Lesteva riparia, Heer, 1839: 193

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

riparia Heer, 1839: 193 ( Lesteva ; Type locality: Genf).

— Heer, 1841: 572 ( Lesteva ; variety of punctata ).

— Kraatz, 1857d: 934 ( Lesteva ; synonym of punctata ).

— Fauvel, 1869: 493 ( Lesteva ; synonym of punctata ).

— Fauvel, 1871a: 131 [= 1872: 105] ( Lesteva ; synonym of punctata ).

— Mulsant and Rey, 1880: 82 ( Lesteva ; synonym of punctata ).

— Ganglbauer, 1895: 714 ( Lesteva ; synonym of punctata ).

— Luze, 1903a: 184 ( Lesteva ; synonym of punctata ).

— Bernhauer and Schubert, 1910: 72 ( Lesteva ; synonym of punctata ).

muscorum Jacquelin du Val, 1859: 37 ( Lesteva ; Type locality: Pyrénées Orientales).

— Fauvel, 1871a: 131 [= 1872: 105] ( Lesteva ; synonym of punctata ).

— Mulsant and Rey, 1880: 82 ( Lesteva ; synonym of punctata ).

— Fowler, 1888: 402 ( Lesteva ; synonym of punctata ).

— Ganglbauer, 1895: 714 ( Lesteva ; synonym of punctata ).

— Luze, 1903a: 184 ( Lesteva ; synonym of punctata ).

— Bernhauer and Schubert, 1910: 72 ( Lesteva ; synonym of punctata ).

punctulata Latreille, 1804 , see: longoelytrata Goeze, 1777 .

reyi Jarrige, 1968 , see: pubescens Mannerheim, 1830 .

riparia Heer, 1839 , see: punctata Erichson, 1839 .

rufimarginata Rougemont, 2000: 168 ( Lesteva ; Type locality: China: Shaanxi, Qin Ling Shan, Mountain W pass at Autoroute at km 70, 46 km S. Xian, 108.47E, 33.51 N, 2300–2500 m).

- DISTRIBUTION: China.

rufitarsis Stephens, 1834 , see: longoelytrata Goeze, 1777 .

rufopunctata Rougemont, 2000: 162 ( Lesteva ; Type locality: China: Zhejiang, Moganshan ). - DISTRIBUTION: China.

sajanensis Zerche, 2000 a: 68 ( Lesteva ; Type locality: Zap. Sajan, Chrebet Kulumys und Ojskij pereval im Ermakovskij Rajon).

- DISTRIBUTION: Russia.

sbordonii Bordoni, 1973 h: 230 ( Lesteva ; subgenus Lestevina ; Type locality: Campania, Grava dei Gentili, S. Angelo a Fasanella, Salerno).

— Zanetti, 1987: 346 ( Lesteva ; subgenus Lestevina ; characters; Italy).

— Bordoni, 1987b: 94 ( Lesteva ; characters).

— Zanetti, 1995: 9 ( Lesteva ; Italy).

— Bordoni and Oromi, 1998: 1150 ( Lesteva ; subgenus Lestevina ; cave species; notes).

- DISTRIBUTION: Italy.

septemmaculata Rougemont, 2000: 167 ( Lesteva ; Type locality: China: Shaanxi, Qin Ling Shan, 118 km E. Xian, 110.06E, 34.27N, 1200–1400 m).

- DISTRIBUTION: China.

sharpi Rye, 1866 , see: monticola Kiesenwetter, 1847 .

shikokuana Watanabe, 1990: 160 ( Lesteva ; Type locality: Japan: Khikoku: Ehime Pref.: Omogokei ).

- DISTRIBUTION: Japan.

shibatai Watanabe, 1990: 158 ( Lesteva ; Type locality: Japan: Honshu: Tochigi Pref.: near Kôtoku-numa ).

— Watanabe and Baba 1973: 260 ( Lesteva ; [Note: Misidentified as plagiata according to Watanabe, 1996a: 9]; collected in gold mines; Japan).

— Watanabe, 1996a: 9 ( Lesteva ; mine collection; Japan).

- DISTRIBUTION: Japan.

sicula Erichson, 1840: 857 ( Lesteva ; Type locality: Sicilia).

— Jacquelin du Val, 1857: pl. 25, fig. 124 ( Lesteva ; [Note: Misidentification: Bernhauer and Schubert, 1910: 71 cited punctata Jacquelin du Val as a synonym of heeri ; Jacquelin du Val did not describe the species as new, he attributed it to Erichson]; illustration).

— Fauvel, 1886: 96 [= 1886a: 88] ( Lesteva ; notes; Algeria).

— Fowler, 1888: 402 ( Lesteva ; characters; Britain).

— Fauvel, 1902b: 56 ( Lesteva ; Algeria; Tunisia).

— Luze, 1903a: 185 ( Lesteva ; characters; Europe).

— Reitter, 1909: 184 ( Lesteva ; characters; Germany).

— Bernhauer and Schubert, 1910: 73 ( Lesteva ; catalog).

— Johansen, 1914: 590 ( Lesteva ; characters; Denmark).

— Scheerpeltz, 1933: 1064 ( Lesteva ; catalog).

— Lohse, 1960: 3 ( Lesteva ; Iberian Peninsula).

— Scheerpeltz, 1961d: 117 ( Lesteva ; Italy).

— Bordoni, 1973h: 234 ( Lesteva ; subgenus Lestevina ).

— Pope, 1977: 22 ( Lesteva ; synonym of heeri ).

— Zanetti, 1987: 342 ( Lesteva ; subgenus Lestevina ; characters; Italy).

— Lucht, 1987: 89 ( Lesteva ; checklist).

— L. Hansen and Sagvolden, 1995: 134 ( Lesteva ; collection notes; Norway).

— Sparacio, 1995: 142 ( Lesteva ; characters; notes; Sicily).

— Zanetti, 1995: 9 ( Lesteva ; Italy).

— Zanetti and Sabella, 1998: 33 ( Lesteva ; Sicily).

- DISTRIBUTION: Europe, Algeria, Tunisia.

heeri Fauvel, 1871 a: 132 [= 1872: 106] ( Lesteva ; Type locality: Lille ; Paris ; Caen ; Morlaix ; Tarbes ; Scandinavia; Britain; Germany).

— Seidlitz, 1875: 241 ( Lesteva ; characters; Baltic region).

— Mulsant and Rey, 1880: 84 ( Lesteva ; characters; France).

— Ganglbauer, 1895: 714 ( Lesteva ; synonym of sicula ).

— Ganglbauer, 1895: 714 ( Lesteva ; [Note: Misidentification: Bernhauer and Schubert, 1910: 71 cited sicula Ganglbauer as a synonym of heeri ; Ganglbauer did not describe the species as new, he attributed it to Erichson]; characters; Europe).

— Luze, 1903a: 185 ( Lesteva ; synonym of sicula ).

— Bernhauer and Schubert, 1910: 71 ( Lesteva ; catalog).

— Portevin, 1929: 438 ( Lesteva ; characters; France).

— Scheerpeltz, 1933: 1063 ( Lesteva ; catalog).

— Palm, 1948: 119 ( Lesteva ; characters; Sweden; Denmark).

— Lohse, 1956a: 54 ( Lesteva ; characters; Germany).

— Horion, 1963: 126 ( Lesteva ; subspecies of sicula ; Spain; France; Belgium; Holland; Britain; Denmark; Sweden; Germany).

— Lohse, 1964: 59 ( Lesteva ; subspecies of sicula ; characters; central Europe).

— Kasule, 1968: 136 ( Lesteva ; larval characters).

— Kasule, 1968a: 70 ( Lesteva ; life history).

— Steel, 1970a: 21 ( Lesteva ; cited as valid species; biological notes).

— Bordoni, 1973h: 234 ( Lesteva ; subgenus Lestevina ).

— Pope, 1977: 22 ( Lesteva ; Britain).

— Hammond, 1980: 136 ( Lesteva ; Ireland).

— Tóth, 1982: 98 ( Lesteva ; subspecies of sicula ; characters).

— Frank, 1982: 8 ( Lesteva ; list of parasites).

