Phloeonomus rhopalocerus, Cameron, 1928

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

rhopalocerus Cameron, 1928 , see: Paraphloeostiba .

ruficollis Bernhauer, 1934 h: 484 ( Phloeonomus ; Type locality: Cape Province: Montagu; Worcester ).

- DISTRIBUTION: South Africa.

rufotestaceus Bernhauer, 1908c: 102 ( Phloeonomus ; Type locality: Britisch-Südafrika, Kapland : Cape flats).

— Bernhauer and Schubert, 1910: 59 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

- DISTRIBUTION: South Africa.

secretus Cameron, 1928 , see: Paraphloeostiba .

sjobergi A. Strand, 1937: 111 ( Phloeonomus ; Type locality: Oslo) .

— Palm, 1948: 86 ( Phloeonomus ; characters; Sweden; Norway; Finland).

— Lohse, 1964: 47 ( Phloeonomus ; subgenus Phloeonomus ; characters; central Europe).

— Smetana, 1964c: 170 ( Phloeonomus ; subgenus Phloeonomus ; characters).

— Smetana, 1981c: 80 ( Phloeonomus ; characters).

— Lucht, 1987: 87 ( Phloeonomus ; in checklist).

— Assing, 1998f: 124 ( Phloeonomus ; note).

— Burakowski, Mroczkowski, and Stefańska, 2000: 41 ( Phloeonomus ; subgenus Phloeonomus ; catalog; Poland).

- DISTRIBUTION: Norway, Sweden, Finland.

sonani Bernhauer, 1943 , see: Paraphloeostiba .

sordidus Sharp, 1887 , see: centralis Blackwelder, 1945 .

specularis Bernhauer, 1915 , see: Paraphloeostiba .

subapicalis Cameron, 1928d: 99 ( Phloeonomus ; Type locality: Sumatra: Fort de Kock ).

— Scheerpeltz, 1933: 1051 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

- DISTRIBUTION: Indonesia.

sublucidus Cameron, 1929b: 441 ( Phloeonomus ; Type locality: Selangor: Bukit Kutu ).

— Scheerpeltz, 1933: 1051 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

- DISTRIBUTION: Malaysia.

subopaca Cameron, 1924 , see: Paraphloeostiba .

suffusus Casey, 1894: 424 ( Omalium ; subgenus Phloeonomus ; Type locality: Alaska: Hunter’s Bay, Prince of Wales Island).

— Bernhauer and Schubert, 1910: 59 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

- DISTRIBUTION: USA.

sumatrensis Bernhauer, 1905 , see: Paraphloeostiba .

tasmanicus Blackburn, 1902: 27 ( Omalium , cited as Homalium ; Type locality: Tasmania).

— Bernhauer and Schubert, 1910: 57 ( Omalium ; subgenus Omalium ; catalog).

— Scheerpeltz, 1933: 1049 ( Phloeonomus ; subgenus Phloeostiba ; catalog).

- DISTRIBUTION: Australia.

temporalis Cameron, 1930: 148 ( Phloeonomus ; subgenus Phloeostiba ; Type locality: Manipur: Shugnu, alt. 3000 feet).

— Scheerpeltz, 1933: 1049 ( Phloeonomus ; subgenus Phloeostiba ; catalog).

- DISTRIBUTION: India.

tonkinensis Cameron, 1940 , see: Paraphloeostiba .

toxopeanus Bernhauer, 1926 , see: Xanthonomus .

tricolor Wollaston, 1865: 523 , 75 appendix ( Omalium , cited as Homalium ; Type locality: Madeira: Ribeira de São Jorge).

— Fauvel, 1902b: 53 ( Omalium , cited as Homalium ; Madeira).

— Luze, 1906: 594 ( Phloeonomus ; characters; Madeira).

— Bernhauer and Schubert, 1910: 58 ( Phloeonomus ; subgenus Xylostiba ; catalog).

tristis Sharp, 1887: 745 ( Omalium ; Type locality: Guatemala, Totonicapam 10,500 feet).

— Bernhauer and Schubert, 1910: 59 ( Phloeonomus ; subgenus Phloeonomus ; catalog).

- DISTRIBUTION: Guatemala.

turneri Bernhauer, 1934 h: 485 ( Phloeonomus ; Type locality: Pondoland : Port St. Johns).

- DISTRIBUTION: South Africa.

ulmi Motschulsky, 1857 , see: Hapalaraea .

usambarae Fauvel, 1904 , see: Phloeostiba .

xanthopterus Fauvel, 1865 , see: praeustus Motschulsky, 1857 .

Phloeostiba

[3 species; Nearctic, Palaearctic, Ethiopian , and Oriental Regions]

Phloeostiba Thomson, 1858: 38 (species included: plana ). TYPE SPECIES: Staphylinus planus Paykull , fixed by monotypy.

— Thomson, 1859: 51 (characters; type species: plana ).

— Thomson, 1861: 208 (characters).

— Mulsant and Rey, 1880: 171 (characters; key to French species).

— Heyden, 1880: 83 (subgenus of Omalium , cited as Homalium ; list of species of Siberian region).

— Seidlitz, 1891: 338 (key to some European species).

— Seidlitz, 1891a: 361 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 120 (subgenus of Omalium ; list of species of Europe and Caucasus).

— Heyden, 1893: 49 (subgenus of Omalium ; list of species of Siberian region).

— Heyden, 1896: 37 (subgenus of Omalium ; list of species of Siberian region).

— Luze, 1906: 589 (synonym of Phloeonomus ).

— Heyden, Reitter, and Weise, 1906: 132 (subgenus of Phloeonomus ; list of species of Europe and Caucasus).

— Reitter, 1909: 189 (characters; key to German species).

— Bernhauer and Schubert, 1910: 58 (subgenus of Phloeonomus ).

— Johansen, 1914: 605 (characters; key to Danish species).

— Cameron, 1925: 11 (subgenus of Omalium ; catalog of Indian species).

— Portevin, 1929: 444 (characters; key to French species).

— Cameron, 1930: 141, 147 (subgenus of Phloeonomus ; characters in key; key to species of British India).

— Joy, 1932: 103 (key to British species).

— Scheerpeltz, 1933: 1048 (world catalog supplement; subgenus of Phloeonomus ).

— Blackwelder, 1943: 51 (subgenus of Phloeonomus ; type species: plana ).

— Kloet and Hincks, 1945: 161 (synonym of Phloeonomus ).

— Tottenham, 1949: 355, 407 (subgenus of Phloeonomus ; type species: plana Paykull ; list of British species).

— Blackwelder, 1952: 306 (type species: plana ).

— Horion, 1963: 90 (subgenus of Phloeonomus ; list of central European species).

— Lohse, 1964: 46 (subgenus of Phloeonomus ; characters; key to central European species).

— Steel, 1970a: 15 (cited as valid genus; larval characters; biological notes).

— Tikhomirova, 1973a: 138 (subgenus of Phloeonomus ; checklist of species of USSR).

— Shibata, 1976: 105 (checklist of species of Japan).

— Pototskaia, 1976: 162 (larval characters).

— Pope, 1977: 23 (list of British species).

— Burakowski, Mroczkowski, and Stefańska, 1979: 47 (catalog; subgenus of Phloeonomus ; Poland).

— Watanabe, 1990: 111 (characters; species of Japan).

— Segers, 1986: 21 (checklist of species; Belgium).

— Zanetti, 1987: 243 (characters; habitus; key to species of Italy). — Zanetti, 1995: 7 (list of species of Italy). — M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 27 (additional records to Danish list of species). — M. Hansen, 1996: 94 (list of Danish species). — Assing, 1996a: 180, 181 (characters; notes). — Assing, 1998f: 124 (valid genus; characters). — Newton, Thayer, Ashe, and Chandler, 2000: 337 ( Nearctic species; notes; characters in key).

