Cordilura albipes ( Fallén, 1819 )

Ozerov, A. L. & Krivosheina, M. G., 2020, A review of the genus Cordilura Fallén, 1810 (Diptera: Scathophagidae) of Russia, Russian Entomological Journal 29 (4), pp. 439-480 : 449-451

Cordilura albipes ( Fallén, 1819)

Figs 12, 21–23 , 33–35 .

albipes Fallén, 1819: 9 ( Cordylura ). Type-locality: “in Quasa Esperöd et in silva Gyllebo Scaniae” (Scåne, Sweden).

bilineata Meigen, 1838: 340 ( Cordylura ). Type-locality: not given (“Hiesige Gegend”,?nr Aachen).

unicolor Loew, 1864: 17 ( Cordylura ). Type-locality: “ Andalusien ” ( Spain).

REMARK. This species was recorded in Russia from European part and from Siberia without indicating specific locality by Gorodkov [1970: 446, 1986: 16]; noted from Karelia [ Humala, Polevoi, 2009: 71], Far East [ Ozerov, 2009: 379; Ozerov, Krivosheina, 2014: 206] and Yakutia [ Bagachanova et al., 2016: 779].

MATERIAL EXAMINED. Adygea: Lagonaki (44.093ºN 40.019ºE), 1725 m, 26–28. VI .2009, K. Tomkovich (5 ♂♂, ZMUM); Dakhovskaya env. (44.201ºN 40.149ºE) , 12–13.VII.2012, K. Tomkovich (1 ♂, ZMUM); Altai: Bol’shoy Yaloman (50.489ºN 86.416ºE) , 7–9.VII.1967, R . Kamenskaya (2 ♂♂, 4 ♀♀, ZMUM); Seminsky Pass , (51.06ºN 85.59ºE) , 1650 m, 27–30. VI .2016, N. Vikhrev (1 ♂, ZMUM); Arkhangelsk Oblast: Vas’kovo (64.413ºN 40.457ºE) , 18.VII.1996, Gorodkov (1 ♂, ZISP); Buryatia: Barguzinsky Nature Reserve, Sosnovka env. (ca. 54.182ºN 109.539ºE) , 4.VIII.1962, Gorodkov (1 ♂, ZISP); Chelyabinsk Oblast: Taganay (55.277ºN 59.795ºE) , 18–24.VII.2008, K. Tomkovich (4 ♂♂, 1 ♀, ZMUM); Crimea: Alushta-Rybachie (44.7–8ºN 34.4–6ºE) , 18–25.IV.2014, N. Vikhrev (2 ♂♂, 1 ♀, ZMUM); Dagestan: 15 km SW of Sergocala (ca. 42.462ºN 47.662ºE) , 15.VII.1983, Nartshuk (1 ♂, ZISP); Irkutsk Oblast: Baykal Port (51.874ºN 104.792ºE) , 25. VI .1965 , Negrobov (1 ♀, ZISP); Listvyanka (51.863ºN 104.866ºE), 21. VI .1965 , Negrobov (1 ♀, ZISP); Karelia: Primorskiy (66.546ºN 33.103ºE), 1–6.VII.2010, A.L. Ozerov (1 ♂, 10 ♀♀, ZMUM); Poyakonda (66.589ºN 33.821ºE) , 7.VII.2010, A.L. Ozerov (1 ♂, ZMUM); Kivach Nature Reserve (ca. 62.295ºN 33.921ºE) , 21. VI .1979 , Gorodkov (3 ♂♂, 2 ♀♀, ZISP); Kemerovo Oblast: Tisul’ (55.759ºN 88.310ºE), 21. VI .1911 , Gorchakovskiy (2 ♂♂, ZISP); Krasnodar Krai: Azish-Tau ridge, “Kamyshanova Polyana” (44.168ºN, 40.044ºE), 1240 m, 10.IV.