— Zanetti, 1987: 342 ( Lesteva ; subgenus Lestevina ; subspecies of sicula ; characters; Italy).

— Lucht, 1987: 89 ( Lesteva ; subspecies of sicula ; checklist; Germany; France; Benelux; Denmark; Sweden).

— Welch, 1993: 227 ( Lesteva ; ovariole number and ovary structure).

— Boháč, 1993: 41 ( Lesteva ; subgenus Lestevina ; subspecies of sicula ; Slovakia).

— M. Hansen, 1996: 95 ( Lesteva ; synonym of sicula ).

— Dauphin, Laguerre, Tamisier, and Teissier, 1997: 20 ( Lesteva ; subspecies of sicula ; France).

soror Smetana, 1967e: 300 ( Lesteva ; Type locality: Türkei: Yeniköy , Toros).

- DISTRIBUTION: Turkey.

staphylinoides Marsham, 1802 , see: longoelytrata Goeze, 1777 .

steeli Lohse, 1982: 203 ( Lesteva ; Type locality: India: Kashmir , Gulmarg).

- DISTRIBUTION: India.

subaptera Mulsant and Rey, 1880 , see: pubescens Mannerheim, 1830 .

submaculata Rougemont, 2000: 160 ( Lesteva ; Type locality: China: Zhejiang Prov., Anji County, ca 500 m, Long Wang Shan N. R.).

- DISTRIBUTION: China.

suturalis Lacordaire, 1835 , see: Geodromicus .

syriaca Luze, 1903 a: 192 ( Lesteva ; Type locality: Syrien: Beirut , Appl).

— Bernhauer and Schubert, 1910: 73 ( Lesteva ; catalog).

- DISTRIBUTION: Lebanon.

szekessyi Lohse and Steel, 1961: 74 ( Lesteva ; Type locality: Crete: Antr. Jovis, Mt. Ida , 2200 m).

- DISTRIBUTION: Crete.

taygetana Lohse, 1955 a: 512 ( Lesteva ; subgenus Lestevidia ; Type locality: Taygetos, Peloponnes).

- DISTRIBUTION: Greece.

torrentum Cameron, 1924b: 171 ( Lesteva ; Type locality: Mussorie District, 6500 to 7000 feet).

— Cameron, 1930: 159 ( Lesteva ; characters; India).

— Scheerpeltz, 1933: 1064 ( Lesteva ; catalog).

— Lohse, 1982: 201, 204 ( Lesteva ; characters; India).

- DISTRIBUTION: India.

transcaspica Bernhauer, 1935 a: 39 ( Lesteva ; Type locality: Transkaspien: Ljutfabad).

— Tikhomirova, 1973a: 140 ( Lesteva ; checklist; Central Asia).

- DISTRIBUTION: Turkmenistan.

truncata Casey, 1885 , see: Unamis .

truncata Lohse, 1960 , see: fontinalis Kiesenwetter, 1850 .

tsushimae Watanabe, 1990: 170 ( Lesteva ; Type locality: Japan: Nagasaki Pref.: Tsushima Is., Izuhara).

- DISTRIBUTION: Japan.

turcica Smetana, 1967e: 305 ( Lesteva ; Type locality: Türkei: Yeniköy , Toros).

— Tikhomirova, 1973a: 140 ( Lesteva ; cited as turcica Smetata, 1937 ; checklist; Central Asia). - DISTRIBUTION: Turkey.

turkestanica Luze, 1904: 76 ( Lesteva ; Type locality: Seravschan: Boschara).

— Bernhauer and Schubert, 1910: 73 ( Lesteva ; catalog).

— Tikhomirova, 1973a: 140 ( Lesteva ; checklist; Central Asia).

— Bordoni, 1987b: 92 ( Lesteva ; notes).

- DISTRIBUTION: Uzbekistan.

uhligi Zanetti, 1984: 79 ( Lesteva ; Type locality: Bulgaria: Stara Planina , 8 km S. of Botev, 1200 m).

- DISTRIBUTION: Bulgaria.

villardi Mulsant and Rey, 1880: 413 ( Lesteva ; Type locality: la grotte du Brudou, à Lenteen-Royans, Drôme).

— Ganglbauer, 1895: 713 ( Lesteva ; characters; France).

— Luze, 1903a: 194 ( Lesteva ; characters; France).

— Bernhauer and Schubert, 1910: 73 ( Lesteva ; catalog).

— Jeannel and Jarrige, 1949: 319 ( Lesteva ; characters; notes; cave species; France).

— Jarrige, 1968b: 75 ( Lesteva ; France).

— Frank, 1982: 8 ( Lesteva ; list of parasites).

— Zanetti, 1987: 361 ( Lesteva ; subgenus Lesteva ; characters; Italy).

— Zanetti, 1995: 9 ( Lesteva ; Italy).

— Bordoni and Oromi, 1998: 1157 ( Lesteva ; facultative cave species; caves in France and Italy). - DISTRIBUTION: France, Italy.

villosa Waltl, 1838 , see: punctata Erichson, 1839 .

yunnanicola Rougemont, 2000: 166 ( Lesteva ; Type locality: China: Yünnan, ca. 100 km NNE Kunming, Liangwang Shan , ca. 2300 m).

- DISTRIBUTION: China.

zolotarevi Roubal, 1910: 99 , 100 ( Lesteva ; Type locality: Caucasus occ.: Kluchor, in der Höhe von etwa 2200 m).

— Scheerpeltz, 1933: 1064 ( Lesteva ; catalog).

— Tikhomirova, 1973a: 140 ( Lesteva ; checklist; western Caucasus).

- DISTRIBUTION: Russia.

Liophilydrodes

[8 species; Palaearctic Region]

Liophilydrodes Nakane, 1983: 148 (replacement name for Lioplax Nakane and Sawada ; [Note: Nakane attributed the name to both himself and Sawada, but in the paragraph preceding the name wrote, “I propose a new name for our Lioplax herein”]). TYPE SPECIES: Philydrodes pullus Nakane and Sawada , fixed by objective synonymy with Lioplax for which the type species had already been fixed.

— Watanabe, 1990: 295 (characters; key to species of Japan).

— Watanabe, 1995: 329 (characters).

Lioplax Nakane and Sawada, 1956 a: 183 (subgenus of Philydrodes ; species included: pulla , puncticollis , troglophila , subtilis ; preoccupied by Troschel, 1857). TYPE SPECIES: Philydrodes pullus Nakane and Sawada , fixed by original designation.

— Nakane, 1983: 148 (synonym of Liophilydrodes ).

— Watanabe, 1990: 295 (synonym of Liophilydrodes ).

flavipennis Watanabe, 1990: 303 ( Liophilydrodes ; Type locality: Japan: Honshu: Yamanashi Pref.: Mt. Senjô-ga-take, S. Jpn. Alps).

- DISTRIBUTION: Japan.

pullus Nakane and Sawada, 1956 a: 184 ( Philydrodes ; subgenus Lioplax ; Type locality: Japan: Mt. Ontake, Nagano).

— Watanabe, 1990: 298 ( Liophilydrodes ; characters; Japan).

- DISTRIBUTION: Japan.

puncticollis Nakane and Sawada, 1956 a: 183 ( Philydrodes ; subgenus Lioplax ; Type locality: Japan: Hokkaido: Nukabira).

— Watanabe and Shibata, 1965: 317 ( Philydrodes , cited as Phylidrodes; subgenus Lioplax ; Japan).

— Watanabe, 1990: 306 ( Liophilydrodes ; characters; Japan).

- DISTRIBUTION: Japan.

subtilis Sharp, 1889: 471 ( Anthophagus ; Type locality: summit of Iwakisan).

— Bernhauer and Schubert, 1910: 81 ( Anthophagus ; catalog).

— Scheerpeltz, 1933: 1067 ( Philydrodes ; catalog).

— Nakane and Sawada, 1956a: 184 ( Philydrodes ; subgenus Lioplax ; characters from original description; Japan).

— Watanabe, 1990: 302 ( Liophilydrodes ; characters; Japan).