Distemmus LeConte, 1861 a: 69 (species included: argus ). TYPE SPECIES: Trogophloeus argus LeConte , fixed by original designation and monotypy. — LeConte, 1863: 26 (list of North American species). — Gemminger and Harold, 1868: 655 (catalog). — Fauvel, 1875a: IV [= 1875b: 206] (catalog; group of Omalium , cited as Homalium ). — Fauvel, 1878d: 205 [= 1878e: 41] (synonym of Omalium , cited as Homalium ). — Luze, 1906: 589 (synonym of Phloeonomus ). — Bernhauer and Schubert, 1910: 57 (synonym of Phloeonomus ). — Blackwelder, 1943: 51 (synonym of Phloeonomus ; type species: argus ). — Blackwelder, 1952: 130 (type species: argus ; synonym of Phloeonomus ). — Watanabe, 1990: 99 (synonym of Phloeonomus ).

argus LeConte, 1850 , see: lapponica Zetterstedt, 1838 .

bipunctata Motschulsky, 1860 , see: plana Paykull, 1792 .

conformis Kraatz, 1857 , see: lapponica Zetterstedt, 1838 .

flavipes Linné, 1758 , see: plana Paykull, 1792 .

lapponica Zetterstedt, 1838: 73 ( Oxytelus ; Type locality: Lapponia Umensis: Lycksele). — Zetterstedt, 1838: 52 ( Omalium ; [Note: Misidentification: Bernhauer and Schubert, 1910: 58 cited planus Zetterstedt as a synonym of lapponicus Zetterstedt ; Zetterstedt did not describe the species as new, he attributed it to Gravenhorst who attributed it to Paykull]; characters; habitat; Lapland; Sweden). — Thomson, 1861: 209 ( Phloeostiba ; synonym of plana ). — Fauvel, 1871a: 95 [= 1872: 69] ( Omalium , cited as Homalium ; characters; France; Lapland; Scandinavia; Britain; Germany; Poland; Switzerland; Siberia; North America ). — Seidlitz, 1875: 233 ( Phloeostiba ; characters; Baltic region). — J. Sahlberg, 1876: 219 ( Phloeostiba ; Finland). — Fauvel, 1878d: 214 [= 1878e: 50] ( Omalium , cited as Homalium ; characters; Alaska; Colorado; Lake Superior). — Mulsant and Rey, 1880: 176 ( Phloeostiba ; characters; France). — Ganglbauer, 1895: 732 ( Phloeonomus ; subgenus Phloeostiba ; characters; north and middle Europe; Siberia; North America ). — Luze, 1906: 598 ( Phloeonomus ; characters; northern and central Europe; Siberia; British Columbia ). — Fall and Cockerell, 1907: 168 ( Omalium , cited as Homalium ; New Mexico). — Poppius, 1909: 4 ( Phloeonomus ; subgenus Phloeostiba ; Lena Valley, Siberia ). — Reitter, 1909: 190 ( Phloeostiba characters; Germany). — Bernhauer and Schubert, 1910: 58 ( Phloeonomus ; subgenus Phloeostiba ; catalog). — Notman, 1919a: 96 ( Phloeonomus ; Ontario). — Fall, 1926: 145 ( Phloeonomus ; Alaska, Yukon Territory). — Portevin, 1929: 445 ( Phloeostiba ; characters; France). — Scheerpeltz, 1933: 1048 ( Phloeonomus ; subgenus Phloeostiba ; catalog). — Palm, 1948: 86 ( Phloeonomus ; characters; Sweden; Denmark; Norway; Finland). — Jarrige, 1950a: 95 ( Phloeonomus , cited as Phlaeonomus; subgenus Phloeostiba ; characters; France). — Hatch, 1957: 81 ( Phloeonomus ; subgenus Phloeostiba ; characters; British Columbia ; Washington). — Horion, 1963: 91 ( Phloeonomus ; subgenus Phloeostiba ; Britain; Norway; Sweden; Finland;

— Lohse, 1964: 46 ( Phloeonomus ; subgenus Phloeostiba ; characters; central Europe).

— Smetana, 1964c: 163 ( Phloeonomus ; subgenus Phloeostiba ; Czechoslovakia).

— Smetana, 1964c: 169 ( Phloeonomus ; subgenus Phloeostiba ; characters).

— Szujecki, 1968a: 711 ( Phloeonomus ; Poland).

— Steel, 1970a: 15 ( Phloeostiba ; biological notes).

— Smetana 1975c: 158 ( Phloeonomus ; Mongolia).

— Pope, 1977: 23 ( Phloeostiba ; Britain).

— Tóth, 1982: 68 ( Phloeonomus ; characters; Hungary).

— Zanetti, 1987: 246 ( Phloeostiba ; characters; Italy).

— Lucht, 1987: 87 ( Phloeonomus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark).

— Welch, 1993: 227 ( Phloeostiba ; ovariole number and ovary structure).

— Boháč, 1993: 41 ( Phloeostiba ; Slovakia; Czech Republic).

— Zanetti, 1995: 7 ( Phloeostiba ; Italy).

— Downie and Arnett, 1996: 438 ( Phloeonomus ; characters; Canada; USA).

— Fowles, Alexander, and Key, 1999: 129 ( Phloeostiba ; saproxylic species; status as endangered or threatened species; Britain).

— Cibuļskis, 1999: 35 ( Phloeonomus ; list; Latvia).

— Ryabukhin, 1999: 18 ( Phloeostiba ; catalog; habitat; distribution; northeastern Russia).

— Hågvar, 1999: 28 ( Phloeonomus ; collected on Fomes fomentarius ; Norway).

— Tronquet, 1999a: 213 ( Phloeostiba ; notes; France).

— Olberg and Andersen, 2000: 223 ( Phloeonomus ; attracted to trunk-window traps baited with

Fomes fomentarius and Phellinus spp. ; Norway).

- DISTRIBUTION: Europe, Russia, Mongolia, Canada, USA. argus LeConte, 1850: 220 ( Trogophloeus ; Type locality: Lake Superior).

— Ganglbauer, 1895: 732 ( Phloeonomus ; synonym of lapponicus ).

— Luze, 1906: 598 ( Phloeonomus ; synonym of lapponicus ).

— Bernhauer and Schubert, 1910: 58 ( Phloeostiba ; synonym of lapponicus ).

— Jarrige, 1950a: 95 ( Phloeonomus , cited as Phlaeonomus; synonym of lapponicus ). planipennis Mäklin, 1853: 198 ( Omalium ; Type locality: insula Sitkha; peninsula Kenai ).

— Fauvel, 1871a: 95 [= 1872: 69] ( Omalium , cited as Homalium ; synonym of lapponicum ).

— LeConte, 1873: 327 ( Omalium ; synonym of pineti ).

— J. Sahlberg, 1876: 219 ( Phloeostiba ; synonym of lapponica ).

— Fauvel, 1878d: 214 [= 1878e: 50] ( Omalium , cited as Homalium ; synonym of lapponicum ).

— Ganglbauer, 1895: 732 ( Phloeonomus ; synonym of lapponicus ).

— Luze, 1906: 598 ( Phloeonomus ; synonym of lapponicus ).

— Bernhauer and Schubert, 1910: 58 ( Phloeostiba ; synonym of lapponica ).

— Jarrige, 1950a: 95 ( Phloeonomus , cited as Phlaeonomus; synonym of lapponicus ). pineti Thomson, 1857 a: 227 ( Omalium ; Type locality: Not cited, presumably Sverige).

— Thomson, 1861: 209 ( Phloeostiba ; characters; Scandinavia).

— Mulsant and Rey, 1880: 177 ( Phloeostiba ; synonym of lapponica ).

— Fowler, 1888: 421 ( Omalium , cited as Homalium ; subgenus Phloeostiba ; characters; Britain).

— Ganglbauer, 1895: 732 ( Phloeonomus ; synonym of lapponicus ).

— Luze, 1906: 598 ( Phloeonomus ; synonym of lapponicus ).

— Bernhauer and Schubert, 1910: 58 ( Phloeostiba ; synonym of lapponica ).

— Jarrige, 1950a: 95 ( Phloeonomus , cited as Phlaeonomus; synonym of lapponicus ).

— Legner and Moore, 1977: 175 ( Phloeonomus ; cited as peneti; found under bark).

— Pope, 1977: 23 ( Phloeostiba ; synonym of lapponica ).

— Zanetti, 1987: 246 ( Phloeostiba ; synonym of lapponica ). conformis Kraatz, 1857d: 987 ( Omalium ; Type locality: Baiern).

— Redtenbacher, 1857: 994 ( Omalium ; characters; Germany).

— Thomson, 1861: 209 ( Phloeostiba ; synonym of plana ).

— Fauvel, 1871a: 95 [= 1872: 69] ( Omalium , cited as Homalium ; synonym of lapponicum ).

— Redtenbacher, 1874: 277 ( Omalium ; characters). — Mulsant and Rey, 1880: 177 ( Phloeostiba ; synonym of lapponica ).

— Ganglbauer, 1895: 732 ( Phloeonomus ; synonym of lapponicus ).

— Luze, 1906: 598 ( Phloeonomus ; synonym of lapponicus ).

— Bernhauer and Schubert, 1910: 58 ( Phloeostiba ; synonym of lapponicus ).

— Jarrige, 1950a: 95 ( Phloeonomus , cited as Phlaeonomus; synonym of lapponicus ).

— Zanetti, 1987: 246 ( Phloeostiba ; synonym of lapponica ).

subtilis Kraatz, 1857d: 987 ( Omalium ; Type locality: Sächsische Schweiz).

— Redtenbacher, 1857: 994 ( Omalium ; characters; Germany).

— Fauvel, 1869: 494 ( Omalium , cited as Homalium ; synonym of lapponicum ).

— Fauvel, 1871a: 95 [= 1872: 69] ( Omalium , cited as Homalium ; synonym of lapponicum ).

— Redtenbacher, 1874: 278 ( Omalium ; characters).

— Ganglbauer, 1895: 732 ( Phloeonomus ; synonym of lapponicus ).

— Luze, 1906: 598 ( Phloeonomus ; synonym of lapponicus ).

— Bernhauer and Schubert, 1910: 58 ( Phloeostiba ; synonym of lapponica ).

— Zanetti, 1987: 246 ( Phloeostiba ; synonym of lapponica ).

pineti Thomson, 1857 , see: lapponica Zetterstedt, 1838 .

planipennis Mäklin, 1853 , see: lapponica Zetterstedt, 1838 .

plana Paykull, 1792: 145 ( Staphylinus ; Type locality: Uplandia).

