Geodromicus subnigritus, Lohse, 1958

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

subnigritus Lohse, 1958 [Note: Nomen nudum], see: plagiatus Fabricius, 1798 .

subsimilis Cameron, 1941: 144 ( Geodromicus ; Type locality: Ghum district : Mangpu; Tiger Hill, 8000– 10,000 feet). - DISTRIBUTION: India.

suensoni Bernhauer, 1936 g: 304 ( Geodromicus ; Type locality: Japan: Kamikochi, North Alps 5000 ft). — Watanabe, 1990: 293 ( Psephidonus ; characters; Japan). - DISTRIBUTION: Japan.

suturalis Lacordaire, 1835: 483 ( Lesteva ; Type locality: Paris). — Erichson, 1840: 852 ( Anthophagus ; synonym of plagiatus ). — Erichson, 1840: 852 ( Anthophagus ; variety b; [Note: Bernhauer and Schubert, 1910: 77 cited plagiatus variety b Erichson as synonym of suturalis ; there is no available name]; characters). — Kraatz, 1857d: 926 ( Anthophagus ; [Note: Bernhauer and Schubert, 1910: 77 cited plagiatus variety a Kraatz as a synonym of suturalis but there is no available name]; characters; Germany). — Jacquelin du Val, 1857: pl. 25, fig. 123 ( Geodromicus ; [Note: Misidentification, cited by Bernhauer and Schubert, 1910: 77 as synonym of suturalis but Jacquelin du Val attributed the species to Fabricius]; illustration). — Fauvel, 1865a: 315 ( Anthophagus ; habitat; France). — Fauvel, 1871a: 133 [= 1872: 107] ( Geodromicus ; variety of plagiatus ; [Note: Bernhauer and Schubert, 1910: 77 cited plagiatus variety α as a synonym of suturalis ; there is no available name]; characters; France; Switzerland; Austria). — Mulsant and Rey, 1880: 57 ( Geodromicus ; synonym of plagiatus ). — Ganglbauer, 1895: 711 ( Geodromicus ; characters; Alps; Pyrenees). — Luze, 1903: 107 ( Geodromicus ; characters; Alps; Pyrenees). — Reitter, 1909: 183 ( Geodromicus ; characters; Germany). — Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog). — Scheerpeltz, 1933: 1066 ( Geodromicus ; catalog). — Fagel, 1958b: 224 ( Geodromicus ; characters; Alpes and Pyrenees). — Smetana, 1959c: 359 ( Geodromicus ; characters; notes; Czechoslovakia). — Horion, 1963: 134 ( Geodromicus ; France; Italy; Switzerland; Germany; Austria; Romania; Czech Republic). — Lohse, 1964: 61 ( Geodromicus ; characters; central Europe). — Tóth, 1982: 102 ( Psephidonus ; characters). — Bordoni, 1984a: 27 ( Geodromicus ; notes). — Zanetti, 1987: 375 ( Psephidonus ; subgenus Psephidonus ; characters; Italy). — Lucht, 1987: 90 ( Geodromicus ; checklist; Germany; Austria; Switzerland). — Boháč, 1993: 41 ( Psephidonus ; Slovakia; Czech Republic). — Zanetti, 1995: 9 ( Geodromicus ; Italy). — Kashcheev, 1999c: 16 ( Geodromicus ; characters in key). - DISTRIBUTION: France, Switzerland, Germany, Czechoslovakia, Austria, Italy.

concolor Luze, 1903: 107 ( Geodromicus ; aberration of suturalis ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; no locality cited). — Bernhauer and Schubert, 1910: 77 ( Geodromicus ; aberration of suturalis ). — Scheerpeltz, 1933: 1066 ( Geodromicus ; synonym of suturalis ). — Tóth, 1982: 102 ( Psephidonus ; aberration of suturalis ; characters). — Zanetti, 1987: 376 ( Psephidonus ; synonym of suturalis ).

svaneticus Iablokov-Khnzorian, 1989: 134 ( Geodromicus ; Type locality: GSSR: Svanetia, okr. Mestii). - DISTRIBUTION: Georgia.

szechwanensis Bernhauer, 1943: 73 ( Geodromicus ; Type locality: China). - DISTRIBUTION: China: Szechwan mer., Mts. Kinfushan, 2000 m, am Flusse Sung-Kanho.

temporalis Casey, 1894: 410 ( Geodromicus ; Type locality: California: Sonoma Co. ).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog).

