Grishin, Zhang & Cong & Shen & Song & Grishin, 2023

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Genomics-based taxonomic rearrangement of Achlyodini and Carcharodini (Lepidoptera: Hesperiidae: Pyrginae), Insecta Mundi 2023 (16), pp. 1-33 : 22

Macarius Grishin , new subgenus

http://zoobank.org/ 3C0E9973-A2BA-4F62-8B65-C9DA18692468

Type species. Pellicia macarius Herrich-Schäffer, 1870 .

Definition. The genus Nisoniades Hübner, 1819 (type species Papilio bromius Stoll, 1787 , a junior subjective synonym of Papilio mimas Cramer, 1775 ) splits into three prominent clades at the subgenus level ( Fig. 4 ). Because no available genus-group names have been associated with Nisoniades , two of three subgenera are new. This new subgenus keys to E.19.3, E19.4, or E.19.12a in Evans (1953) and is distinguished from its relatives by the following characters: vein RS on ventral hindwing in males usually runs parallel to vein M 1 from its origin to the end of the swollen portion, if divergent then right harpe is bifid or vein RS origin is closer to the end of cell than to the wing base and dorsal hindwing tuft in males is inconspicuous, harpe of right valva is frequently bifid, hindwing is frequently quadrate and angled at vein CuA 1, harper of left valva frequently does not extend past ampulla. In DNA, a combination of the following nuclear genomic base pairs is diagnostic: aly6517.5.2:T1476C, aly6517.5.2:G1699T, aly4975.2.1:C43T, aly4975.2.1:T73G, aly23605.23.23:A145G.

Etymology. The name is a masculine noun in the nominative singular, tautonymous with the type species name.

Species included. Pellicia laurentina R. Williams and E. Bell, 1939 , Nisoniades evansi Steinhauser, 1989 , Nisoniades supra Steinhauser, 1989 , Pellicia maura Mabille and Boullet, 1917 , Pellicia hora Hayward, 1939 , Pellicia criton Mabille, 1898 , Pellicia hesperia Hayward, 1939 , Arteurotia castolus Hewitson, 1878 , Pellicia haywardi R. Williams and E. Bell, 1939 , Pellicia rimana E. Bell, 1942 , Pellicia macarius Herrich-Schäffer, 1870 , and Achlyodes nyctineme A. Butler, 1877 .

Parent taxon. Genus Nisoniades Hübner, [1819] .

Figures

Figure 4. The nuclear genome-based phylogeny of Carcharodini, first segment (continues in Fig. 5, 6). See Fig.1 for notations.