— Paykull, 1800: 405 ( Staphylinus ; characters; Sweden).

— Gravenhorst, 1802: 112 ( Omalium ; characters).

— Latreille, 1804: 370 ( Omalium ; characters; Germany; Sweden).

— Gravenhorst, 1806: 204 ( Omalium ; characters).

— Gyllenhal, 1810: 221 ( Omalium ; characters; Sweden).

— Olivier, 1811: 475 ( Omalium ; characters; Europe).

— Gyllenhal, 1827: 466 ( Omalium ).

— Zetterstedt, 1828: 53 ( Omalium ; characters; Lapland).

— Curtis, 1829: 29 ( Omalium ; catalog; Britain).

— C. Sahlberg, 1830: 284 ( Omalium ; characters; Finland).

— Mannerheim, 1830: 54 ( Anthobium ; Finland; Sweden).

— Mannerheim, 1831: 468 ( Anthobium ; Finland; Sweden).

— Stephens, 1834: 346 ( Omalium ; characters; England).

— Lacordaire, 1835: 475 ( Anthobium ; characters; France).

— Erichson, 1839a: 636 ( Omalium ; characters; Germany).

— Heer, 1839: 175 ( Xylodromus ; characters; Switzerland).

— Stephens, 1839: 427 ( Omalium ; characters; England).

— Erichson, 1840: 886 ( Omalium ; characters; Europe).

— Redtenbacher, 1849: 753 ( Omalium ; characters; Austria).

— Thomson, 1857a: 227 ( Omalium ; characters).

— Fairmaire and Laboulbène, 1856: 644 ( Omalium ; characters; France).

— Redtenbacher, 1857: 253 ( Omalium ; characters; Austria).

— Kraatz, 1857d: 986 ( Omalium ; characters; Germany).

— Thomson, 1858: 38 ( Phloeostiba ; type species).

— Thomson, 1859: 51 (type species of Phloeostiba ).

— Thomson, 1861: 208 ( Phloeostiba ; characters; Scandinavia).

— Fauvel, 1871a: 96 [= 1872: 70] ( Omalium , cited as Homalium ; characters; France).

— Redtenbacher, 1874: 276 ( Omalium ; characters; Austria).

— Seidlitz, 1875: 233 ( Phloeostiba ; characters; Baltic region).

— J. Sahlberg, 1876: 219 ( Phloeostiba ; Finland).

— Mulsant and Rey, 1880: 173 ( Phloeostiba ; characters; France).

— Gozis, 1886: 16 (type species of Omalium ).

— Fowler, 1888: 421 ( Omalium , cited as Homalium ; subgenus Phloeostiba ; characters; Britain). — Ganglbauer, 1895: 732 ( Phloeonomus ; subgenus Phloeostiba ; characters; Europe; Siberia). — Luze, 1906: 596 ( Phloeonomus ; characters; Europe; Siberia).

— Sainte-Claire Deville, 1906: 70 ( Omalium , cited as Homalium ; Corsica).

— Bernhauer and Schubert, 1910: 58 ( Phloeonomus ; subgenus Phloeostiba ; catalog).

— Johansen, 1914: 606 ( Phloeostiba ; characters; Denmark).

— Portevin, 1929: 444 ( Phloeostiba ; characters; France).

— Scheerpeltz, 1933: 1049 ( Phloeonomus ; subgenus Phloeostiba ; catalog).

— Hinton, 1945: 50 ( Phloeonomus ; subgenus Phloeostiba ; characters).

— Palm, 1948: 85 ( Phloeonomus ; characters; Sweden; Denmark; Norway; Finland).

— Tottenham, 1949: 407 ( Phloeonomus ; checklist; Britain).

— Jarrige, 1950a: 94 ( Phloeonomus , cited as Phlaeonomus; subgenus Phloeostiba ; characters; France).

— Horion, 1963: 90 ( Phloeonomus ; subgenus Phloeostiba ; Transbaical; Scotland; Lapland; France to Poland and Russia; Iberian Peninusula; Sardinia; Italy; Balkans to Greece;

Germany; Austria).

— Lohse, 1964: 46 ( Phloeonomus ; subgenus Phloeostiba ; characters; central Europe).

— Smetana, 1964c: 169 ( Phloeonomus ; subgenus Phloeostiba ; characters).

— Szujecki, 1968a: 711 ( Phloeonomus ; Poland).

— Scheerpeltz, 1968a: 11 ( Phloeonomus ; catalog; Austria).

— Steel, 1970a: 15 ( Phloeostiba ; biological notes).

— Shibata, 1974: 34 ( Phloeostiba ; Japan).

— Legner and Moore, 1977: 175 ( Phloeonomus ; found under bark).

— Pope, 1977: 23 ( Phloeostiba ; Britain).

— Muona, 1979: 18 ( Phloeonomus ; Finland; Russia; Sweden; Norway; Denmark; Baltic States).

— Burakowski, Mroczkowski, and Stefańska, 1979: 46 ( Phloeonomus ; catalog; Poland).

— Hammond, 1980: 139 ( Phloeostiba ; Ireland).

— Tóth, 1982: 67 ( Phloeonomus ; characters; Hungary).

— Zanetti, 1987: 244 ( Phloeostiba ; characters; Italy).

— Ádám, 1987: 137 ( Phloeostiba ; collecting notes; Hungary).

— Filatova, 1981: 17 ( Phloeonomus ; Russia).

— Muona and Viramo, 1986: 15 ( Phloeonomus ; Finland).

— Lucht, 1987: 87 ( Phloeonomus ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark).

— Gusarov, 1989: 5 ( Phloeonomus ; habitat notes; Ukraine).

— Watanabe, 1990: 114 ( Phloeonomus ; characters; Japan).

— Silfverberg, 1992: 21 ( Phloeonomus ; Finland; Russia; Sweden; Norway; Denmark; Estonia; Lithuania).

— Boháč, 1993: 41 ( Phloeostiba ; Slovakia; Czech Republic).

— Sparacio, 1995: 141 ( Phloeostiba ; characters; summary of distribution; notes; Sicily).

— Zanetti, 1995: 7 ( Phloeostiba ; Italy).

— Sparacio, 1995: 141 ( Phloeostiba ; characters; notes; Sicily).

— Zanetti, 1995: 7 ( Phloeostiba ; Italy).

— Franzen, 1996: 206 ( Phloeonomus ; collecting notes; Germany).

— M. Hansen, 1996: 94 ( Phloeostiba ; checklist; Denmark).

— Ádám, 1996: 238 ( Phloeostiba ; synonym of flavipes ).

— Eisinger, 1997: 169 ( Phloeonomus ; collecting notes; Germany).

— T. Wagner, 1997: 234 ( Phloeonomus ; collecting notes; Germany).

— Anderson, Nash, and O’Connor, 1997: 15 ( Phloeostiba ; Ireland).

— Fowles, Alexander, and Key, 1999: 129 ( Phloeostiba ; saproxylic species; status as endangered or threatened species; Britain).

— Cibuļskis, 1999: 35 ( Phloeonomus ; list; Latvia).

— Ryabukhin, 1999: 19 ( Phloeostiba ; catalog; habitat; distribution; northeastern Russia).

— Herman, 2001: 21 ( Phloeostiba ; prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.3).

- DISTRIBUTION: Iceland, Europe, Russia, Japan.

flavipes Linné, 1758: 423 ( Staphylinus ; Type locality: Europa; [Note: This name is older than plana . Bernhauer and Schubert (1910, 1911, 1912, 1914, 1916, 1926) listed no flavipes described by Linné however they cited flavipes Fabricius twice; flavipes Fabricius, 1793: 527 was listed in Phloeonomus (Bernhauer and Schubert, 1910: 58) and flavipes Fabricius, 1775: 268 was cited in Tachinus (Bernhauer and Schubert, 1916: 483) . Fabricius credited the species to Linné (1767: 685) who attributed it to his earlier description (1758: 423). This confusion must be resolved by examination of the type]).

— Linné, 1760: 233 ( Staphylinus ; characters; Sweden).

— O. Müller, 1764: 23 ( Staphylinus ; characters).

— Linné, 1767: 685 ( Staphylinus ; characters; Europe).

— Füessly, 1775: 21 ( Staphylinus ).

— O. Müller, 1776: 97 ( Staphylinus ; characters).

— Goeze, 1777: 721 ( Staphylinus ; characters).

— Fabricius, 1793: 527 ( Staphylinus ; [Note: This reference was cited by Bernhauer and Schubert (1910: 58) as the original description for flavipes but Fabricius attributed the species to Linné]; characters; Selandia).

— Fabricius, 1801: 599 ( Staphylinus ; characters).

— Erichson, 1839a: 636 ( Omalium ; [Note: Attributed to Fabricius who attributed it to Linné]; characters; Germany).

— Heer, 1839: 175 ( Xylodromus ; [Note: Attributed flavipes to Fabricius who attributed it to Linné]; synonym of planus ).