— Hatch, 1957: 70 ( Geodromicus ; characters; British Columbia ; Washington; Oregon).

- DISTRIBUTION: Canada, USA.

torrentis Puthz, 1973 j: 517 ( Geodromicus ; Type locality: Pakistan: Swat Region : Guhral

Spring).

- DISTRIBUTION: Pakistan.

transversus Motschulsky, 1860 , see: Trichodromeus .

trapezipennis Coiffait, 1979d: 563 ( Geodromicus ; Type locality: Nouristan, vallée de l’Agôr

Khwar, 2000–3000 m).

- DISTRIBUTION: Afghanistan.

trisulcatus Coiffait, 1983: 278 ( Geodromicus ; Type locality: Cachemire: Pahalgam, Kolakoi,

3400 m).

- DISTRIBUTION: India.

tumidus Luze, 1903 , see: Trichodromeus .

verticalis Say, 1831 , see: plagiatus Fabricius, 1798 .

yushanensis Hayashi, 1992 , see: ohkurai Hayashi, 1992 .

ziganensis Coiffait, 1978 , see: jarrigeanus Fagel, 1976 .

zwickianus Fagel, 1976: 387 ( Geodromicus ; Type locality: S.O. Anatolien: Ost Taurus bei

Sürgü).

— Bordoni, 1984a: 37 ( Geodromicus ; cited as zwikianus; Turkey).

- DISTRIBUTION: Turkey.

Hygrodromicus

[14 species; Palaearctic Region] Hygrodromicus Tronquet, 1981: 72 (species included: obesus ; [Note: When Tronquet published his new species, Hygrodromicus obesus , the generic name was cited in a footnote as “ Hygrodromicus Coiffait , genre nouveau (sous presse)” but Tronquet published a few generic characters at the end of his description of obesus thereby validating the name. Tronquet’s use of the generic name appeared on September 9,

1981 and Coiffait’s on October 30, 1981]). TYPE SPECIES: Hygrodromicus obesus

Tronquet, fixed by monotypy.

— Zerche, 1992a: 107, 112, 120, 139 (key to species; phylogenetic relationships). Hygrodromicus Coiffait, 1981c: 258 , 259 (species included: meurguesae ). TYPE SPECIES:

Hygrodromicus meurguesae Coiffait , fixed by original designation and monotypy.

— Bordoni, 1990a: 100 (characters).

— Zerche, 1992a: 107 (synonym of Hygrodromicus Tronquet ). behnei Zerche, 1992 a: 117 ( Hygrodromicus ; Type locality: Asia cent., Dushanbe, Tadshik.,

Warsob-Tal/Hissar-Geb., 2000–3000 m).

- DISTRIBUTION: Tajikistan. cachemiricus Coiffait, 1983: 286 ( Hygrodromicus ; Type locality: Cachemire: Badrinath,

3500).

- DISTRIBUTION: India. coriaceus Cameron, 1924b: 172 , 173 ( Geodromicus ; Type locality: Chakrata District; Chulli

Khud and Binal Gad, 7000 to 8000 feet).

— Cameron, 1930: 162 ( Geodromicus ; characters; India).

— Scheerpeltz, 1933: 1065 ( Geodromicus ; catalog).

— Zerche, 1992a: 120 ( Hygrodromicus ).

- DISTRIBUTION: India. klapperichi Coiffait, 1982b: 83 ( Hygrodromicus ; Type locality: Afghanistan: Nouristan: ledouxi Coiffait, 1983: 285 ( Hygrodromicus ; Type locality: Cachemire: Himachal Pradesh,

Mari, 3500 m).