- DISTRIBUTION: Japan.

suzukii Watanabe, 1990: 305 ( Liophilydrodes ; Type locality: Japan: Honshu : Gifu Pref.:

Kagami-daira on Mt. Yumiori-dake, N. Jpn. Alps).

troglophilus Nakane and Sawada, 1956 a: 184 ( Philydrodes ; subgenus Lioplax ; Type locality: Japan: Tokyo, Okutama, Nippara cave).

— Watanabe, 1990: 306 ( Liophilydrodes ; characters; Japan).

— Watanabe, 1996a: 9 ( Liophilydrodes ; cave collection; Japan).

- DISTRIBUTION: Japan.

uenoi Watanabe, 1995: 329 ( Liophilydrodes ; Type locality: Jiuzhai Gou, Chang Hai , 3100 m, Nanping Xian, Sichuan Sheng, Southwest China).

- DISTRIBUTION: China.

yamanakai Watanabe, 1990: 301 ( Liophilydrodes ; Type locality: Japan: Honshu: Toyama Pref.: Mt. Tateyama ).

- DISTRIBUTION: Japan.

Mannerheimia

[19 species; Palaearctic and Nearctic Regions]

Mannerheimia Mäklin, 1880: 80 (species included: divergens , affine , confusum , saginatum ). TYPE SPECIES: Omalium divergens Mäklin , fixed by subsequent designation by R. Lucas, 1920: 392.

— Heyden, 1880: 82 (list of species of Siberian region).

— Mäklin, 1881: 39 (characters; [Note: Published without citation of earlier reference as though a new genus]).

— Duvivier, 1883: 196 (catalog).

— Heyden, Reitter, and Weise, 1891: 119 (subgenus of Deliphrum ; list of species of Europe and Caucasus).

— Ganglbauer, 1895: 725 (characters).

— Luze, 1905c: 251 (characters; key to species).

— Heyden, Reitter, and Weise, 1906: 133 (list of species of Europe and Caucasus).

— Bernhauer and Schubert, 1910: 63 (world catalog; 3 species).

— Gridelli, 1924c (notes; key to italian species).

— Winkler, 1925: 332 (catalog for Palaearctic region).

— Porta, 1926: 24 (characters; key to species of Italy).

— Cameron, 1930: 149 (characters).

— Scheerpeltz, 1933: 1054 (world catalog supplement).

— Palm, 1948: 94 (characters; key to species of Sweden).

— Blackwelder, 1952: 230 (type species).

— Horion, 1963: 100 (list of central European species).

— Székessy, 1963: 18 (characters).

— Lohse, 1964: 50 (characters; key to central European species).

— Scheerpeltz, 1968a: 12 (catalog of Austrian species).

— Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Muona, 1979: 18 (list of Scandinavian species).

— Coiffait, 1982: 150 (key to species of Kashmir and Nepal).

— Zanetti, 1983: 429 (characters).

— Zanetti, 1987: 273 (characters; habitus; key to species of Italy).

— Muona and Viramo, 1986: 15 (list of species of northeastern Finland).

— Watanabe, 1990: 77 (characters; key to species of Japan).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Siitonen, 1993: 230 (list of Finnish species).

— Zanetti, 1995: 8 (list of species of Italy).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (1 undescribed Nearctic species; notes; characters in key).

affinis Mäklin, 1878 , see: arctica Erichson, 1840 .

afghanica Coiffait, 1982b: 82 ( Mannerheimia ; Type locality: Afghanistan: Nouristan: Kamdesch , 2000 m).

aprutiana Gridelli, 1924c: 135 , 136 ( Mannerheimia ; Type locality: Maiella , Abruzzo).

— Scheerpeltz, 1933: 1054 ( Mannerheimia ; catalog).

— Zanetti, 1987: 276 ( Mannerheimia ; characters; Italy).

— Zanetti, 1995: 8 ( Mannerheimia ; Italy).

- DISTRIBUTION: Italy.

arctica Erichson, 1840: 873 ( Deliphrum ; Type locality: Lapland).

— Thomson, 1861: 198 ( Deliphrum ; characters; Scandinavia).

— Seidlitz, 1875: 236 ( Deliphrum ; characters; Baltic region).

— J. Sahlberg, 1876: 214 ( Deliphrum ; Finland).

— J. Sahlberg, 1880: 108 ( Mannerheimia ; characters; Siberia).

— Ganglbauer, 1895: 725 ( Mannerheimia ; characters; Palaearctic region).

— Luze, 1905c: 253 ( Mannerheimia ; characters; Sweden; Norway; Finland; Lapland; Arctic Siberia; southern Alps).

— Bernhauer and Schubert, 1910: 63 ( Mannerheimia ; catalog).

— Gridelli, 1924c: 135, 136 ( Mannerheimia ; key characters; notes; Italy).

— Scheerpeltz, 1933: 1054 ( Mannerheimia ; catalog).

— Horion, 1935: 173 ( Mannerheimia ; characters; northern Europe; Siberia; southern Alps; Germany).

— Palm, 1948: 96 ( Mannerheimia ; characters; Sweden; Norway; Finland).

— Horion, 1963: 100 ( Mannerheimia ; Norway; Sweden; Finland; Russia; Austria).

— Lohse, 1964: 50 ( Mannerheimia ; characters; central Europe).

— Tikhomirova, 1973a: 139 ( Mannerheimia ; checklist; Alps; northern Europe; Siberia).

— Zanetti, 1987: 274 ( Mannerheimia ; characters; Italy).

— Lucht, 1987: 88 ( Mannerheimia ; checklist; Austria).

— Zanetti, 1995: 8 ( Mannerheimia ; Italy).

— Kashcheev, 1999: 143 ( Mannerheimia ; characters).

- DISTRIBUTION: Sweden, Norway, Finland, Russia, Austria, Italy.

affinis Mäklin, 1878: 27 [= 1878a: 303] ( Omalium , cited as Homalium ; [preoccupied]; Type locality: Werschininskoj, 68°45′ n. lat.).

— J. Sahlberg, 1880: 108 ( Mannerheimia ; synonym of arctica ).

— Mäklin, 1881: 40 ( Mannerheimia ; [Note: Cited by Bernhauer and Schubert, 1910: 63 as the first reference for the species]; characters; Siberia).

— Ganglbauer, 1895: 725 ( Mannerheimia ; synonym of arctica ).

— Luze, 1905c: 253 ( Mannerheimia ; synonym of arctica ).

— Bernhauer and Schubert, 1910: 63 ( Mannerheimia ; synonym of arctica ).

confusa Mäklin, 1878: 27 [= 1878a: 303] ( Omalium , cited as Homalium ; Type locality: Chantaiskoj, 68°5′ n. lat.).

— J. Sahlberg, 1880: 108 ( Mannerheimia ; synonym of arctica ).

— Mäklin, 1881: 40 ( Mannerheimia ; [Note: Cited by Bernhauer and Schubert, 1910: 63 as the first reference for the species]; characters; Siberia).

— Ganglbauer, 1895: 725 ( Mannerheimia ; synonym of arctica ).

— Luze, 1905c: 253 ( Mannerheimia ; synonym of arctica ).

— Bernhauer and Schubert, 1910: 63 ( Mannerheimia ; synonym of arctica ).

saginata Mäklin, 1878: 28 [= 1878a: 304] Omalium , cited as Homalium ; Type locality: Krasnojarsk och Tomsk).

— J. Sahlberg, 1880: 108 ( Mannerheimia ; synonym of arctica ).

— Mäklin, 1881: 41 ( Mannerheimia ; [Note: Cited by Bernhauer and Schubert, 1910: 63 as the first reference for the species]; characters; Siberia).

— Ganglbauer, 1895: 725 ( Mannerheimia ; synonym of arctica ).

— Luze, 1905c: 253 ( Mannerheimia ; synonym of arctica ).

— Bernhauer and Schubert, 1910: 63 ( Mannerheimia ; synonym of arctica ).

asiatica Kashcheev, 1999: 142 ( Mannerheimia ; Type locality: Tian’shan’, Khr. Kungei-Alatau, ushch., Kul’bastau, 2500-3000 m).

brevipennis Motschulsky, 1860: 545 ( Omalium ; [preoccupied]; Type locality: Mt. Ula Tau, Kazakh Uplands ; [Note: See lectotype designation by Hammond, 1970]).