–3. V .2010, Kustov (1 ♂, 2 ♀♀, ZMUM); Sochi, Mt. Akhun (43.523ºN 39.879ºE) , 25.IV.2008, N. Vikhrev (1 ♂, ZMUM); Sochi, Estosadok env., Mt. Psekhako (43.691ºN 40.366ºE) , 14–18. VI .2008 , K. Tomkovich (2 ♂♂, 2 ♀♀, ZMUM); Dakhovskaya env. (44.201ºN 40.149ºE), 12–13.VII.2012, K. Tomkovich (1 ♂, ZMUM); Lagonaki (44.093ºN 40.019ºE) , 1725 m, 26–28. VI .2009, K. Tomkovich (5 ♂♂, ZMUM); Teberdinsky Nature Reserve (ca. 43.443ºN 41.738ºE) , 15 and 20. V .1964 , Gorodkov (3 ♂♂, 3 ♀♀, ZISP); Teberdinsky Nature Reserve (ca. 43.405ºN 41.721ºE), 3–19. V .1964 , Gorodkov (2 ♂♂, 2 ♀♀, ZISP); Krasnoyarsk Krai: Kryuchkovo station (56.096ºN 92.109ºE), 14–23.VII.2009, K. Tomkovich (5 ♂♂, 5 ♀♀, ZMUM); Ergaki Nature Park (52.84ºN 93.25ºE) , 1450 m, 27– 29. VI .2017, N. Vikhrev (3 ♂♂, ZMUM); Leningrad Oblast: Luga (ca. 58.73ºN 29.84ºE) , 21. VI . and 8.VIII.1955 , A. Stackelberg (2 ♂♂, ZISP); Rozhdestveno (59.322ºN 29.946ºE), 15. VI .1957 , A. Stackelberg (2 ♀♀, ZISP); Sablino (ca. 59.63ºN 30.76ºE), 16. VI .1923 , A. Stackelberg (1 ♂, ZISP); Mordovia: Mordovsky Nature Reserve, Cordon Novin’kovsky (54.931ºN, 43.421ºE), 4–7.VII.2020, K. Tomkovich (1 ♂, ZMUM); Moscow and Moscow Oblast: Bittsa (55.641ºN 37.570ºE), 24. VI ., 26– 31.VII.1936, B. Rohdendorf (1 ♂, 7 ♀♀, ZMUM); same place , 29. VI .1936 , E. Smirnov (2 ♂♂, 2 ♀♀, ZMUM); Izmaylovo (55.800ºN 37.832ºE, 55.802ºN 37.840ºE), 5.IX.1981, 26– 30. V. and 5. VI.1982, 17.IV, 26. V. and 2.VII.1983, 18. V .2008 , A.L. Ozerov (9 ♂♂, 5 ♀♀, ZMUM); Kuntsevo (55.720ºN 37.473ºE), 17, 19 and 26. V., 18. VI. and 2.VII.1983, 13.V.1992, 16.VI.1993, A.L. Ozerov (4 ♂♂, 2 ♀♀, ZMUM); Tsaritsyno (55.616ºN 37.683ºE) , 5–10.X.2009, K. Tomkovich (1 ♂, ZMUM); 20 km WSW of Volokolamsk (55.983ºN 35.616ºE) , 13.VII.1999, A.L. Ozerov (1 ♂, ZMUM); 35 km NNE of Moscow (56.200ºN 37.833ºE) , 26. VI .1993 , A.L. Ozerov (2 ♂♂, 1 ♀, ZMUM); Abramtsevo (56.230ºN 37.956ºE), 13.VIII.1955, E.S. Smirnov (1 ♀, ZMUM); Andreevskoe (55.974ºN 35.604ºE) , 4.VIII.2005, A.L. Ozerov (2 ♀♀, ZMUM); Burtsevo env. (55.981ºN 35.597ºE, 55.975ºN 35.589ºE, 55.978ºN 35.589ºE, 55.975ºN 35.586ºE), 25. V ., 2– 3 and 18. VI.2006, 25.VII. and 1.VIII.2007, 29. V .2010 , A.L. Ozerov (4 ♂♂, 4 ♀♀, ZMUM); Vostochnyy (55.366ºN 36.498ºE), 20. V .2010 , D. Gavryushin (2 ♂♂, ZMUM); Golitsyno (55.649ºN 37.011ºE), 23.V.1976, 21.V.1977, 9.IX.1979, 7.VI.1980, 29. V . and 5. VI.1981, 7.VIII.1983, 3.VIII.1989, A. Shatalkin (10 ♂♂, 1 ♀, ZMUM); Dmitrov env. (56.316ºN 37.725ºE), 10. VI . and 29.VII.2006, 19. V .2007 , N. Vikhrev (4 ♂♂, ZMUM); Zosimova Pustyn’ (55.412ºN 36.814ºE), 24 and 28. V .2010 , D. Gavryushin (1 ♂, 1 ♀, ZMUM); Ivanovskoe (55.939ºN 35.621ºE), 12. VI .2006 , A.L. Ozerov (1 ♂, ZMUM); Ignatkovo (55.939ºN 35.620ºE), 13. VI .2008 , A.L. Ozerov (1 ♂ ZMUM); Mamontovka (55.999ºN 37.816ºE), 3 and 16.VII.1956, E. Smirnov (1 ♂, ZMUM); Molzhaninovka (55.936ºN 37.385ºE) , 13. V .2010 , A.L. Ozerov (1 ♂, ZMUM); Naro-Fominsk env. (55.357ºN 36.736ºE, 55.364ºN 36.738ºE, 55.364ºN 36.741ºE, 55.393ºN 36.779ºE, 55.375ºN 36.756ºE, 55.372ºN 36.757ºE), 4 and 16. VI.2006, 22. V., 3 and 7.VI.2007, 13.V.2008, 8. VI . and 2.VIII.2010, 18. V .2012 , D. Gavryushin (5 ♂♂, 5 ♀♀, ZMUM); Ozhigovo (55.453ºN 36.884ºE), 26.VII.2010, D. Gavryushin (1 ♀, ZMUM); Rodinka (55.774ºN 36.291ºE) , 5. VI .2012 , D. Gavryushin (1 ♂, ZMUM); Stepan’kovo (55.004ºN 35.624ºE), 3. VI .2007 , A.L. Ozerov (2 ♂♂, ZMUM); Murmansk Oblast: Kolvitsa (67.082ºN 33.183ºE), 8.VIII.1995, Gorodkov (2 ♀♀, ZISP); Nizhegorod Oblast: Dzerzhinsk (56.21ºN 43.62ºE) , 17.VIII.2009, N. Vikhrev (1 ♀, ZMUM); North Ossetia – Alania: Bakhty-Laparyrag range (42.938ºN 44.287ºE) , 1770 m, 1–14.VI.1989 and 28–30.VI.1990, A.L. Ozerov, A. Shatalkin (4 ♂♂, 6 ♀♀, ZMUM); Buron env. (43.793ºN 43.922ºE) , 20– 29.VI.1988, 6–7.VI.1989, 28. VI . and 5–6.VII.1990, A.L. Ozerov, A. Shatalkin (6 ♂♂, 7 ♀♀, ZMUM); Alagir env. (43.015ºN 44.224ºE) , 17.VII.1990, A.L. Ozerov (1 ♂, ZMUM); Vladikavkaz (43.044ºN 44.682ºE) , 16. IV.1899, Demokidov (1 ♀, ZISP); Novgorod Oblast: Ramen’e (58.374ºN 33.430ºE), Okulovka env ., 4. VI .1988 , Gorodkov (1 ♂, ZISP); Novosibirsk Oblast: Novosibirsk (54.842ºN 83.114ºE), 14. VI .2008 , O. Kosterin (1 ♂, ZMUM); 4 km S of Novososedovo (54.616ºN 83.985ºE), 7.VII.2008, O. Kosterin (1 ♂, ZMUM); (55.52ºN 83.24ºE) , 22. V .2011 , O. Kosterin (3 ♂♂, ZMUM); Primorsky Krai: Kedrovaya Pad’ Nature Reserve (43.104ºN 131.512ºE), 8–16. VI .2007 , A. Ovchinnikov (1 ♀, ZISP); Rostov Oblast: Kamensk-Shakhtinsky (48.293ºN 40.257ºE), 25. V .2011 , D. Gavryushin (2 ♀♀, ZMUM); Ryazan’ Oblast: Kasimov (54.94ºN 41.34ºE), 25.VII.2013, N. Vikhrev (1 ♂, ZMUM); Sakhalin Oblast: Sakhalin I., Mt. Chekhov (47.005ºN 142.839ºE) , 29. VI .1973 , Kasparyan (1 ♀, ZISP); Tatarstan: Volzhsko-Kamsky Nature Reserve, Lake Raifa (55.897ºN 48.733ºE), 19.VII.2007, Basov (1 ♂, ZMUM); Tomsk Oblast: Basandayka (56.283ºN 85.489ºE) , 23. VI .1967 , Gorodkov (5 ♂♂, ZISP); Tyumen’ Oblast: (63.766ºN 59.716ºE), 1–4.VII.2010, K. Tomkovich (2 ♂♂, ZMUM); (63.818ºN 59.562ºE) , 1–4.VII.2010, K. Tomkovich (3 ♂♂, 1 ♀, ZMUM); River Soromlorop’yavin (62.650ºN 71.891ºE) , 26.VII.2010, K. Tomkovich (1 ♀, ZMUM); Sob’ env. (67.07ºN 65.46ºE) , 26–31.VII.2011, K. Tomkovich (1 ♂, ZMUM); Surgut (61.259ºN 73.418ºE) , 27.VII.1977, Gorodkov (3 ♀♀, ZISP); Tuva: Kyzyl env. (51.7ºN 94.7ºE) , 5–7.VII.2017, N. Vikhrev (1 ♂, ZMUM); Zabaykalsky Krai: Baley (51.579ºN 116.634ºE) , 18.VII.1971, V . Richter (1 ♂, ZISP) .