— Erichson, 1840: 886 ( Omalium ; [Note: Attributed to Linné]; synonym of planum ).

— Fairmaire and Laboulbène, 1856: 644 ( Omalium ; [Note: Attributed to Linné]; synonym of planum ).

— Kraatz, 1857d: 986 ( Omalium ; synonym of planum ).

— Ganglbauer, 1895: 732 ( Phloeonomus ; attributed to Fabricius; synonym of planus ).

— Luze, 1906: 597 ( Phloeonomus ; synonym of planus ).

— Bernhauer and Schubert, 1910: 58 ( Phloeonomus ; synonym of planus ).

— Watanabe, 1990: 114 ( Phloeonomus ; synonym of planus ).

— Ádám, 1996: 238 ( Phloeostiba ; valid name; Hungary).

— Herman, 2001: 21 ( Phloeostiba ; synonym of plana ).

bipunctata Motschulsky, 1860: 545 ( Xylodromus ; Type locality: Pologne).

— Hochhuth, 1862: 105 ( Omalium ; characters; Poland).

— Luze, 1906: 486, 597 ( Phloeonomus ; synonym of planus ).

— Bernhauer and Schubert, 1910: 58 ( Phloeonomus ; synonym of planus ).

— Watanabe, 1990: 114 ( Phloeonomus ; synonym of planus ).

subtilis Kraatz, 1857 , see: lapponica Zetterstedt, 1838 .

usambarae Fauvel, 1904d: 291 ( Omalium , cited as Homalium ; subgenus Phloeostiba ; Type locality: Usambara: Derema, 850 m).

— Bernhauer and Schubert, 1910: 58 ( Phloeonomus ; subgenus Phloeostiba ; catalog).

— Bernhauer, 1942c: 349 ( Phloeonomus ; Cameroon).

— Cameron, 1950g: 8 ( Phloeonomus ; subgenus Xylostiba ; Belgian Congo).

— Fagel, 1957a: 27 ( Phloeostiba ; Ruanda).

- DISTRIBUTION: Tanzania, Rwanda, Congo, Cameroon.

Phyllodrepa

[57 species; Neotropical, Nearctic, Palaearctic , Oriental, and Australian Regions] Phyllodrepa Thomson, 1859: 52 (species included: floralis ). TYPE SPECIES: Staphylinus floralis Paykull , fixed by original designation and monotypy.

— Thomson, 1861: 214 (characters).

— Mulsant and Rey, 1880: 230 (characters; key to French species).

— Heyden, 1880: 83 (subgenus of Omalium , cited as Homalium ; list of species of Siberian region).

— Fowler, 1888: 417 (notes; group of Omalium ; key to British species).

— Seidlitz, 1891: 341 (key to some European species).

— Seidlitz, 1891a: 363 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 120 (subgenus of Omalium ; list of species of Europe and Caucasus).

— Heyden, 1893: 49 (subgenus of Omalium ; list of species of Siberian region).

— Ganglbauer, 1895: 738 (characters; key to middle European species).

— Heyden, 1896: 37 (list of species of Siberian region).

— Luze, 1906: 547 (characters; key to species).

— Heyden, Reitter, and Weise, 1906: 130 (list of species of Europe and Caucasus).

— Sainte-Claire Deville, 1907: 28 (key to and annotated list of species of Seine Basin, France).

— Reitter, 1909: 192 (characters; key to German species).

— Bernhauer and Schubert, 1910: 46 (world catalog; 46 species).

— Petri, 1912: 52 (list of species of Siebenbürgen).

— Johansen, 1914: 613 (characters; key to Danish species).

— Leng, 1920: 94 (catalog of North American species).

— Winkler, 1925: 328 (catalog for Palaearctic region).

— Cameron, 1925: 103 (catalog of Indian species).

— Porta, 1926: 20 (characters; key to species of Italy).

— Portevin, 1929: 448 (characters; key to French genera).

— Cameron, 1930: 136 (characters; key to British Indian species).

— Roubal, 1930: 305 (catalog; Slovakia).

— Joy, 1932: 104 (key to British species).

— Scheerpeltz, 1933: 1039 (world catalog supplement).

— Normand, 1935: 357 (list of species of Tunisia).

— Scheerpeltz, 1937b: 187 (list of Bulgarian species).

— Jeannel, 1940: 116 (characters in key; type species: florale Paykull).

— Blackwelder, 1944: 102 (checklist of species from Mexico, Central and South America , and the West Indies).

— Kloet and Hincks, 1945: 160 (list of British species).

— Palm, 1948: 66 (characters; Swedish species).

— Tottenham, 1949: 355, 406 (subgenus of Hapalaraea ; type species: floralis Paykull ; list of British species).

— Blackwelder, 1952: 307 (type species: floralis ).

— Hatch, 1957: 76 (subgenus of Hapalaraea ; characters; key to species of Pacific Northwest).

— Smetana, 1959a: 196 (list of species collected in Albania).

— Horion, 1963: 59 (list of central European species).

— Lohse, 1964: 38 (characters; key to central European species).

— Scheerpeltz, 1968a: 8 (catalog of Austrian species).

— Steel, 1970a: 12 (larval characters; biological notes).

— Tikhomirova, 1973a: 137 (checklist of species of USSR).

— Pope, 1977: 23 (list of British species).

— Topp, 1978: 308 (larval characters).

— Burakowski, Mroczkowski, and Stefańska, 1979: 30 (catalog; subgenus of Hapalaraea ; Poland).

— Muona, 1979: 17 (synonym of Hapalarea).

— Uhlig, 1979: 242 (some species collected in Germany).

— Uhlig and Vogler, 1981: 84 (list of some species collected in Germany; habitat and collecting notes).

— Zanetti, 1983a: 461 (characters; valid genus).

— Segers, 1986: 19 (checklist of species; Belgium).

— Zanetti, 1987: 168 (characters; habitus; key to species of Italy).

— Lohse and Lucht, 1989: 126 (note).

— Angelini, 1991: 192 (list of species from high elevations of Calabria, Italy).

— M. Hansen, Kristensen, Mahler, Pedersen, 1991: 106 (list of Danish species).

— Dettner and Reissenweber, 1991 (defensive secretions: chemistry and systematics).

— Silfverberg, 1992: 21 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Boháč, 1993: 40 (subgenus of Hapalaraea ; list of species of Czech Republic and Slovakia).

— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 27 (additional records to Danish list of species). — M. Hansen, 1996: 94 (list of Danish species).

— M. Hansen, Mahler, Palm, and Pedersen, 1996: 240 (additions to list of Danish species).

— M. Hansen, Palm, Pedersen, and Runge, 1998: 69 (list of species collected in 1997; Denmark).

— Newton, Thayer, Ashe, and Chandler, 2000: 337 (11 Nearctic species; notes; characters in key).

Dropephylla Mulsant and Rey, 1880: 242 (subgenus of Phyllodrepa ; species included: lucida , linearis , heeri , scabriuscula , brevicornis , vilis , gracilicornis ). TYPE SPECIES: Omalium lucida Erichson , fixed by subsequent designation by Blackwelder, 1952: 136.

— Luze, 1906: 547 (synonym of Phyllodrepa ).

— Heyden, Reitter, and Weise, 1906: 130 (subgenus of Phyllodrepa ; list of species of Europe and Caucasus).

— Bernhauer and Schubert, 1910: 47 (subgenus of Phyllodrepa ; catalog).

— Tottenham, 1939d: 229 (type species designation: iopterum ).

— Palm, 1948: 66 (subgenus of Phyllodrepa ; characters; species of Sweden).

— Tottenham, 1949: 355, 406 (subgenus of Hapalaraea ; type species: ioptera Stephens = lucidum Erichson ; list of British species).

— Lohse, 1964: 39 (subgenus of Phyllodrepa ; characters; key to species of central Europe).

— Steel, 1970a: 11 (valid genus; larval characters; biological notes).

— Tikhomirova, 1973a: 138 (subgenus of Phyllodrepa ; checklist of species of USSR).

— Pope, 1977: 23 (list of British species).

— Ádám, 1985: 251 (cited as subgenus of Hapalaraea ; list of species from southeast Hungary).

— Segers, 1986: 19 (cited as valid genus; checklist of species; Belgium).

— Zanetti, 1987: 171, 181 (subgenus of Phyllodrepa ; characters; habitus; key to species of Italy).

— Welch, 1993 (ovariole structure and development).

— Boháč, 1993: 40 (subgenus of Hapalaraea ; list of species of Czech Republic and Slovakia).

— Newton, Thayer, Ashe, and Chandler, 2000: 336 (2 Nearctic species; appears to be distinct group; cited as valid genus; notes; characters in key).

afghanica Coiffait, 1982b: 80 ( Phyllodrepa ; subgenus Phyllodrepa ; Type locality: Afghanistan: Nouristan: Kamdesch, 2200 m).

- DISTRIBUTION: Afghanistan.

akbesiana Pic, 1898 , see: aspera Eppelsheim, 1889 .

almorensis Champion, 1920: 241 ( Omalium ; Type locality: W. Almora in Kumaon).

— Cameron, 1930: 137 ( Phyllodrepa ; subgenus Dropephylla ; characters; India).

