CoL , 118 (species included: africana , costipennis View in CoL , arenaria View in CoL , rufa View in CoL , madecassus View in CoL , hova View in CoL ). TYPE SPECIES: Omalium africanum Fauvel , by original designation.

— Blackwelder, 1952: 273 (type species: africanus View in CoL ).

— Fagel, 1956: 2 (synonym of Omalium View in CoL ).

— Coiffait and Saiz, 1968: 451 (valid genus; characters).

— Newton, 1985: 199 (notes on austral distribution).

— Newton, Thayer, Ashe, and Chandler, 2000: 337 (1 undescribed Nearctic species; notes; characters in key).

africana Fauvel, 1907: 12 ( Omalium View in CoL , cited as Homalium View in CoL ; Type locality: Kilimandjaro: Kilema).

— Bernhauer and Schubert, 1910: 51 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Bernhauer, 1940: 130 ( Omalium View in CoL ; Uganda; Kenya).

— Jeannel, 1940: 116, 118 ( Omaliopsis View in CoL ).

— Fagel, 1956: 11 ( Omalium View in CoL ; characters; German East Africa).

- DISTRIBUTION: Tanzania, Uganda, Kenya.

arenaria Bernhauer, 1904: 16 View in CoL ( Omalium View in CoL ; Type locality: Kap der guten Hoffnung).

— Bernhauer and Schubert, 1910: 51 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Jeannel, 1940: 118 ( Omaliopsis View in CoL ).

- DISTRIBUTION: South Africa.

costipennis Fauvel, 1904e: 301 View in CoL ( Omalium View in CoL , cited as Homalium View in CoL ; Type locality: Tananarive).

— Bernhauer and Schubert, 1910: 52 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Jeannel, 1940: 118 ( Omaliopsis View in CoL ).

- DISTRIBUTION: Madagascar.

flavescens Motschulsky, 1857 View in CoL , see: rufum Sachse, 1852 View in CoL .

hova Fauvel, 1904e: 300 View in CoL ( Omalium View in CoL , cited as Homalium View in CoL ; Type locality: Tananarive).

— Bernhauer and Schubert, 1910: 54 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Jeannel, 1940: 118 ( Omaliopsis View in CoL ; cited as howa View in CoL ).

- DISTRIBUTION: Madagascar.

madegassa Bernhauer, 1902: 173 View in CoL ( Omalium View in CoL ; Type locality: Port Daufin , Madagaskar).

— Bernhauer and Schubert, 1910: 54 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Jarrige, 1957: 112 ( Omaliopsis View in CoL ; Réunion).

- DISTRIBUTION: Madagascar, Réunion.

picipennis Fauvel, 1867 View in CoL a: 44 [= 1868: 49] ( Omalium View in CoL ; Type locality: Chili).

— Bernhauer and Schubert, 1910: 55 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Coiffait and Saiz, 1968: 452 ( Omaliopsis View in CoL ; characters; Chile).

- DISTRIBUTION: Chile.

rufa Sachse, 1852: 148 View in CoL ( Omalium View in CoL ; [preoccupied]; Type locality: Vorgeb. d. g. Hoffnung).

— Bernhauer and Schubert, 1910: 56 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Jeannel, 1940: 118 ( Omaliopsis View in CoL ).

— Herman, 2001: 37 ( Omaliopsis View in CoL ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: South Africa.

flavescens Motschulsky, 1857 View in CoL a: 492 ( Phloeonomus View in CoL ; Type locality: Cap de bonne Esperance).

— Bernhauer and Schubert, 1910: 56 ( Omalium View in CoL ; synonym of rufum View in CoL ).

russata Fairmaire and Germain, 1861: 453 View in CoL ( Omalium View in CoL ; Type locality: Chiloé).

— Fauvel, 1867a: 45 [= 1868: 50] ( Omalium View in CoL ; characters; Chile).

— Bernhauer and Schubert, 1910: 56 ( Omalium View in CoL ; subgenus Omalium View in CoL ; catalog).

— Scheerpeltz, 1933: 1047 ( Omalium View in CoL ; catalog).

— Coiffait and Saiz, 1968: 451 ( Omaliopsis View in CoL ; characters; Chile).

— Saiz, 1969: 4 ( Omaliopsis View in CoL ; characters; collecting notes; Chile).

- DISTRIBUTION: Chile.