— Bernhauer and Schubert, 1910: 51 ( Omalium ; subgenus Omalium ; catalog).

— Hammond, 1970: 67, 70 ( Mannerheimia , transferred from Omalium ; lectotype designation:

Mt. Ula Tau, Kazakh Uplands ; characters; northern Scandinavia, northern Russia, northern Siberia; Alps; Tyrol; Apennines, Altai Range, Himalaya).

— Herman, 2001: 37 ( Mannerheimia ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Russia, Sweden, Austria, Italy, Himalaya, Altai region.

divergens Mäklin, 1878: 26 [= 1878a: 302] ( Omalium , cited as Homalium ; Type locality: Seliwaninskoj, 65°55′ n. lat; Baklanowskij, 64° 25 n. lat.).

— J. Sahlberg, 1880: 108 ( Mannerheimia ; characters; Siberia).

— Mäklin, 1881: 39 ( Mannerheimia ; characters; Siberia).

— Luze, 1905c: 254 ( Mannerheimia ; characters; Siberia; Sweden; Altai region).

— Bernhauer and Schubert, 1910: 52 ( Omalium ; subgenus Omalium ; catalog).

— Bernhauer and Schubert, 1910: 63 ( Mannerheimia ; [Note: The species was cited twice by Bernhauer and Schubert, once in Omalium and once in Mannerheimia but for the latter the original reference is cited incorrectly as Mäklin, 1880]; catalog).

— Gridelli, 1924c: 135, 137 ( Mannerheimia ; characters; Altai region; Siberia; Sweden).

— Scheerpeltz, 1933: 1054 ( Mannerheimia ; catalog).

— Palm, 1948: 96 ( Mannerheimia ; characters; Sweden).

— Hammond, 1970: 67, 70 ( Mannerheimia ; synonym of brevipenne ).

— Kashcheev, 1999: 143 ( Mannerheimia ; cited as valid species; characters).

cachemirica Coiffait, 1982: 146 ( Mannerheimia ; Type locality: Inde: Cachemire , Aru).

- DISTRIBUTION: India.

confusa Mäklin, 1878 , see: arctica Erichson, 1840 .

curtella Sharp, 1889: 473 ( Omalium ; Type locality: Miyanoshita).

— Bernhauer and Schubert, 1910: 52 ( Omalium ; subgenus Omalium ; catalog).

— Cameron, 1949b: 460 ( Omalium ; Japan).

— Watanabe, 1990: 81 ( Mannerheimia ; characters; Japan).

- DISTRIBUTION: Japan.

divergens Mäklin, 1878 , see: brevipennis Motschulsky, 1860 .

doderoi Gridelli, 1924: 135 , 137 ( Mannerheimia ; Type locality: Valsavaranche, Valle d’Aosta, Alpi Graie ).

— Scheerpeltz, 1933: 1054 ( Mannerheimia ; catalog).

— Horion, 1963: 101 ( Mannerheimia ; Italy).

— Lohse, 1964: 50 ( Mannerheimia ; characters; central Europe).

— Hammond, 1970: 68 ( Mannerheimia ; may be synonym of divergens ).

— Zanetti, 1987: 277 ( Mannerheimia ; characters; Italy).

— Lucht, 1987: 88 ( Mannerheimia ; checklist; Austria).

— Zanetti, 1995: 8 ( Mannerheimia ; Italy).

— Kashcheev, 1999: 143 ( Mannerheimia ; characters).

— Assing, Frisch, Kahlen, et al., 1998: 124 ( Mannerheimia ; Austria).

- DISTRIBUTION: Austria, Italy.

emodensa Coiffait, 1977: 266 ( Mannerheimia ; Type locality: Environs de Alm Darghari près de Maharigaon, 4000 m région de Jumla, Népal occidentale).

— Coiffait, 1982: 42 ( Mannerheimia ; Nepal).

— Coiffait, 1982: 150 ( Mannerheimia ; characters).

- DISTRIBUTION: Nepal.

goriensis Champion, 1925: 103 ( Mannerheimia ; Type locality: Gori Valley, E. Kumaon,

alt. 7000 ft.).

— Cameron, 1930: 150 ( Mannerheimia ; characters; India).

— Coiffait, 1982: 150 ( Mannerheimia ; characters).

— Coiffait, 1984: 118 ( Mannerheimia ; Pakistan).

- DISTRIBUTION: India, Pakistan.

japonica Watanabe, 1990: 87 ( Mannerheimia ; Type locality: Japan: Honshu : Miyagi Pref.:

Mt. Ushiroeboshi-dake on Zaô Mts.).

- DISTRIBUTION: Japan.

evias Watanabe, 1990: 84 ( Mannerheimia ; Type locality: Japan: Honshu: Gifu Pref.: Mt. Sugoroku-dake on northern Japanese Alps ).

- DISTRIBUTION: Japan.

kirschenblatti Khnzorian, 1956: 138 ( Mannerheimia ; Type locality: ArmSSR: Verkhnii Gukasian [Gukasianskii r-n], na zapalnykh kechutskikh ros, 2500 m).

— Tikhomirova, 1973a: 139 ( Mannerheimia ; checklist; Armenia).

— Kashcheev, 1999: 143 ( Mannerheimia ; characters).

- DISTRIBUTION: Armenia.

obscura Coiffait, 1982: 148 ( Mannerheimia ; Type locality: Inde: Cachemire , Chandanwari, près Pahalgam, 2800–3100 m).

— Coiffait, 1983: 274 ( Mannerheimia ; Kashmir).

- DISTRIBUTION: India.

micros Fauvel, 1900d: 222 ( Deliphrum ; subgenus Mannerheimia ; Type locality: Turkestan: Monts Alai).

— Luze, 1905c: 255 ( Mannerheimia ; characters; Turkestan).

— Bernhauer and Schubert, 1910: 63 ( Mannerheimia ; catalog).

— Gridelli, 1924c: 138 ( Mannerheimia ; characters; Turkestan).

— Scheerpeltz, 1933: 1054 ( Mannerheimia ; catalog).

— Hammond, 1970: 68 ( Mannerheimia ; may be synonym of divergens ).

— Tikhomirova, 1973a: 139 ( Mannerheimia ; checklist; Central Asia).

— Kashcheev, 1999: 143 ( Mannerheimia ; characters).

- DISTRIBUTION: Kyrgyzstan.

saginata Mäklin, 1878, see: arctica Erichson, 1840 .

scutellata Coiffait, 1982: 149 ( Mannerheimia ; Type locality: Inde: Cachemire , Sonamarg , Himalaya-Hauptkette, Nichinai Tal, 3100–3200 m).

- DISTRIBUTION: India.

tsurumii Watanabe, 1990: 83 ( Mannerheimia ; Type locality: Japan: Honshu: Tochigi Pref.: near Yunoko Lake, Oku-Nikkô ).

- DISTRIBUTION: Japan.

vicina Coiffait, 1982: 146 ( Mannerheimia ; Type locality: Inde: Cachemire , Chandanwari,

près Pahalgam, 2800–3100 m).

- DISTRIBUTION: India.

yasudai Watanabe, 1990: 86 ( Mannerheimia ; Type locality: Japan: Hokkaido: Mt. Kurodake on Daisetsu Mts. ).

- DISTRIBUTION: Japan.

Microedus

[7 species; Nearctic Region]

Microedus LeConte, 1874: 273 (species included: austinianus ). TYPE SPECIES: Microedus austinianus LeConte , fixed by monotypy.

— Fauvel, 1878d: 250 [= 1878e: 86] (characters).

— Duvivier, 1883: 194 (catalog).

— Bernhauer and Schubert, 1910: 75 (world catalog; 2 species).

— Leng, 1920: 95 (catalog of North American species).

— Blackwelder, 1952: 245 (type species: austinianus ).

— Moore, 1966: 48, 49 (characters; notes).

— Moore and Legner, 1972a: 75 (key to species).

— Moore and Legner, 1975: 197 (catalog; North America north of Mexico).