— Scheerpeltz, 1933: 1041 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Coiffait, 1984: 118 ( Phyllodrepa ; Pakistan).

- DISTRIBUTION: India, Pakistan.

alutacea Reitter, 1909: 193 ( Phyllodrepa ; subgenus Hapalaraea ; Type locality: Circassien).

— Bernhauer and Schubert, 1910: 49 ( Phyllodrepa ; subgenus Hapalaraea ; catalog).

— Tikhomirova, 1973a: 138 ( Phyllodrepa ; subgenus Hapalaraea ; checklist; Central Asia).

- DISTRIBUTION: Caucasus.

amanni Bernhauer, 1940b: 624 ( Phyllodrepa ; subgenus Dropephylla ; Type locality: Ötztal in Tirol, Ochsengarten, 1557 m).

— Horion, 1963: 67 ( Phyllodrepa ; subgenus Dropephylla ; cited as ammanni ; Austria).

— Lohse, 1964: 40 ( Phyllodrepa ; subgenus Dropephylla ; cited as ammanni ; characters; central Europe).

— Tikhomirova, 1973a: 138 ( Phyllodrepa ; subgenus Drepophylla ; cited as ammanni ; checklist; Tirol).

— Zanetti, 1987: 186 ( Phyllodrepa ; subgenus Dropephylla ; cited as ammanni ; characters; Italy).

— Lucht, 1987: 87 ( Phyllodrepa ; checklist; Austria).

— Zanetti, 1995: 6 ( Phyllodrepa ; cited as ammanni ; Italy).

- DISTRIBUTION: Austria, Italy.

angustata Mäklin, 1878: 28 [= 1878a: 304] ( Omalium , cited as Homalium ; Type locality: Mesenkin).

— J. Sahlberg, 1880: 111 ( Phyllodrepa ; characters; Siberia).

— J. Sahlberg, 1897: 365 ( Phyllodrepa ; Siberia).

— Luze, 1906: 555 ( Phyllodrepa ; characters; Arctic Siberia).

— Poppius, 1909: 4 ( Phyllodrepa ; Lena Valley, Siberia ).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; subgenus Phyllodrepa ; catalog).

— Munster, 1925: 5 ( Phyllodrepa ; notes; Novaya Zemlya).

— Scheerpeltz, 1933: 1040 ( Phyllodrepa ; subgenus Phyllodrepa ; catalog).

— Ryabukhin, 1999: 16 ( Hapalaraea ; subgenus Phyllodrepa ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Russia.

obscuricornis J. Sahlberg, 1897: 365 ( Phyllodrepa ; variety of angustata ; Type locality: Novaja Semlia ad Maly Karmakuli).

— Luze, 1906: 491, 555 ( Phyllodrepa ; aberration of angustata ).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; synonym of angustata ).

— Scheerpeltz, 1933: 1040 ( Phyllodrepa ; variety of angustata ).

obscurata Luze, 1906: 491 , 555 ( Phyllodrepa ; aberration of angustata ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; no locality cited).

apicalis Bernhauer, 1939 , see: Stenomalium .

armena Khnzorian, 1959 , see: Dialycera .

aspera Eppelsheim, 1889: 183 ( Omalium , cited as Homalium ; subgenus Phyllodrepa ; Type locality: Syrien).

— Luze, 1906: 485, 581 ( Dialycera ; characters; Syria).

— Bernhauer and Schubert, 1910: 49 ( Phyllodrepa ; subgenus Dialycera ; catalog).

— Khnzorian, 1959: 68 ( Phyllodrepa ; subgenus Dialycera ; characters).

- DISTRIBUTION: Syria.

akbesiana Pic, 1898: 91 ( Omalium , cited as Homalium ; Type locality: Akbès dans la Haute Syrie).

— Luze, 1906: 485, 581 ( Omalium ; synonym of asperum ).

— Bernhauer and Schubert, 1910: 50 ( Phyllodrepa ; synonym of aspera ).

atra Casey, 1894: 420 ( Omalium ; [preoccupied]; Type locality: California: Santa Cruz Co.).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; subgenus Phyllodrepa ; catalog).

— Herman, 2001: 39 ( Phyllodrepa ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: USA.

atricapilla Bernhauer, 1903b: 191 ( Phyllodrepa , cited as Phylodrepa; Type locality:

Central Altai).

— Luze, 1906: 565 ( Phyllodrepa ; characters; Altai).

— Bernhauer and Schubert, 1910: 47 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Pototskaia, 1967: 21 ( Hapalaraea ; larval characters).

— Tikhomirova, 1973a: 138 ( Phyllodrepa ; subgenus Drepophylla ; checklist; Altai).

- DISTRIBUTION: Altai region.

baicalensis Bernhauer, 1903c: 591 ( Phloeonomus ; Type locality: Baikalsee).

— Luze, 1906: 485, 562 ( Phyllodrepa ; characters; eastern Siberia).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; subgenus Phyllodrepa ; catalog).

— Palm, 1948: 72 ( Phyllodrepa ; characters; Sweden; Norway).

— Tikhomirova, 1973a: 138 ( Phyllodrepa ; subgenus Phyllodrepa ; checklist; northern Europe; eastern Siberia).

- DISTRIBUTION: Norway, Sweden, Russia.

beieri Scheerpeltz, 1958c: 387 ( Phyllodrepa ; subgenus Dropephylla ; Type locality: bei Nisista in den Xerovuni im südlichen Epirus, etwa 800 m Seehöhe).

- DISTRIBUTION: Greece.

bolsonensis Scheerpeltz, 1972: 50 ( Phyllodrepa ; subgenus Phyllodrepa ; Type locality:

bonariensis Bernhauer, 1912d: 29 ( Phyllodrepa ; Type locality: Argentinien: Prov. Buenos Aires).

— Scheerpeltz, 1933: 1040 ( Phyllodrepa ; subgenus Phyllodrepa ; catalog).

- DISTRIBUTION: Argentina.

brevicornis Erichson, 1840 , see: vilis Erichson, 1840 .

brunnea Paykull, 1789 , see: melanocephala Fabricius, 1787 .

cacti Schwarz, 1899: 9 ( Omalium ; subgenus Phyllodrepa ; Type locality: Tucson, Arizona).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; subgenus Phyllodrepa ; catalog).

— Newton, Thayer, Ashe, and Chandler, 2000: 336 ( Dropephylla ).

- DISTRIBUTION: USA.

caseyi Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; subgenus Phyllodrepa ; replacement name for pacifica Casey ).

— Hatch, 1957: 76 ( Hapalaraea subgenus Phyllodrepa ; characters; British Columbia ; Oregon). - DISTRIBUTION: USA.

pacifica Casey, 1894: 421 ( Omalium ; [preoccupied]; Type locality: California: Siskiyou Co.).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; synonym of caseyi ).

caucasica Kolenati, 1846: 28 ( Omalium ; Type locality: Caucasus et trans Caucasus).

— Hochhuth, 1849: 205 ( Omalium ; characters; Caucasus).

— Luze, 1906: 568 ( Phyllodrepa ; characters; Caucasus; Turkestan; Asia Minor).

— Bernhauer and Schubert, 1910: 48 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Pototskaia, 1967: 21 ( Hapalaraea ; larval characters).

— Tikhomirova, 1973a: 138 ( Phyllodrepa ; subgenus Drepophylla ; checklist; Asia Minor; Central Asia).

- DISTRIBUTION: Caucasus, Turkestan, Turkey.

clavigera Luze, 1906: 566 ( Phyllodrepa ; Type locality: Nord-Finland: Kittilä; Mittle-Finland, Jisalmi).

— Bernhauer and Schubert, 1910: 48 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Scheerpeltz, 1933: 1041 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Y. Kangas, 1937 ( Phyllodrepa ; natural history).

— Palm, 1948: 73 ( Phyllodrepa ; subgenus Dropephylla ; characters; Sweden; Finland).

— Silfverberg, 1992: 21 ( Hapalaraea ; Finland; Sweden; Norway).

- DISTRIBUTION: Finland, Sweden, Norway.

crassicornis A. Matthews, 1863, see: salicis Gyllenhal, 1810 .

crassicornis J. Sahlberg, 1876 , see: sahlbergi Luze, 1906 .

cribripennis Fauvel, 1878: 88 [= 1878a: 8] ( Omalium , cited as Homalium ; Type locality: Daya; Bou-Saada; Caucase).

— Fauvel, 1886: 14 [= 1886a: 6] ( Omalium , cited as Homalium ; Algeria).

— Eppelsheim, 1892: 346 ( Omalium ; Turkestan).

— Fauvel, 1902b: 52 ( Omalium , cited as Homalium ; Algeria).

— Luze, 1906: 558 ( Phyllodrepa ; characters; notes; Algeria).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; subgenus Phyllodrepa ; catalog).

- DISTRIBUTION: Algeria, Uzbekistan, Caucasus.

curticollis Eppelsheim, 1889: 182 ( Omalium , cited as Homalium ; subgenus Phyllodrepa ; Type locality: Beyrut, Syrien).

— Luze, 1906: 573 ( Phyllodrepa ; characters; Lebanon).