- DISTRIBUTION: India. lestevoides Reitter, 1900 , see: reitteri Scheerpeltz, 1933 . meurguesae Coiffait, 1981c: 260 ( Hygrodromicus ; Type locality: Cachemire, Badrinath,

3500 m).

— Coiffait, 1983: 275 ( Hygrodromicus ; Kashmir).

- DISTRIBUTION: India. obesus Tronquet, 1981: 71 ( Hygrodromicus ; Type locality: Afghanistan: versant nord du col de Salang, 2500 m, Hindu-Kush).

- DISTRIBUTION: Afghanistan. ovalis Luze, 1904: 78 ( Geodromicus ; Type locality: Turkestan: Jagnob Kol; [Note: See lectotype designation by Zerche, 1992a]. Also cited from Iskander-kul: Karakul-darja).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog).

— Zerche, 1992a: 119 ( Hygrodromicus ; lectotype designation: TRKST. Jagnob Kol; characters;

Uzbekistan; Tajikistan).

- DISTRIBUTION: Uzbekistan, Tajikistan. palliditarsis Coiffait, 1983: 287 ( Hygrodromicus ; Type locality: Cachemire: Gangria,

3500 m).

- DISTRIBUTION: India. punctatissimus Coiffait, 1983: 286 ( Hygrodromicus ; Type locality: Cachemire: Gangria,

3500 m).

- DISTRIBUTION: India. reitteri Scheerpeltz, 1933: 1066 ( Geodromicus ; replacement name for lestevoides Reitter ).

— Zerche, 1992a: 114 ( Hygrodromicus ; characters; Turkestan; Tadzhikistan).

- DISTRIBUTION: Tajikistan, Uzbekistan. lestevoides Reitter, 1900b: 228 ( Geodromicus ; [preoccupied]; Type locality: Bokhara,

Turkestan; [Note: See lectotype designation by Zerche, 1992a]).

— Luze, 1903: 116 ( Geodromicus ; characters; Russian central Asia: Buchara).

— Scheerpeltz, 1933: 1066 ( Geodromicus ; synonym of reitteri ).

— Zerche, 1992a: 107, 114 ( Hygrodromicus ; lectotype designation: Bokhara, Turkestan; synonym of reitteri ). splendidus Zerche, 1992 a: 116 ( Hygrodromicus ; Type locality: USSR: Tadzhikistan, Pamir,

Muksu-Gebiet, Hochfläche bei Kishlak Ljarsch, 3000–4000 m).

- DISTRIBUTION: Tajikistan. wittmeri Coiffait, 1984: 145 ( Hygrodromicus ; Type locality: Cachemire: Pahalgam,

2200–3000 m).

- DISTRIBUTION: India. wrasei Zerche, 1992 a: 113 ( Hygrodromicus ; Type locality: Asia centr., Duschanbe, Tad-

schikistan, Warsob-Tal/Hissar-Gebirge, 2000–2500 m).

- DISTRIBUTION: Tajikistan.

Hygrogeus

[12 species; Palaearctic Region]

Hygrogeus Mulsant and Rey, 1880: 56 (species included: aemulus ). TYPE SPECIES: Anthophagus aemulus Rosenhauer , fixed by monotypy.

— Heyden, Reitter, and Weise, 1891: 118 (subgenus of Geodromicus ; list of species of Europe and Caucasus).

— Ganglbauer, 1895: 710 (characters).

— Luze, 1902d: 529 (characters).

— Heyden, Reitter, and Weise, 1906: 135 (list of species of Europe and Caucasus).

— Cameron, 1925: 104 (catalog of Indian species).

— Porta, 1926: 30 (characters; key to species of Italy).

— Cameron, 1930: 165 (characters; key to species of British India).

— Scheerpeltz, 1933: 1067 (world catalog supplement).

— Blackwelder, 1952: 193 (type species: aemulus ).