— Campbell, 1978: 1 (moved to Anthophagini ).

— Moore and Legner, 1979: 184 (characters; notes).

— Campbell and Davies, 1991: 91 (checklist of species; Alaska and Canada).

— Downie and Arnett, 1996: 435 (characters; species of Northeastern North America ).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (7 Nearctic species; notes; characters in key).

austinianus LeConte, 1874: 273 ( Microedus ; Type locality: White Mountains [New Hampshire], Vancouver Island ).

— Fauvel, 1878d: 250 [= 1878e: 86] ( Microedus ; characters; New Hampshire; Vancouver Island).

— Bernhauer and Schubert, 1910: 75 ( Microedus ; catalog).

— Van Dyke, 1924: 16 ( Microedus ; cited as austianus; Alaska).

— Fall, 1926: 145 ( Microedus ; Alaska).

— Hatch, 1957: 71 ( Microedus ; characters; British Columbia ; Idaho; Washington; Oregon).

— Moore and Legner, 1972a: 75 ( Microedus ; characters).

— Downie and Arnett, 1996: 435 ( Microedus ; characters; Canada; USA).

- DISTRIBUTION: Canada, USA.

ewingi Hatch, 1957: 72 ( Microedus ; Type locality: Washington: Mt. Rainier, below Sluskin Falls ).

— Moore and Legner, 1972a: 75 ( Microedus ; characters).

- DISTRIBUTION: USA.

fenderi Hatch, 1957: 72 ( Microedus ; Type locality: Washington: Olympic Hot Springs. Also cited from British Columbia ).

— Moore and Legner, 1972a: 75 ( Microedus ; characters).

- DISTRIBUTION: Canada, USA.

giulianii Moore, 1972 a: 75 ( Microedus ; Type locality: South Fork of Big Pine Creek , Inyo County, California, 8500 feet).

- DISTRIBUTION: USA.

laticollis Mannerheim, 1843b: 234 ( Anthophagus ; Type locality: insula Sitkha).

— Mannerheim, 1852: 319 ( Anthophagus ; characters; Sitka).

— Fauvel, 1878d: 251 [= 1878e: 87] ( Microedus ; characters; Alaska).

— Bernhauer and Schubert, 1910: 75 ( Microedus ; catalog).

— Fall, 1926: 145 ( Microedus ; Alaska).

— Hatch, 1957: 71 ( Microedus ; characters; British Columbia ; Washington; Oregon; Idaho).

— Moore and Legner, 1972a: 75 ( Microedus ; characters).

- DISTRIBUTION: Canada, USA.

porterae Hatch, 1957: 71 ( Microedus ; Type locality: Washington: Olympic Hot Springs. Also cited from Oregon, Idaho, and British Columbia ).

— Moore and Legner, 1972a: 75 ( Microedus ; cited as porteri ; characters).

- DISTRIBUTION: Canada, USA.

rogersi Hatch, 1957: 71 ( Microedus ; Type locality: Washington: Olympic Hot Springs. Also cited from British Columbia ).

— Moore and Legner, 1972a: 75 ( Microedus ; characters).

- DISTRIBUTION: Canada, USA.

Olophrum

[54 species (1 doubtful); Nearctic and Palaearctic Regions]

Olophrum Erichson, 1839 a: 622 (species included: piceum , fuscum , assimile ). TYPE SPECIES: Omalium piceum Gyllenhal , fixed by subsequent designation by Westwood, 1840: 156.

— Heer, 1839: 189 (characters).

— Stephens, 1839: 422 (characters).

— Redtenbacher, 1849: 749 (characters).

— Schaum, 1852: 30 (list of species; Europe).

— Lacordaire, 1854: 140 (characters; notes; list of species).

— Fairmaire and Laboulbène, 1856: 634 (characters).

— Gistel, 1856: 389 (list of species).

— Jacquelin du Val, 1857: 74 (characters).

— Redtenbacher, 1857: 247 (characters).

— Kraatz, 1857d: 939 (characters).

— Thomson, 1858: 37 (characters).

— G. Waterhouse, 1858: 31 (catalog of British species).

— Thomson, 1859: 49 (characters; type species: piceum ).

— Schaum, 1859: 31 (catalog; European species).

— LeConte, 1861a: 70 (characters).

— Thomson, 1861: 192 (characters).

— Gredler, 1863: 123 (list of species from Tirol).

— LeConte, 1863: 26 (list of North American species).

— Wencker and Silbermann, 1866: 36 (list of species; collecting notes; France).

— Gemminger and Harold, 1868: 660 (catalog).

— Jacquelin du Val, 1868: 80 (catalog; European species).

— Fauvel, 1871a: 123 [= 1872: 97] (characters; key to species).

— Sharp, 1871d: 14 (list of British species).

— Redtenbacher, 1874: 267 (characters; key to Austrian species).

— Fauvel, 1875a:VI [= 1875b: 208] (catalog).

— Fauvel, 1878d: 242 [= 1878e: 78] (key to North American species).

— Mulsant and Rey, 1880: 132 (characters; key to French species).

— Heyden, 1880: 81 (list of species of Siberian region).

— Duvivier, 1883: 195 (catalog).

— Fowler, 1888: 404 (notes; key to British species).

— Seidlitz, 1891: 345 (key to some European species).

— Seidlitz, 1891a: 367 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 119 (list of species of Europe and Caucasus).

— Heyden, 1893: 48 (list of species of Siberian region).

— Ganglbauer, 1895: 719 (characters; key to middle European species).

— Heyden, 1896: 37 (list of species of Siberian region).

— Luze, 1905: 33 (characters; revision of and key to Palaearctic species).

— Heyden, Reitter, and Weise, 1906: 133 (list of species of Europe and Caucasus).

— Sainte-Claire Deville, 1907: 44 (characters; key to and annotated list of species of Seine Basin, France).

— Reitter, 1909: 185 (characters; key to German species).

— Bernhauer and Schubert, 1910: 65 (world catalog; 18 species).

— Blatchley, 1910: 478 (characters).

— Petri, 1912: 54 (list of species of Siebenbürgen).

— Johansen, 1914: 592 (characters; key to Danish species).

— Leng, 1920: 94 (catalog of North American species).

— Winkler, 1925: 333 (catalog for Palaearctic region).

— Porta, 1926: 26 (characters; key to species of Italy).

— Scheerpeltz, 1929 (monograph; discussion; taxonomic history; key to species groups and to species; characters; illustrations; zoogeography; catalog).

— Portevin, 1929: 440 (characters; key to French genera).

— Roubal, 1930: 313 (catalog; Slovakia).

— Joy, 1932: 95 (key to British species).

— Scheerpeltz, 1933: 1056 (world catalog supplement).

— Munster, 1935 (discussion of Scheerpeltz’s revision; key to some species).

— Palm, 1948: 103 (characters; key to species of Sweden).

— Tottenham, 1949: 357, 407 (type species: piceum Gyllenhal ; list of British species).

— Tottenham, 1954: 30 (characters; key to British species).

— Hatch, 1957: 50, 61 (characters; key to species of Pacific Northwest).

— Horion, 1963: 109 (list of central European species).

— Székessy, 1963: 20 (characters).

— Lohse, 1964: 53 (characters; key to central European species).

— Smetana, 1964c: 170 (key to Czechoslovakian species).

— Moore, 1966: 48, 55 (characters; notes).

— Pototskaia, 1967: 20 (larval characters).

— Kasule, 1968: 133, 134 (larval characters).

— Scheerpeltz, 1968a: 14 (catalog of Austrian species).

— Steel, 1970a: 18 (larval characters; biological notes).

— Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Moore and Legner, 1975: 199 (catalog; North America north of Mexico).

— Shibata, 1976: 108 (checklist of species of Japan).

— Pope, 1977: 22 (list of British species).

— Coiffait, 1978f: 277 (key to species of France).

— Coiffait, 1978f: 277 ( Olophrum ; characters; France).

— Topp, 1978: 308 (larval characters).

— Burakowski, Mroczkowski, and Stefańska, 1979: 57 (catalog; Poland).

— Moore and Legner, 1979: 186 (characters; notes).

— Muona, 1979: 18 (list of Scandinavian species).