— Bernhauer and Schubert, 1910: 48 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

- DISTRIBUTION: Lebanon.

devillei Bernhauer, 1902d: 705 ( Phyllodrepa ; cited as Phylodrepa; Type locality: Alpes maritimes, Mont Macaron).

— Luze, 1906: 566 ( Phyllodrepa ; characters; Maritime Alps).

— Luze, 1906: 563 ( Phyllodrepa ; characters; cited as gobanzi ; Sardinia [Note: Misidentification of devillei , according to Zanetti, 1987: 191]).

— Bernhauer and Schubert, 1910: 48 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Zanetti, 1987: 189 ( Phyllodrepa ; subgenus Dropephylla ; characters; Italy).

— Whitehead, 1990b: 190 ( Dropephylla ; collecting notes; Wales).

— Zanetti, 1995: 6 ( Phyllodrepa ; Italy).

— Owen, 1997: 302 ( Dropephylla ; Britain).

— Fowles, Alexander, and Key, 1999: 129 ( Dropephylla ; saproxylic species; status as endangered or threatened species; Britain).

— Owen, 2000: 251 ( Dropephylla ; pitfall trap collections from underground at roots of trees; Britain).

- DISTRIBUTION: Italy, France, Britain.

grandiloqua Luze, 1910 a: 394 ( Phyllodrepa ; Type locality: Schottland, Nordengland ).

— Scheerpeltz, 1933: 1041 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Steel, 1970a: 11 ( Dropephylla ; biological notes).

— Pope, 1977: 23 ( Dropephylla ; Britain).

— Hammond, 1980: 137 ( Dropephylla ; notes; Ireland).

— Zanetti, 1987: 189 ( Phyllodrepa ; synonym of devillei ).

— Luff, 1996: 30 ( Dropephylla ; collecting notes; Isle of Man).

— Owen, 1997: 302 ( Dropephylla ; synonym of devillei ).

propinqua Bernhauer, 1943 a: 172 ( Phyllodrepa ; subgenus Dropephylla ; Type locality: Schottland: Oban).

— Pope, 1977: 23 ( Dropephylla ; synonym of grandiloqua ).

elegans Kraatz, 1857d: 989 ( Omalium ; Type locality: Grafschaft Glatz).

— Redtenbacher, 1857: 994 ( Omalium ; characters).

— Redtenbacher, 1874: 278 ( Omalium ; characters).

— Ganglbauer, 1895: 741 ( Phyllodrepa ; subgenus Dropephylla ; characters; Europe).

— Luze, 1906: 492, 564 ( Phyllodrepa ; synonym of linearis ).

— Bernhauer and Schubert, 1910: 48 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Scheerpeltz, 1933: 1041 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Horion, 1963: 67 ( Phyllodrepa ; species unclear).

— Horion, 1965: 176 ( Phyllodrepa ; discussed as doubtful: Grafschaft Glatz, Galtzer Schneeberg, Biskiden).

— Lohse, 1964: 40 ( Phyllodrepa ; subgenus Dropephylla ; characters; only holotype from Glatz; perhaps a specimen of linearis ; central Europe).

— Lucht, 1987: 87 ( Phyllodrepa ; in checklist).

— Boháč, 1993: 40 ( Hapalaraea ; subgenus Dropephylla ; Czech Republic).

- DISTRIBUTION: Poland, Czech Republic.

flavipennis Cameron, 1930: 138 ( Phyllodrepa ; subgenus Dropephylla ; Type locality:

Dehra Dun).

— Scheerpeltz, 1933: 1041 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Legner and Moore, 1977: 175 ( Hapalaraea ; found under bark).

- DISTRIBUTION: India.

floralis Paykull, 1789: 67 ( Staphylinus ; Type locality: Suecia).

— Fabricius, 1793: 530 ( Staphylinus ; characters).

— Panzer, 1795: 358 ( Staphylinus ; characters; Germany).

— Paykull, 1800: 406 ( Staphylinus ; characters; Sweden).

— Fabricius, 1801: 604 ( Stenus ; characters; France).

— Turton, 1802: 514 ( Staphylinus ; characters; France).

— Gravenhorst, 1802: 118 ( Omalium ; characters).

— Latreille, 1804: 373 ( Omalium ; characters; France; Germany; Sweden).

— Gravenhorst, 1806: 210 ( Omalium ; characters).

— Olivier, 1811: 477 ( Omalium ; characters; France; Germany; Sweden).

— Curtis, 1829: 29 ( Omalium ; catalog; Britain).

— Mannerheim, 1830: 54 ( Anthobium ; Russia).

— Stephens, 1834: 350 ( Omalium ; characters; England).

— Erichson, 1839a: 633 ( Omalium ; characters; Germany).

— Heer, 1839: 177 ( Omalium ; characters; Switzerland).

— Stephens, 1839: 429 ( Omalium ; characters; England).

— Erichson, 1840: 879 ( Omalium ; characters; Europe).

— P. Lucas, 1846: 127 ( Anthobium ; notes; Algeria).

— Hochhuth, 1849: 204 ( Omalium ; characters).

— Redtenbacher, 1849: 752 ( Omalium ; characters; Austria).

— Kiesenwetter, 1851a: 435 ( Omalium ; France).

— Hardy, 1851: 61 ( Omalium ; notes; England).

— Thomson, 1857a: 225 ( Omalium ; characters).

— Fairmaire and Laboulbène, 1856: 642 ( Omalium ; characters; France).

— Redtenbacher, 1857: 252 ( Omalium ; characters; Austria).

— Kraatz, 1857d: 996 ( Omalium ; characters; Germany).

— Thomson, 1859: 52 (type species of Phyllodrepa ).

— Thomson, 1861: 214 ( Phyllodrepa ; characters; Scandinavia).

— Hochhuth, 1862: 108 ( Omalium ; Russia).

— Fauvel, 1871a: 88 [= 1872: 62] ( Omalium , cited as Homalium ; [Note: Misidentification: Bernhauer and Schubert, 1910: 46 cited rufipes Fauvel as a synonym of floralis ; Fauvel did not describe the species as new, he attributed it to Fourcroy. I have assumed that the species cited by Fourcroy referred to the rufipes of Linné a species that is now in Tachinus ]; characters; France; Algeria).

— Redtenbacher, 1874: 275 ( Omalium ; characters; Austria).

— Seidlitz, 1875: 235 ( Omalium ; characters; Baltic region).

— Fauvel, 1878d: 210 [= 1878e: 46] ( Omalium , cited as Homalium ; [Note: Misidentification: Bernhauer and Schubert, 1910: 46 cited rufipes Fauvel as a synonym of floralis ; Fauvel did not describe the species as new, he attributed it to Fourcroy. I have assumed that the species cited by Fourcroy referred to the rufipes of Linné and species that is now in Tachinus ]; characters; Lake Superior; Michigan; Pennsylvania).

— Mulsant and Rey, 1880: 234 ( Phyllodrepa ; characters; France).

— Fowler, 1888: 418 ( Omalium , cited as Homalium ; cited as rufipes Fourcroy ; subgenus Phyllodrepa ; characters; Britain).

— Ganglbauer, 1895: 740 ( Phyllodrepa ; characters; Europe; Algeria; Caucasus; North America ).

— Fauvel, 1902b: 52 ( Omalium , cited as Homalium ; Algeria).

— Luze, 1906: 560 ( Phyllodrepa ; characters; northern and central Europe; Caucasus; Asia Minor; Algeria).

— Sainte-Claire Deville, 1906: 69 ( Omalium , cited as Homalium ; Corsica).

— Reitter, 1909: 192 ( Phyllodrepa ; characters; Germany).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; subgenus Phyllodrepa ; catalog).

— Blatchley, 1910: 480 ( Omalium , cited as Homalium ; characters; Indiana).

— Johansen, 1914: 614 ( Phyllodrepa ; subgenus Phyllodrepa ; characters; Denmark).

— Portevin, 1929: 448 ( Phyllodrepa ; cited as rufipes ; characters; France).

— Portevin, 1929: 452 ( Anthobium ; characters; France).

— Scheerpeltz, 1933: 1040 ( Phyllodrepa ; subgenus Phyllodrepa ; catalog).

— C. Koch, 1940: 373 ( Phyllodrepa ; Spain).

— Palm, 1948: 71 ( Phyllodrepa ; characters; Sweden; Denmark; Norway; Finland).

— Jeannel and Jarrige, 1949: 313 ( Phyllodrepa ; cave species; Algeria).

— Hatch, 1957: 76 ( Hapalaraea subgenus Phyllodrepa ; characters; British Columbia ; Idaho; Washington; Oregon).

— Larsson and Gígja, 1959: 58 ( Phyllodrepa ; collecting notes; Iceland).

— Horion, 1963: 65 ( Phyllodrepa ; subgenus Phyllodrepa ; Algeria; Turkey; Caucasus; Lapland to Mediterranean region; Germany; Austria).

— Lohse, 1964: 38 ( Phyllodrepa ; subgenus Phyllodrepa ; characters; central Europe).

— Smetana, 1964d: 53 ( Phyllodrepa ; subgenus Phyllodrepa ; Czechoslovakia).