Omalium View in CoL

[141 species; (11 doubtful); Neotropical, Nearctic, Palaearctic, Ethiopian, Madagascan , Oriental, and Australian Regions]

Omalium Gravenhorst, 1802: 111 View in CoL (species included: planum , depressum View in CoL , brunneum View in CoL , crenatum View in CoL , brachypterum View in CoL , rufum View in CoL , rugosum View in CoL , rivulare , viburni View in CoL , florale, ranunculi View in CoL , striatum View in CoL ). TYPE SPECIES: Staphylinus rivularis Paykull View in CoL , fixed by subsequent designation by Latreille, 1810: 427.

— Latreille, 1804: 286 (characters).

— Gravenhorst, 1806: 204 (characters).

— Latreille, 1806: 297 (characters).

— Latreille, 1810: 182, 427 (characters; type species).

— Gyllenhal, 1810: 198 (characters).

— Olivier, 1811: 472 (characters).

— Leach, 1815: 91 (characters).

— Samouelle, 1819: 174 (characters).

— Billberg, 1820: 14 (cited as Homalium View in CoL ; characters; list of some species).

— Dejean, 1821: 26 (list of species in collection).

— Latreille, 1825: 245 (cited as Omalie; characters).

— Berthold, 1827: 332 (characters).

— C. Sahlberg, 1827: 277 (characters).

— Zetterstedt, 1828: 48 (characters).

— Curtis, 1829: 29 (catalog; Britain).

— Stephens, 1829a: 25 (catalog; Britain).

— Stephens, 1829: 297 (catalog; Britain).

— Latreille, 1829: 439 (characters).

— Mannerheim, 1830: 10, 51 (characters).

— Mannerheim, 1830: 10, 53 (key; characters; [Note: Misidentification: Bernhauer and Schubert, 1910: 50 cited Anthobium Mannerheim View in CoL as a synonym of Omalium View in CoL ; Mannerheim did not describe the genus as new, he attributed it to Leach]).

— Mannerheim, 1831: 424, 465 (characters).

— Mannerheim, 1831: 424, 467 (characters; [Note: Misidentification: Bernhauer and Schubert, 1910: 50 cited Anthobium Mannerheim View in CoL as a synonym of Omalium View in CoL ; Mannerheim did not describe the genus as new, he attributed it to Leach]).

— Dejean, 1833: 69 (list of species).

— Stephens, 1834: 345 (characters; England).

— Gistel, 1834: 9 (list of species).

— Lacordaire, 1835: 484 (characters).

— Westwood, 1838a: 18 (characters; type species).

— Zetterstedt, 1838: 49 (characters).

— Erichson, 1839a: 628 (characters).

— Heer, 1839: 175 (characters).

— Stephens, 1839: 422 (characters).

— Erichson, 1840: 874 (characters).

— Laporte, 1840: 191 (characters).

— Redtenbacher, 1849: 750 (characters).

— Schaum, 1852: 30 (list of species; Europe).

— Lacordaire, 1854: 143 (characters; notes; list of species).

— Wollaston, 1854: 612 (characters; notes).

— Fairmaire and Laboulbène, 1856: 638 (characters).

— Giebel, 1856: 74 (fossil species).

— Gistel, 1856: 389 (cited as Homalium View in CoL ; list of species).

— Jacquelin du Val, 1857: 76 (characters).

— Redtenbacher, 1857: 249 (characters).

— Kraatz, 1857d: 974 (characters).

— Thomson, 1858: 38 (characters).

— G. Waterhouse, 1858: 31 (catalog of British species).

— Thomson, 1859: 51 (characters; type species: rivulare ).

— Schaum, 1859: 32 (catalog; European species).

— LeConte, 1861a: 70 (characters).

— Thomson, 1861: 211 (characters).

— Gredler, 1863: 124 (list of species from Tirol).

— LeConte, 1863: 26 (list of North American species).

— Fauvel, 1865c: 5 [= 1866: 13] (characters; list of species).

— Wencker and Silbermann, 1866: 36 (list of species; collecting notes; France).

— Fauvel, 1867a: 43 [= 1868: 48] (characters; key to Chilean species).

— Jacquelin du Val, 1868: 81 (catalog; European species).

— Sharp, 1871d: 14 (cited as Homalium View in CoL ; list of British species).

— Fauvel, 1871a: 83 [= 1872: 57] (characters; key to species).

— Redtenbacher, 1874: 273 (characters; key to Austrian species).

— Fauvel, 1875a: [= 1875b:] (catalog; cited as Homalium View in CoL ).