ADDITIONAL MATERIAL EXAMINED. Armenia: Bashkend [now = Artsvashen] (40.641ºN 45.523ºE), 24.VII.1976, Nartshuk (1 ♂, ZISP); Belarus: Belovezhskaya Pushcha (52.707ºN 23.862ºE), 14 and 17.VII ., 5.VIII.1961, Nartshuk (1 ♂, 2 ♀♀, ZISP); Korolevo (55.303ºN 30.212ºE) , 27. VI .1905 , Birulya (1 ♂, ZISP); Georgia : Tsagveri (41.800ºN 43.481ºE), 1200 m, 8 and 12. V .1969 , Gorodkov (7 ♂♂, 2 ♀♀, ZISP); Borzhomi park (41.891ºN 43.321ºE), 880–900 m, 16. V .1969 , Gorodkov (1 ♂, ZISP); Mongolia: Ulan- Bator env. (ca. 47.907ºN 106.931ºE), 24–29. VI .1970 , Nartshuk (2 ♂♂, ZISP).

DIAGNOSTIC DESCRIPTION. Male. Female. Body-length 3.8–6.2 mm. Head. Frontal vitta from yellow to brownish; fronto-orbital plate yellowish. Face, parafacial and gena yellowish, with weak whitish re-

39 40

41

flection. Postcranium yellow completely or yellow in lower part and black in upper half or third. 2 orbital and 2–4 frontal setae present. Antenna pale yellow. Postpedicel approximately 2 times as long as wide. Arista plumose, longest hairs approximately as long as width of postpedicel. Palpus yellow. Thorax yellow in ground colour; scutum varies in colour from shiny black completely to almost yellow completely, usually yellow with two blackish stripes along dorsocentral lines. Acrostichals setulose in two rows, dorsocentrals 3+3 (including scapular seta), intra-alars (0–1)+(0–2), supra-alars 1+2, 2 postpronotals. Scutellum with a pair of strong discal setae only; apical scutellar setae as setulae or absent ( Fig. 12). Legs shining, usually yellow completely, sometimes mid and hind femora slightly darkened. Wing tinged with brownish; veins blackish. Vein R 1 setulose on apical third of dorsal surface. Abdomen from yellow to black, shining or subshining. Male sternite 4 rectangular, longer than wide. Male sternite 5, epandrium and surstyli as in Figs 33–35 . Female ovipositor as in Figs 21–23 .

DISTRIBUTION. Russia: Adygea, Altai, Arkhangelsk Oblast, Buryatia, Chelyabinsk Oblast, Crimea, Dagestan, Irkutsk Oblast, Karelia, Kemerovo Oblast, Krasnodar Krai, Krasnoyarsk Krai, Leningrad Oblast, Mordovia, Moscow Oblast, Murmansk Oblast, Nizhegorod Oblast, North Ossetia – Alania, Novgorod Oblast, Novosibirsk Oblast, Primorsky Krai, Rostov Oblast, Ryazan’ Oblast, Sakhalin Oblast, Tatarstan, Tomsk Oblast, Tyumen’ Oblast, Tuva, Yakutia, Zabaykalsky Krai. — Europe, including Belarus (first record), Armenia (first record), Georgia (first record), Iran [ Khaghaninia, Gharajedaghi, 2014: 391], Mongolia, Japan [ Iwasa, 2020: 472].

Figures

Figs 21–23. Cordilura albipes (Fallén): 21 — end of female abdomen, dorsal view; 22 — same, lateral view; 23 — same, ventral view. Рис 21–23. Cordilura albipes (Fallén): 21 — конец брюШка самки, сверху; 22 — то же, сбоку; 23 — то же, сниЗу.

Figs 33–41. Cordilura albipes (Fallén) (33–35), C. amurensis Ozerov (36–38) and C. atrata (Zetterstedt) (39–41): 33, 36, 39 — male sternite 5; 34, 37, 40 — epandrium and surstyli, lateral view; 35, 38, 41 — same, dorsal view. Рис. 33–41. Cordilura albipes (Fallén) (33–35), C. amurensis Ozerov (36–38) и C. atrata (Zetterstedt) (39–41): 33, 36, 39 — стернит 5 самца; 34, 37, 40 — Эпандрий и сурстили, сбоку; 35, 38, 41 — то же, сверху.