— Szujecki, 1968a: 711 ( Hapalaraea ; Poland). — Pope, 1977: 23 ( Phyllodrepa ; Britain).

— Hammond, 1980: 137 ( Phyllodrepa ; Ireland).

— Tóth, 1982: 53 ( Hapalaraea ; characters; Hungary).

— Zanetti, 1987: 180 ( Phyllodrepa ; subgenus Phyllodrepa ; characters; Italy).

— Ádám, 1987: 137 ( Phyllodrepa ; collecting notes; Hungary).

— Lucht, 1987: 86 ( Phyllodrepa ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Dettner and Reissenweber, 1991 ( Phyllodrepa ; defensive secretions: chemistry and systematics). — Boháč, 1993: 40 ( Hapalaraea ; subgenus Phyllodrepa ; Czech Republic; Slovakia).

— Sparacio, 1995: 140 ( Phyllodrepa ; characters; notes; Sicily).

— Zanetti, 1995: 6 ( Phyllodrepa ; Italy).

— Downie and Arnett, 1996: 436 ( Hapalaraea ; characters; USA).

— Zanetti and Sabella, 1998: 28 ( Phyllodrepa ; Sicily).

— Cibuļskis, 1999: 34 ( Hapalaraea ; list; Latvia).

- DISTRIBUTION: Iceland, Europe, Russia, Turkey, Algeria, Caucasus, Canada, USA. viburni Gravenhorst, 1802: 117 ( Omalium ; Type locality: Not cited, but presumably Brunsvigae).

— Latreille, 1804: 373 ( Omalium ; characters; Germany).

— Gravenhorst, 1806: 210 ( Omalium ; characters).

— Gyllenhal, 1810: 224 ( Omalium ; characters; Sweden).

— Olivier, 1811: 477 ( Omalium ; characters; Germany).

— Gyllenhal, 1827: 467 ( Omalium ).

— Mannerheim, 1830: 54 ( Anthobium ; Finland; Sweden; Russia).

— Mannerheim, 1831: 468 ( Anthobium ; Finland; Sweden; Russia).

— Stephens, 1834: 351 ( Omalium ; characters; England).

— C. Sahlberg, 1830: 285 ( Omalium ; characters; Finland).

— Runde, 1835: 23 ( Anthobium ; characters; Germany).

— Lacordaire, 1835: 472 ( Anthobium ; characters; France).

— Erichson, 1839a: 633 ( Omalium ; synonym of floralis ).

— Heer, 1839: 177 ( Omalium ; synonym of florale).

— Stephens, 1839: 429 ( Omalium ; variety of florale; characters; England).

— Erichson, 1840: 879 ( Omalium ; synonym of florale).

— Laporte, 1840: 192 ( Anthobium ; characters; France).

— Hardy, 1851: 61 ( Omalium ; synonym of florale).

— Thomson, 1857a: 225 ( Omalium ; synonym of florale).

— Fairmaire and Laboulbène, 1856: 642 ( Omalium ; synonym of florale).

— Kraatz, 1857d: 996 ( Omalium ; synonym of florale).

— Thomson, 1861: 214 ( Phyllodrepa ; synonym of floralis ).

— Mulsant and Rey, 1880: 235 ( Phyllodrepa ; synonym of floralis ).

— Ganglbauer, 1895: 740 ( Phyllodrepa ; synonym of floralis ).

— Luze, 1906: 560 ( Phyllodrepa ; synonym of floralis ).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; synonym of floralis ).

ruficornis Stephens, 1834: 352 ( Omalium ; Type locality: Suffolk; Norfolk; near Bristol).

— Stephens, 1839: 429 ( Omalium ; characters; England).

— G. Waterhouse, 1858: 32 ( Omalium ; synonym of florale).

— Fauvel, 1871a: 88 [= 1872: 62] ( Omalium , cited as Homalium ; synonym of rufipes ).

— Ganglbauer, 1895: 740 ( Phyllodrepa ; synonym of floralis ).

— Luze, 1906: 560 ( Phyllodrepa ; synonym of floralis ).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; synonym of floralis ).

fahraei Zetterstedt, 1838: 52 ( Omalium ; Type locality: Norvegiae Dowre).

— Erichson, 1840: 888 ( Omalium ; characters; Lapland).

— Thomson, 1857a: 225 ( Omalium ; synonym of florale).

— Thomson, 1861: 214 ( Phyllodrepa ; synonym of floralis ).

— Fauvel, 1871a: 88 [= 1872: 62] ( Omalium , cited as Homalium ; synonym of rufipes ).

— Luze, 1906: 560 ( Phyllodrepa ; synonym of floralis ).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; synonym of floralis ).

maculicornis Heer, 1839: 178 ( Omalium ; Type locality: monts Jura).

— Redtenbacher, 1857: 253 ( Omalium ; characters; Austria).

— Fauvel, 1871a: 88 [= 1872: 62] ( Omalium , cited as Homalium ; synonym of rufipes ).

— Redtenbacher, 1874: 277 ( Omalium ; characters).

— Mulsant and Rey, 1880: 235 ( Phyllodrepa ; synonym of floralis ).

— Ganglbauer, 1895: 740 ( Phyllodrepa ; synonym of floralis ).

— Luze, 1906: 560 ( Phyllodrepa ; synonym of floralis ).

— Bernhauer and Schubert, 1910: 46 ( Phyllodrepa ; synonym of floralis ).

— Scheerpeltz, 1933: 1040 ( Phyllodrepa ; synonym of floralis ).

— Zanetti, 1987: 180 ( Phyllodrepa ; synonym of floralis ).

fahraei Zetterstedt, 1838, see: floralis Paykull, 1789 .

gagliardii C. Koch, 1937 , see: ioptera Stephens, 1832 .

gobanzi Ganglbauer, 1904: 650 ( Phyllodrepa ; subgenus Dropephylla ; Type locality: Insel Meleda, Dalmatia).

— Luze, 1906: 563 ( Phyllodrepa ; characters; Meleda Island; Greece; Sardinia [Note: Misidentification of devillei , according to Zanetti, 1987: 191]).

— Bernhauer and Schubert, 1910: 48 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Zanetti, 1987: 191 ( Phyllodrepa ; subgenus Dropephylla ; characters; possibly in Italy).

- DISTRIBUTION: Croatia, Greece.

gracilicornis Fairmaire and Laboulbène, 1856: 642 ( Omalium ; Type locality: Fontainebleau).

— Kraatz, 1857d: 993 ( Omalium ; characters).

— Fauvel, 1871a: 90 [= 1872: 64] ( Omalium , cited as Homalium ; characters; France; Britain).

— Mulsant and Rey, 1880: 257 ( Phyllodrepa ; subgenus Dropephylla ; characters; France).

— Fowler, 1888: 420 ( Omalium , cited as Homalium ; subgenus Phyllodrepa ; characters; Britain).

— Ganglbauer, 1895: 742 ( Phyllodrepa ; subgenus Dropephylla ; characters; western middle Europe).

— Luze, 1906: 571 ( Phyllodrepa ; characters; western central Europe).

— Reitter, 1909: 193 ( Phyllodrepa ; subgenus Dropephylla ; characters; Germany).

— Bernhauer and Schubert, 1910: 48 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— J. Sahlberg, 1913: 58 ( Phyllodrepa ; Anatolia).

— Johansen, 1914: 617 ( Phyllodrepa ; subgenus Dropephylla ; characters; Denmark).

— Portevin, 1929: 449 ( Phyllodrepa ; characters; France).

— Scheerpeltz, 1933: 1041 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Palm, 1948: 74 ( Phyllodrepa ; subgenus Dropephylla ; characters; Sweden; Denmark).

— Horion, 1963: 68 ( Phyllodrepa ; subgenus Dropephylla ; Britain; Denmark; Sweden; Holland; France; Switzerland; Spain; Poland; Germany; Austria).

— Lohse, 1964: 40 ( Phyllodrepa ; subgenus Dropephylla ; characters; central Europe).

— Legner and Moore, 1977: 175 ( Hapalaraea ; found under bark).

— Pope, 1977: 23 ( Dropephylla ; Britain).

— Hammond, 1980: 137 ( Dropephylla ; notes; Ireland).

— Tóth, 1982: 56 ( Hapalaraea ; characters).

— Zanetti, 1987: 188 ( Phyllodrepa ; subgenus Dropephylla ; characters; Italy).

— Lucht, 1987: 87 ( Phyllodrepa ; checklist; Germany; Austria; Switzerland; Benelux; Denmark).

— Allen, 1988b: 210 ( Dropephylla ; collecting notes; Britain).

— Boháč, 1993: 40 ( Hapalaraea ; subgenus Dropephylla ; Czech Republic).

— Zanetti, 1995: 6 ( Phyllodrepa ; Italy).

— Morgan, 1996: 60 ( Dropephylla ; Britain).

— M. Hansen, 1996: 94 ( Phyllodrepa ).

— Assing, Frisch, Kahlen, et al., 1998: 122 ( Phyllodrepa ; note).

- DISTRIBUTION: Europe, Turkey.

hiemalis Fuss, 1868: 355 ( Omalium ; Type locality: Cleve, Thiergarten).