— Sharp, 1876: 402 (notes).

— Provancher, 1877: 261 (characters).

— Fauvel, 1878d: 205 [= 1878e: 41] (cited as Homalium View in CoL ; key to North American species). — Broun, 1880: 115 (characters).

— Mulsant and Rey, 1880: 189 (characters; key to French species).

— Heyden, 1880: 82 (cited as Homalium View in CoL ; list of species of Siberian region).

— Duvivier, 1883: 198 (catalog; cited as Homalium View in CoL ).

— Gozis, 1886: 16 (type species).

— Sharp, 1887: 744 (notes; Mexican and Central American species).

— Fowler, 1888: 409 (notes; cited as Homalium View in CoL ; key to British species).

— Seidlitz, 1891: 339 (key to some European species).

— Seidlitz, 1891a: 362 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 120 (list of species of Europe and Caucasus).

— Heyden, 1893: 49 (list of species of Siberian region).

— Ganglbauer, 1895: 733 (characters; key to middle European species).

— Heyden, 1896: 37 (list of species of Siberian region).

— Fauvel, 1897c: 230 (key to species).

— Fauvel, 1897d: 244 (catalog of species of Barbary; Azores; Madeira; Canaries). — Heyden, Reitter, and Weise, 1906: 131 (list of species of Europe and Caucasus). — Sainte-Claire Deville, 1907: 32 (cited as Homalium View in CoL ; key to and annotated list of species of

Seine Basin, France). — Broun, 1909a: 97 (characters). — Reitter, 1909: 190 (characters; key to German species). — Bernhauer and Schubert, 1910: 50 (world catalog; 120 species). — Blatchley, 1910: 479 (characters; key to Indiana species). — Petri, 1912: 53 (list of species of Siebenbürgen). — Johansen, 1914: 608 (characters; key to Danish species). — Leng, 1920: 94 (catalog of North American species). — Winkler, 1925: 330 (catalog for Palaearctic region). — Cameron, 1925: 10, 104 (catalog of Indian species). — Porta, 1926: 18 (characters; key to species of Italy). — Portevin, 1929: 445 (characters; key to French species). — Cameron, 1930: 138 (characters; key to British Indian species). — Roubal, 1930: 307 (catalog; Slovakia). — Scheerpeltz, 1933: 1043 (world catalog supplement). — Normand, 1935: 357 (list of species of Tunisia). — Scheerpeltz, 1937b: 187 (list of Bulgarian species). — Jeannel, 1940: 115 (characters in key; type species: rivulare Paykull ). — Blackwelder, 1943: 53 (characters; type species: rivulare ; key to species of West Indies). — Blackwelder, 1944: 101 (checklist of species from Mexico, Central and South America View in CoL , and the West Indies). — Jeannel and Jarrige, 1949: 311 (cave species; type species: rivulare ). — Kloet and Hincks, 1945: 161 (list of British species). — Scheerpeltz and Höfler, 1948: 152 (key to species in fungus in Europe). — Palm, 1948: 75 (characters; key to species of Sweden). — Tottenham, 1949: 355, 406 (type species: rivularis Paykull View in CoL ; list of British species; notes). — Blackwelder, 1952: 273 (type species). — Tottenham, 1954: 23 (characters; key to British species). — Hatch, 1957: 52, 80 (characters; key to species of Pacific Northwest). — Kocher, 1958: 93 (checklist of species; Morocco). — Smetana, 1959a: 196 (list of species collected in Albania). — Ferreira, 1962a: 9 (list of Portuguese species). — Székessy, 1963: 14 (characters). — Lohse, 1964: 41 (characters; key to central European species). — Pototskaia, 1967: 20 (larval characters). — Kasule, 1968: 133, 136 (larval characters). — Scheerpeltz, 1968a: 9 (catalog of Austrian species). — Bolov, 1969a: 328 (collecting notes for some species; Russia). — Steel, 1970a: 13 (larval characters; biological notes). — Bordoni, 1973j: 656 (list of species from islands near Sicily). — Tikhomirova, 1973a: 138 (checklist of species of USSR). — Moore and Legner, 1975: 201 (catalog; North America View in CoL north of Mexico). — Shibata, 1976: 101 (checklist of species of Japan). — Moore and Legner, 1976: 536 (includes seashore species; notes). — Pope, 1977: 23 (list of British species). — Topp, 1978: 308 (larval characters). — Burakowski, Mroczkowski, and Stefańska, 1979: 38 (catalog; Poland). — Moore and Legner, 1979: 208 (characters; notes). — Muona, 1979: 18 (list of Scandinavian species). — Uhlig, 1979: 242 (some species collected in Germany). — Uhlig, Vogel, and Sieber, 1980: 240 (some species collected in Germany). — Waterston, Owen, Welch, Bacchus and Hammond, 1981: 273 (list of species of Outer

Hebrides).