— Uhlig, 1979: 242 (some species collected in Germany).

— M. Dvořák, 1979: 109 (some species collected in Slovakia).

— Uhlig, Vogel, and Sieber, 1980: 241 (some species collected in Germany).

— Uhlig and Vogler, 1981: 87 (list of some species collected in Germany; habitat and collecting notes).

— Waterston, Owen, Welch, Bacchus and Hammond, 1981: 273 (list of species of Outer Hebrides).

— Spahr, 1981: 95 (references to specimens from amber and copal).

— Tóth, 1982: 83 (characters; habitus; key to species of Hungary).

— Campbell, 1983 (characters; habitus; revision of and key to North American species; discussion).

— Ádám, 1985: 251 (list of species from southeast Hungary).

— Segers, 1986: 21 (checklist of species; Belgium).

— Zanetti, 1987: 298 (characters; habitus; key to species of Italy).

— Ádám, 1987: 137 (list of two species with collecting notes; Hungary).

— Muona and Viramo, 1986: 15 (list of species of northeastern Finland).

— Lohse and Lucht, 1989: 126 (notes).

— Watanabe, 1990: 126 (characters; key to species of Japan).

— Ryabukhin, 1991b: 13 (charactes; key to species of northeastern Russia).

— Campbell and Davies, 1991: 89 (checklist of species; Alaska and Canada).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Siitonen, 1993: 230 (list of Finnish species).

— Welch, 1993: 227 (ovariole number and ovary structure).

— Boháč, 1993: 41 (list of species of Czech Republic and Slovakia).

— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 27 (additional records to Danish list of species).

— Zanetti, 1995: 8 (list of species of Italy).

— M. Hansen, 1996: 95 (list of Danish species).

— Downie and Arnett, 1996: 432 (characters; key to species of Northeastern North America ).

— M. Hansen, Palm, Pedersen, and Runge, 1998: 70 (list of species collected in 1997; Denmark).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (7 Nearctic species; notes; characters in key).

Lathrium LeConte, 1850: 221 (species included: convexicolle ). TYPE SPECIES: Lathrium convexicolle LeConte , fixed by monotypy.

— Lacordaire, 1854: 142 (characters; notes; list of species).

— Gemminger and Harold, 1868: 660 (synonym of Olophrum ).

— Fauvel, 1871a: 123 [= 1872: 97] (synonym of Olophrum ).

— Bernhauer and Schubert, 1910: 66 (synonym of Olophrum ).

— Blackwelder, 1952: 209 (type species: convexicolle ).

— Campbell, 1983: 579 (synonym of Olophrum ).

alpestre Erichson, 1840 , see: consimile Gyllenhal, 1810 .

alpinum Heer, 1839 , see: consimile Gyllenhal, 1810 .

aragatzense Khnzorian, 1962: 103 ( Olophrum ; Type locality: ArmSSR: Aragats u ozera Karilich, 3200 m).

- DISTRIBUTION: Armenia.

arcanum † Scudder, 1900a: 86 ( Olophrum ; pleistocene fossil; Type locality: Scarborough).

- DISTRIBUTION: Canada.

arrowi Scheerpeltz, 1929: 105 ( Olophrum ; Type locality: Japan).

— Nakane, 1963a: 82 ( Olophrum , cited as Omalium ; characters; habitus photograph; Japan).

— Hasegawa and Kanie, 1992: 19 ( Olophrum ; seashore collection; Japan).

— Watanabe, 1990: 130 ( Olophrum ; characters; Japan).

- DISTRIBUTION: Japan.

arvernicum Coiffait, 1978f: 275 ( Olophrum ; Type locality: Puy de Sancy, Puy-de-Dôme).

- DISTRIBUTION: France.

assimile Paykull, 1800: 409 ( Staphylinus ; Type locality: Suecia).

— Gyllenhal, 1810: 202 ( Omalium ; characters).

— Gyllenhal, 1827: 463 ( Omalium ).

— C. Sahlberg, 1827: 278 ( Omalium ; characters; Finland).

— Curtis, 1829: 28 ( Anthobium ; catalog; Britain).

— Mannerheim, 1830: 51 ( Omalium ; Sweden).

— Mannerheim, 1831: 465 ( Omalium ; Sweden).

— Erichson, 1839a: 624 ( Olophrum ; characters; Germany).

— Heer, 1839: 189 ( Olophrum ; characters; Switzerland).

— Erichson, 1840: 865 ( Olophrum ; characters; Germany; Sweden).

— Laporte, 1840: 191 ( Omalium ; characters; Sweden; France).

— Redtenbacher, 1849: 749 ( Olophrum ; characters; Austria).

— Fairmaire and Laboulbène, 1856: 635 ( Olophrum ; characters; France).

— Redtenbacher, 1857: 248 ( Olophrum ; characters; Austria).

— Kraatz, 1857d: 942 ( Olophrum ; characters; Germany).

— Thomson, 1861: 194 ( Olophrum ; characters; Scandinavia).

— Hochhuth, 1862: 103 ( Olophrum ; Russia).

— Fauvel, 1871a: 124 [= 1872: 98] ( Olophrum ; characters; France; Scandinavia; Germany; Switzerland; Austria; Russia; Syria).

— Redtenbacher, 1874: 268 ( Olophrum ; characters; Austria).

— Seidlitz, 1875: 240 ( Olophrum ; characters; Baltic region).

— J. Sahlberg, 1876: 213 ( Olophrum ; Finland).

— Mulsant and Rey, 1880: 139 ( Olophrum ; characters; France).

— Ganglbauer, 1895: 720 ( Olophrum ; characters; north and middle Europe).

— Luze, 1905: 41 ( Olophrum ; characters; central and northern Europe; Caucasus).

— Reitter, 1909: 186 ( Olophrum ; characters; Germany).

— Bernhauer and Schubert, 1910: 66 ( Olophrum ; catalog).

— Johansen, 1914: 593 ( Olophrum ; characters; Denmark).

— Scheerpeltz, 1929: 113 ( Olophrum ; characters; Bosnia, Serbia, Dalmatia, Romania, Ukraine, Alps, Austria, Poland, Germany, Holland, Belgium, France, Denmark, Sweden, Norway, Finland, Russia, Caucasus).

— Portevin, 1929: 440 ( Olophrum ; characters; France).

— Munster, 1936: 101 ( Olophrum ; characters; notes).

— Horion, 1963: 113 ( Olophrum ; Caucasus; Syria; Russia; Finland; Sweden; Norway; Scotland; Denmark; Baltic states; Poland; Czechslovakia; Hungary; Romania; Yugoslavia; Slovenia; Italy?; France; Switzerland; Belgium; Germany; Austria).

— Lohse, 1964: 54 ( Olophrum ; characters; central Europe).

— Smetana, 1964c: 171 ( Olophrum ; characters).

— Smetana, 1964d: 54 ( Olophrum ; Czechoslovakia).

— Iablokoff-Khnzorian, 1964: 159 ( Olophrum ; characters).

— Szujecki, 1968a: 712 ( Olophrum ; Poland).

— Pope, 1977: 22 ( Olophrum ; Britain).

— Coiffait, 1978f: 277 ( Olophrum ; characters; France).

— Tóth, 1982: 88 ( Olophrum ; characters; Hungary).

— Zanetti, 1987: 303 ( Olophrum ; characters; Italy).

— Lucht, 1987: 89 ( Olophrum ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux; Denmark; Sweden).

— Boháč, 1993: 41 ( Olophrum ; Slovakia; Czech Republic).

— Zanetti, 1995: 8 ( Olophrum ; Italy?).

— Cibuļskis, 1999: 35 ( Olophrum ; list; Latvia).

— Assing, 2001: 75 ( Olophrum ; collected from xerothermous site; Germany).

- DISTRIBUTION: Europe, Russia, Georgia, Syria.

austriacum Scheerpeltz, 1929: 35 ( Olophrum ; Type locality: Traunauen bei Ebelsberg bei Linz in Oberösterreich).

— Horion, 1935: 171 ( Olophrum ; characters; Germany).

— Horion, 1963: 110 ( Olophrum ; Germany; Austria).