— Fauvel, 1871a: 93 [= 1872: 67] ( Omalium , cited as Homalium ; characters; France).

— Redtenbacher, 1874: 277 ( Omalium ; characters). — Ganglbauer, 1895: 742 ( Phyllodrepa ; synonym of gracilicornis ). — Luze, 1906: 571 ( Phyllodrepa ; synonym of gracilicornis ). — Bernhauer and Schubert, 1910: 48 ( Phyllodrepa ; synonym of gracilicornis ). — Scheerpeltz, 1933: 1041 ( Phyllodrepa ; synonym of gracilicornis ).

graeca Bernhauer, 1929 g: 178 ( Phyllodrepa ; subgenus Dropephylla ; Type locality: Griechenland: Parnass). — Scheerpeltz, 1933: 1041 ( Phyllodrepa ; subgenus Dropephylla ; catalog). - DISTRIBUTION: Greece.

grandiloqua Luze, 1910 , see: devellei Bernhauer, 1902.

heeri Heer, 1841: 571 ( Phloeonomus ; cited as heerii ; Type locality: Salève, Pittous 4200 ̍). — Fauvel, 1871a: 90 [= 1872: 65] ( Omalium , cited as Homalium ; cited as heeri ; synonym of vile ). — Mulsant and Rey, 1880: 248 ( Phyllodrepa ; subgenus Dropephylla ; cited as heeri ; characters; France). — Ganglbauer, 1895: 742 ( Phyllodrepa ; cited as heeri ; synonym of vilis ). — Luze, 1906: 572 ( Phyllodrepa ; cited as heeri ; synonym of vilis ). — Bernhauer and Schubert, 1910: 49 ( Phyllodrepa ; cited as heeri ; synonym of vilis ). — Scheerpeltz, 1933: 1042 ( Phyllodrepa ; cited as heeri ; synonym of vilis ). — Pope, 1977: 23 ( Dropephylla ; cited as heeri ; cited as valid species; Britain). — Fowles, Alexander, and Key, 1999: 129 ( Dropephylla ; saproxylic species; status as endangered or threatened species; Britain). - DISTRIBUTION: France, Britain.

hiemalis Fuss, 1868 , see: gracilicornis Fairmaire and Laboulbène, 1856 .

himalayica Cameron, 1924 , see: Phloeonomus .

hispanica Bernhauer, 1929 g: 177 ( Phyllodrepa ; subgenus Dropephylla ; Type locality: Nordspanien: Caboalles, Cancas). — Scheerpeltz, 1933: 1041 ( Phyllodrepa ; subgenus Dropephylla ; catalog). - DISTRIBUTION: Spain.

hummleri Bernhauer, 1936 g: 303 ( Phyllodrepa ; Type locality: Bosnien: Hranisava ). - DISTRIBUTION: Bosnia Hercegovina.

ioptera Stephens, 1832: 349 ( Omalium ; Type locality: Hertford; London; New Forest; Devonshire; Suffolk). — Paykull, 1789: 64 ( Staphylinus ; variation β of brunneus is described; cited as synonym of ioptera by Bernhauer and Schubert, 1910: 48; Sweden). — Gravenhorst, 1806: 206 ( Omalium ; [Note: Misidentification: Bernhauer and Schubert, 1910: 48 cited brunnea variety 1 Gravenhorst as a synonym of ioptera ; there is no available name]; characters). — Gyllenhal, 1810: 227 ( Omalium ; [Note: Misidentification: Bernhauer and Schubert, 1910: 48 cited brunnea variety b Gyllenhal as a synonym of ioptera ; there is no available name]; characters; Sweden). — Stephens, 1834: 349 ( Omalium ; characters; England). — Stephens, 1839: 428 ( Omalium ; characters; England). — Hardy, 1851: 58 ( Phloeonomus ; notes; England). — Fauvel, 1871a: 90 [= 1872: 64] ( Omalium , cited as Homalium ; characters; France; Scandinavia; Britain; Germany; Switzerland; Italy; Austria). — Seidlitz, 1875: 235 ( Omalium ; characters; Baltic region). — J. Sahlberg, 1876: 223 ( Phyllodrepa ; Finland). — Fauvel, 1878: 88 [= 1878a: 8] ( Omalium , cited as Homalium ; Algeria). — Fauvel, 1886: 14 [= 1886a: 6] ( Omalium , cited as Homalium ; Algeria). — Fowler, 1888: 420 ( Omalium , cited as Homalium ; subgenus Phyllodrepa ; characters; Britain). — Ganglbauer, 1895: 741 ( Phyllodrepa ; subgenus Dropephylla ; characters; north and middle Europe; Italy). — Luze, 1906: 569 ( Phyllodrepa ; characters; northern and central Europe).

— Reitter, 1909: 193 ( Phyllodrepa ; subgenus Dropephylla ; characters; Germany).

— Bernhauer and Schubert, 1910: 48 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Johansen, 1914: 616 ( Phyllodrepa ; subgenus Dropephylla ; characters; Denmark).

— Portevin, 1929: 449 ( Phyllodrepa ; characters; France).

— Scheerpeltz, 1933: 1041 ( Phyllodrepa ; subgenus Dropephylla ; catalog).

— Roubal, 1933a: 83 ( Phyllodrepa ; characters; notes).

— Tottenham, 1939d: 229 (type species of Dropephylla ).

— Palm, 1948: 73 ( Phyllodrepa ; subgenus Dropephylla ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 68 ( Phyllodrepa ; subgenus Dropephylla ; Britain; Denmark; Norway; Sweden; Finland; Russia; Poland; Czech Republic; Romania; Serbia; Bosnia Hercegovina; Macedonia; Dalmatia; Italy; Switzerland; Belgium; Holland; Germany; Austria).

— Lohse, 1964: 39 ( Phyllodrepa ; subgenus Dropephylla ; characters; central Europe).

— Steel, 1970a: 11 ( Dropephylla ; biological notes).

— Legner and Moore, 1977: 175 ( Hapalaraea ; found under bark).

— Pope, 1977: 23 ( Dropephylla ; Britain).

— Tóth, 1982: 55 ( Hapalaraea ; characters; Hungary).

— Zanetti, 1987: 193 ( Phyllodrepa ; subgenus Dropephylla ; characters; Italy).

— Lucht, 1987: 87 ( Phyllodrepa ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Boháč, 1993: 40 ( Hapalaraea ; subgenus Dropephylla ; Czech Republic; Slovakia).

— Zanetti, 1995: 6 ( Phyllodrepa ; Italy).

— M. Hansen, 1996: 94 ( Phyllodrepa ).

— Cibuļskis, 1999: 34 ( Hapalaraea ; list; Latvia).

— Owen, 1999b: 61 ( Dropephylla ; captured underground at roots of old trees; Britain).

— Fowles, Alexander, and Key, 1999: 129 ( Dropephylla ; saproxylic species; status as endangered or threatened species; Britain).

— Owen, 2000: 251 ( Dropephylla ; pitfall trap collections from underground at roots of trees; Britain).

- DISTRIBUTION: Europe, Algeria.

lucida Erichson, 1839 a: 634 ( Omalium ; Type locality: Mark Brandenburg).

— Heer, 1839: 185 ( Phloeonomus ; characters; Switzerland).

— Erichson, 1840: 881 ( Omalium ; characters; Germany; France; Sweden).

— Redtenbacher, 1849: 751 ( Omalium ; characters; Austria).

— Thomson, 1857a: 226 ( Omalium ; characters).

— Fairmaire and Laboulbène, 1856: 643 ( Omalium ; characters; France).

— Redtenbacher, 1857: 250 ( Omalium ; characters; Austria).

— Kraatz, 1857d: 995 ( Omalium ; characters; Germany).

— Thomson, 1861: 215 ( Phyllodrepa ; characters; Scandinavia).

— Fauvel, 1871a: 90 [= 1872: 64] ( Omalium , cited as Homalium ; synonym of iopterum ).

— Redtenbacher, 1874: 274 ( Omalium ; characters; Austria).

— Mulsant and Rey, 1880: 242 ( Phyllodrepa ; subgenus Dropephylla ; characters; France).

— Ganglbauer, 1895: 741 ( Phyllodrepa ; synonym of ioptera ).

— Bernhauer and Schubert, 1910: 48 ( Phyllodrepa ; synonym of ioptera ).

— Scheerpeltz, 1933: 1041 ( Phyllodrepa ; synonym of ioptera ).

— Zanetti, 1987: 192 ( Phyllodrepa ; synonym of ioptera ).

medioglabra Roubal, 1933 a: 85 ( Phyllodrepa ; form of ioptera ; Type locality: Slovakia: Levice, B. Bystrica, Gemer).

gagliardii C. Koch, 1937: 84 ( Phyllodrepa ; subgenus Dropephylla ; subspecies of ioptera ; Type locality: Assergi, Gran Sasso; Mte. Giovi, Firenze).

— Zanetti, 1986: 95 ( Phyllodrepa ; synonym of ioptera ).

— Zanetti, 1987: 193 ( Phyllodrepa ; synonym of ioptera ).