— Uhlig and Vogler, 1981: 85 (list of some species collected in Germany; habitat and collecting notes).

— Spahr, 1981: 95 (references to specimens from amber and copal).

— Tóth, 1982: 57 (characters; habitus; key to species of Hungary).

— Zanetti, 1985: 27 (key to some Italian species).

— Ádám, 1985: 251 (list of species from southeast Hungary).

— Frank, 1986a: 365 (checklist of species of Florida).

— Muona and Viramo, 1986: 15 (list of species of northeastern Finland).

— Segers, 1986: 20 (checklist of species; Belgium).

— Ádám, 1987: 137 (list of four species with collecting notes; Hungary).

— Zanetti, 1987: 205 (characters; habitus; species group classification; key to species of Italy).

— Watanabe, 1990: 88 (characters; key to species of Japan).

— Dettner and Reissenweber, 1991 (defensive secretions: chemistry and systematics).

— Frank, 1991: 344 (larval characters).

— Campbell and Davies, 1991: 88 (checklist of species; Alaska and Canada).

— Silfverberg, 1992: 21 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Siitonen, 1993: 230 (list of Finnish species).

— Welch, 1993 (ovariole structure and development).

— Boháč, 1993: 41 (list of species of Czech Republic and Slovakia).

— Hernández, Outerelo, and Gamarra, 1994: 187 (checklist of species; Canary Islands).

— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 27 (additional records to Danish list of species).

— Zanetti, 1995: 7 (list of species of Italy).

— M. Hansen, 1996: 94 (list of Danish species).

— M. Hansen, Mahler, Palm, and Pedersen, 1996: 240 (additions to list of Danish species).

— Downie and Arnett, 1996: 437 (characters; key to species of Northeastern North America View in CoL ).

— Klimaszewski, Newton, and Thayer, 1996: 146 (species in New Zealand and nearby islands probably belong in another genus).

— M. Hansen, Palm, Pedersen, and Runge, 1998: 69 (list of species collected in 1997; Denmark).

— Lundgren, 1998: 38 (list of species from Florida; records for other States listed for each species).

— Newton, Thayer, Ashe, and Chandler, 2000: 337 (14 Nearctic species; notes; characters in key).

Homalium Ljungh, 1804 View in CoL a: 74 (emendation of Omalium View in CoL ). TYPE SPECIES: Staphylinus rivularis Paykull View in CoL , fixed by objective synonymy with Omalium View in CoL .

— Blackwelder, 1952: 188 (type species).

Homalium Agassiz, 1847: 258 View in CoL (emendation of Omalium View in CoL ). TYPE SPECIES: Staphylinus rivularis Paykull View in CoL , fixed by objective synonymy with Omalium View in CoL .

— Blackwelder, 1952: 188 (type species).

Homalium Gemminger and Harold, 1868: 665 View in CoL (emendation of Omalium View in CoL ). TYPE SPECIES: Staphylinus rivulare Paykull , fixed by objective synonymy with Omalium View in CoL .

— Blackwelder, 1952: 189 (type species).

Ochthexenus Motschulsky, 1860: 546 View in CoL (species included: clavicornis View in CoL , rivularis View in CoL ). TYPE SPECIES: Staphylinus rivularis Paykull View in CoL , fixed by subsequent designation by Gozis, 1886: 16.

— Gemminger and Harold, 1868: 665 (catalog).

— Fauvel, 1869: 493 (synonym of Omalium View in CoL ).

— Heyden, 1880: 82 (subgenus of Omalium View in CoL , cited as Homalium View in CoL ; list of species of Siberian region).

— Gozis, 1886: 16 (type species).

— Bernhauer and Schubert, 1910: 50 (synonym of Omalium View in CoL ).

— Cameron, 1930: 138 (synonym of Omalium View in CoL ).

— Blackwelder, 1943: 53 (synonym of Omalium View in CoL ; type species: clavicornis View in CoL ).

— Blackwelder, 1952: 266 (type species: rivularis View in CoL ).

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