— Horion, 1963: 137 (list of central European species).

— Lohse, 1964: 63 (characters; key to central European species).

— Scheerpeltz, 1968a: 17 (catalog of Austrian species).

— Tikhomirova, 1973a: 141 (checklist of species of USSR).

— Zanetti, 1987: 384 (characters; habitus; species of Italy).

— Zerche, 1992a: 139 (phylogenetic relationship).

— Zanetti, 1995: 9 (list of species of Italy).

aemulus Rosenhauer, 1847: 12 ( Anthophagus ; Type locality: Steinach in Tyrol).

— Kraatz, 1857d: 929 ( Anthophagus ; characters; Austria).

— Fauvel, 1865a: 316 [= 1865b: 66] ( Anthophagus ; characters; France).

— Fauvel, 1871a: 139 [= 1872: 113] ( Anthophagus ; characters; France; Italy).

— Redtenbacher, 1874: 266 ( Geodromicus ; characters; Austria).

— Fauvel, 1876a: 54 [= 1876: 239] ( Anthophagus ; Switzerland).

— Mulsant and Rey, 1880: 61 ( Geodromicus ; subgenus Hygrogeus ; characters; France).

— Ganglbauer, 1895: 710 ( Hygrogeus ; characters; Alps).

— Luze, 1902d: 529 ( Hygrogeus ; characters; Alps).

— Bernhauer and Schubert, 1910: 78 ( Hygrogeus ; catalog).

— Portevin, 1929: 433 ( Anthophagus ; characters; France).

— Scheerpeltz, 1933: 1067 ( Hygrogeus ; catalog).

— Horion, 1935: 170 ( Hygrogeus ; characters; Alps; Austria).

— Horion, 1963: 137 ( Hygrogeus ; France; Switzerland; Italy; Germany; Austria).

— Lohse, 1964: 63 ( Hygrogeus ; characters; central Europe).

— Jarrige, 1968b: 76 ( Hygrogeus ; France).

— Zanetti, 1987: 384 ( Hygrogeus ; characters; Italy).

— Lucht, 1987: 90 ( Hygrogeus ; checklist; Germany; Austria).

— Zanetti, 1995: 9 ( Hygrogeus ; Italy).

- DISTRIBUTION: Germany, France, Switzerland, Italy, Austria.

cenisius Fairmaire, 1861 a: 647 ( Anthophagus ; Type locality: Mont Cenis).

— Fauvel, 1865a: 316 [= 1865b: 66] ( Anthophagus ; synonym of aemulus ).

— Fauvel, 1871a: 139 [= 1872: 113] ( Anthophagus ; synonym of aemulus ).

— Mulsant and Rey, 1880: 61 ( Geodromicus ; synonym of aemulus ).

— Ganglbauer, 1895: 710 ( Hygrogeus ; synonym of aemulus ).

— Luze, 1902d: 529 ( Hygrogeus ; synonym of aemulus ).

— Bernhauer and Schubert, 1910: 78 ( Geodromicus ; synonym of aemulus ).

alticola Coiffait, 1983: 283 ( Hygrogeus ; Type locality: Cachemire: Bradinath [Note: Should spelling be Badrinath], 3500 m).

- DISTRIBUTION: India.

armatus Cameron, 1941 , see: Trichodromeus .

atrocaeruleus Champion, 1925 , see: cyanipennis Cameron, 1924 .

brevisulcus Champion, 1925 , see: Trichodromeus .

cenisius Fairmaire, 1861 , see: aemulus Rosenhauer, 1847 .

coeruleipennis Coiffait, 1983: 280 ( Hygrogeus ; Type locality: Cachemire: Badrinath, 3500 m). - DISTRIBUTION: India.

cyanipennis Cameron, 1924b: 174 ( Hygrogeus ; Type locality: Chulli Khud , Charata District, 8000 feet).

— Cameron, 1930: 165 ( Hygrogeus ; characters; India).

— Scheerpeltz, 1933: 1067 ( Hygrogeus ; catalog).