— Lohse, 1964: 54 ( Olophrum ; characters; central Europe).

— Smetana, 1964c: 171 ( Olophrum ; characters).

— Tóth, 1982: 85 ( Olophrum ; characters; Hungary).

— Lucht, 1987: 89 ( Olophrum ; checklist; Germany; Austria).

- DISTRIBUTION: Germany, Austria, Hungary.

bernhaueri Scheerpeltz, 1929: 80 ( Olophrum ; Type locality: Kaukasus, Kasbekp.).

- DISTRIBUTION: Georgia.

bernhauerianum Scheerpeltz, 1929 , see: consimile Gyllenhal, 1810 .

blattoides Ahrens, 1812 , see: piceum Gyllenhal, 1810 .

bodemeyeri Scheerpeltz, 1929: 94 ( Olophrum ; Type locality: Luristan, Persien).

- DISTRIBUTION: Iran.

boreale Paykull, 1792: 146 ( Staphylinus ; Type locality: Bothnia occidentali).

— Paykull, 1800: 411 ( Staphylinus ; [Note: Bernhauer and Schubert, 1910: 66 cited the female of boreale Paykull, 1800 as a synonym of consimile ]; characters; Sweden).

— Gravenhorst, 1806: 213 ( Omalium ; synonym of quadrum Gravenhorst ).

— Gyllenhal, 1810: 198 ( Omalium ; characters; western Bothnia region).

— Gyllenhal, 1827: 461 ( Omalium ).

— C. Sahlberg, 1827: 277 ( Omalium ; characters; Lapland).

— Zetterstedt, 1828: 48 ( Omalium ; characters; Lapland).

— Mannerheim, 1830: 51 ( Omalium ; Lapland).

— Mannerheim, 1831: 465 ( Omalium ; Lapland).

— Zetterstedt, 1838: 49 ( Omalium ; characters; habitat; Lapland).

— Erichson, 1840: 866 ( Olophrum ; characters; Finland).

— Thomson, 1861: 193 ( Olophrum ; characters; Scandinavia).

— Seidlitz, 1875: 240 ( Olophrum ; characters; Baltic region).

— J. Sahlberg, 1876: 212 ( Olophrum ; Finland).

— Luze, 1905: 43 ( Olophrum ; characters; Lapland; Finland; Norway).

— Poppius, 1909: 5 ( Olophrum ; Lena Valley, Siberia ).

— Bernhauer and Schubert, 1910: 66 ( Olophrum ; catalog).

— Scheerpeltz, 1929: 124 ( Olophrum ; characters; Norway, Sweden, Finland, Russia).

— Palm, 1948: 107 ( Olophrum ; characters; Sweden; Norway; Finland).

— Ashworth, 1972: 214 ( Olophrum ; late-glacial subfossil; Britain).

— Campbell, 1983: 602 ( Olophrum ; characters; Alberta; British Columbia ; Manitoba; Northwest Territories; Quebec; Yukon Territory; Alaska; Montana; Utah; Wyoming).

— Lohse and Lucht, 1989: 126 ( Olophrum ; note).

— Ryabukhin, 1991b: 14 ( Olophrum ; characters; Russia).

— Elias, 1991: 556 ( Olophrum ; Quaternary fossil collection; Colorado).

— Downie and Arnett, 1996: 432 ( Olophrum ; characters; Canada; USA).

— Lemdahl and Gustavsson, 1997: 181 ( Olophrum ; collected in lateglacial and middle Holocene deposits; Sweden).

— Ryabukhin, 1999: 23 ( Olophrum ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Sweden, Norway, Finland, Russia, Canada, USA.

nigropiceum Motschulsky, 1860b: 119 ( Olophrum ; Type locality: Kamtschatka).

— Motschulsky, 1859: 491 ( Olophrum ; Amur River).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; catalog).

— Scheerpeltz, 1929: 120 ( Olophrum ; characters; Siberia, Norway).

— Munster, 1935: 29, 30 ( Olophrum ; characters; variety of boreale ; notes).

— Steel, 1964a: 248 ( Olophrum ; confirmation of synonymy with nigropiceum ).

— Campbell, 1983: 602 ( Olophrum ; synonym of boreale ).

minor Poppius, 1909 a: 15 ( Olophrum ; variety of boreale ; [preoccupied]; Type locality: Tarhanowa).

helleni Scheerpeltz, 1929: 128 ( Olophrum ; Type locality: nördlichstes Norwegen: Finmarken , Varanger-Fjord , Tana-Fjord , Vadsö ; nördlichstes Schweden: Torne-Träsk ; nördlichstes Finnland: Enare-See ; Halbinse Kola; Halbinsel Kanin).

— Munster, 1935: 29, 30 ( Olophrum ; variety of boreale ; notes).

— Campbell, 1983: 603 ( Olophrum ; synonym of boreale ).

quesneli Hatch, 1957: 62 ( Olophrum ; Type locality: B. C. [ British Columbia ]: Barkerville).

— Campbell, 1983: 603 ( Olophrum ; synonym of boreale ).

brevicolle Bernhauer, 1903 , see: latum Mäklin, 1853 .

bulgaricum Scheerpeltz, 1929: 37 ( Olophrum ; Type locality: Kalofer, Tundza, Nordbulgarien).

- DISTRIBUTION: Bulgaria.

cascadense Campbell, 1983: 596 ( Olophrum ; Type locality: California: Mt. Lassen N. P., Kings Creek, 7500 feet. Also cited from Oregon).

- DISTRIBUTION: USA.

caucasicum Fauvel, 1875 a:VII [= 1875b: 209] ( Olophrum ; Type locality: Caucase).

— Luze, 1905: 39 ( Olophrum ; characters; Caucasus).

— Bernhauer and Schubert, 1910: 66 ( Olophrum ; catalog).

— Scheerpeltz, 1929: 67 ( Olophrum ; characters; Caucasus).

— Tikhomirova, 1973a: 139 ( Olophrum ; checklist; Caucasus).

- DISTRIBUTION: Georgia, Azerbaijan.

celatum † Scudder, 1900a: 86 ( Olophrum ; pleistocene fossil; Type locality: Scarborough).

- DISTRIBUTION: Canada.

consimile Gyllenhal, 1810: 199 ( Omalium ; Type locality: Bothnia occidentalis).

— Paykull, 1800: 411 ( Staphylinus ; [Note: Misidentification: Bernhauer and Schubert, 1910: 66 cited the female of boreale Paykull, 1800 as a synonym of consimile ]; characters; Sweden).

— Gyllenhal, 1827: 461 ( Omalium ).

— C. Sahlberg, 1827: 277 ( Omalium ; characters; Finland).

— Zetterstedt, 1828: 48 ( Omalium ; characters; Lapland).

— Mannerheim, 1830: 51 ( Omalium ; Finland).

— Mannerheim, 1831: 465 ( Omalium ; Finland).

— Stephens, 1834: 341 ( Anthobium ; characters; Scotland).

— Stephens, 1839: 427 ( Olophrum ; characters; England).

— Erichson, 1840: 867 ( Olophrum ; characters; Lapland; Russia).

— Redtenbacher, 1857: 993 ( Olophrum ; characters; Austria).

— Kraatz, 1857d: 941 ( Olophrum ; characters; Germany).

— Thomson, 1861: 193 ( Olophrum ; characters; Scandinavia).

— Fauvel, 1871a: 125 [= 1872: 99] ( Olophrum ; characters; Britain; Germany; Russia; Lapland; Scandinavia).

— LeConte, 1873: 327 ( Omalium ; characters).

— Redtenbacher, 1874: 268 ( Olophrum ; characters).

— Seidlitz, 1875: 240 ( Olophrum ; characters; Baltic region).

— J. Sahlberg, 1876: 212 ( Olophrum ; Finland).

— Fowler, 1888: 405 ( Olophrum ; characters; Britain).

— Ganglbauer, 1895: 721 ( Olophrum ; characters; north Europe; England; Germany).

— Luze, 1905: 45 ( Olophrum ; characters; Germany; Finland; Sweden; Norway; England; Lapland; Russia; Siberia).

— Poppius, 1909: 5 ( Olophrum ; Lena Valley, Siberia ).

— Reitter, 1909: 186 ( Olophrum ; characters; Germany).

— Bernhauer and Schubert, 1910: 66 ( Olophrum ; catalog).

— Notman, 1919a: 95 ( Olophrum ; Ontario).

— Bernhauer, 1926g: 2 ( Olophrum ; Kamchatka region).

— Scheerpeltz, 1929: 155 ( Olophrum ; characters; Germany; Sweden; Norway; Finland; Russia; Urals; Siberia; Alaska; Vancouver; Alberta; Quebec; Maine; New York).

— Munster, 1936: 97 ( Olophrum ; characters; notes).

— Palm, 1948: 107 ( Olophrum ; characters; Sweden; Norway; Finland).

— Hatch, 1957: 61 ( Olophrum ; characters; British Columbia ; Oregon; Idaho).

— Horion, 1963: 115 ( Olophrum ; Russia; Finland; Sweden; Norway; Britain; Poland; Czech Republic; Germany).

— Lohse, 1964: 54 ( Olophrum ; characters; central Europe).

— Steel, 1964a: 248 ( Olophrum ; notes).

— Smetana, 1964c: 171 ( Olophrum ; characters).

— Pope, 1977: 22 ( Olophrum ; Britain).

— Schwert and Morgan, 1980: 101 ( Olophrum ; late glacial fossil; New York).

— Miller and Morgan, 1982: 264 ( Olophrum ; Postglacial fossils; New York).

— Campbell, 1983: 586 ( Olophrum ; characters; Alberta; British Columbia ; Manitoba; Northwest Territories; Quebec; Saskatchewan; Yukon Territory; Alaska; Arizona; Colorado; Idaho; Michigan; Montana; Nevada; Oregon; South Dakota; Utah; Washington; Wyoming).

— Schwert, Anderson, Morgan, Morgan, and Karrow, 1985: 216 ( Olophrum ; Quaternary fossil; Ontario).

— Lucht, 1987: 89 ( Olophrum ; checklist; Germany; Poland; Czechoslovakia; Austria; Sweden).

— Schülke and Uhlig, 1988: 3 ( Olophrum ; notes; Germany).

— Ryabukhin, 1991b: 17 ( Olophrum ; characters; Russia).

— Elias, 1991: 556 ( Olophrum ; Quaternary fossil collection; Colorado).

— Boháč, 1993: 41 ( Olophrum ; Czech Republic).

— Zanetti, 1995: 8 ( Olophrum ; Italy).

— Downie and Arnett, 1996: 433 ( Olophrum ; characters; Canada; USA).

— Lavoie, Elias, and Payette, 1997: 231 ( Olophrum ; Holocene fossil; subarctic Quebec).

— Ryabukhin, 1999: 22 ( Olophrum ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Europe, Russia, Canada, USA.

marginatum Kirby, 1837: 89 ( Omalium ; [preoccupied]; Type locality: America Borealis : Lat. 65°).

— Fauvel, 1878d: 244 [= 1878e: 80] ( Olophrum ; characters; Alaska; Lake superior; Michigan).

— LeConte, 1879: 510 ( Olophrum ; characters; Colorado; Alaska).

— Bernhauer and Schubert, 1910: 65 ( Lathrimaeum ; synonym of sordidum ).

— Blair, 1933: 94 ( Olophrum ; synonym of boreale ).

— Moore, 1966: 55 ( Anthobium ). alpinum Heer, 1839: 190 ( Acidota ; Type locality: Seeloch auf der Alp Mühlebach; auf dem Luchmanier; Zaportalp; Canälpass am Gletscherrand; St. Bernhard; Chamouni).

— Fairmaire and Laboulbène, 1856: 635 ( Olophrum ; characters; France).

— Kraatz, 1857d: 941 ( Olophrum ; synonym of alpestre ).

— Fauvel, 1871a: 125 [= 1872: 99] ( Olophrum ; characters; France; Switzerland; Germany).

— Mulsant and Rey, 1880: 141 ( Olophrum ; characters; France).

— Ganglbauer, 1895: 720 ( Olophrum ; characters; Alps; Pyrenees; Sudeten).

— Luze, 1905: 44 ( Olophrum ; characters; Alps; Pyrenees; Sudeten).

— Reitter, 1909: 186 ( Olophrum ; characters; Germany).

— Bernhauer and Schubert, 1910: 66 ( Olophrum ; catalog).

— Scheerpeltz, 1929: 132 ( Olophrum ; characters; Alps).

— Portevin, 1929: 440 ( Olophrum ; characters; France).

— Munster, 1936: 97–100 ( Olophrum ; variety of consimile ; discussion).

— Palm, 1948: 107 ( Olophrum ; characters; Sweden).

— Horion, 1963: 114 ( Olophrum ; Italy; France; northern Carpathians; Czech Republic; Austria). — Lohse, 1964: 55 ( Olophrum ; characters; central Europe).

— Smetana, 1964c: 171 ( Olophrum ; characters).

— Coiffait, 1978f: 278 ( Olophrum ; characters; France).

— Tóth, 1982: 86 ( Olophrum ; characters).

— Campbell, 1983: 586 ( Olophrum ; synonym of consimile ).

— Zanetti, 1987: 305 ( Olophrum ; characters; Italy).

— Lucht, 1987: 89 ( Olophrum ; checklist; Germany; Poland; Czechoslovakia; Austria).

— Boháč, 1993: 41 ( Olophrum ; cited as valid species; Slovakia; Czech Republic).

— Zanetti, 1995: 8 ( Olophrum ; synonym of consimile ).

alpestre Erichson, 1840: 867 ( Olophrum ; Type locality: Helvetiae alpes).

— Heer, 1841: 571 ( Olophrum ; synonym of alpina ).

— Redtenbacher, 1849: 828 ( Olophrum ; characters; Austria).

— Fairmaire and Laboulbène, 1856: 635 ( Olophrum ; synonym of alpinum ).

— Redtenbacher, 1857: 248 ( Olophrum ; characters; Austria).

— Kraatz, 1857d: 941 ( Olophrum ; characters; Germany).

— Fauvel, 1865a: 318 [= 1865b: 68] ( Olophrum ; habitat; France).

— Fauvel, 1871a: 125 [= 1872: 99] ( Olophrum ; synonym of alpinum ).

— Redtenbacher, 1874: 268 ( Olophrum ; characters; Austria).

— Seidlitz, 1875: 240 ( Olophrum ; characters; Baltic region).

— Mulsant and Rey, 1880: 142 ( Olophrum ; synonym of alpinum ).

— Ganglbauer, 1895: 720 ( Olophrum ; synonym of alpinum ).

— Luze, 1905: 44 ( Olophrum ; synonym of alpinum ).

— Bernhauer and Schubert, 1910: 66 ( Olophrum ; synonym of alpinum ).

— Campbell, 1983: 586 ( Olophrum ; synonym of consimile ).

— Zanetti, 1987: 305 ( Olophrum ; synonym of alpinum ).

parvulum Mäklin, 1853: 195 ( Olophrum ; Type locality: lacum Thkujabna in peninsula Kenai ; [Note: See lectotype designation by Campbell, 1983]).

— Fauvel, 1878d: 245 [= 1878e: 81] ( Olophrum ; characters; Alaska).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; catalog).

— Scheerpeltz, 1929: 149 ( Olophrum ; characters; Alaska).

— Steel, 1964a: 248, 250 ( Olophrum ; appears to be small form and synonym of consimile ).

— Campbell, 1983: 586 ( Olophrum ; lectotype designation; synonym of consimile ).

marginatum Mäklin, 1853: 196 ( Olophrum ; [preoccupied]; Type locality: peninsula Kenai ; [Note: See lectotype designation by Campbell, 1983]).

— Fauvel, 1869: 493 ( Olophrum ; synonym of consimile ).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; catalog).

— Scheerpeltz, 1929: 152 ( Olophrum ; characters; Alaska).

— Munster, 1936: 100 ( Olophrum ; variety of consimile ).

— Steel, 1964a: 248 ( Olophrum ; synonym of consimile ).