Centetodon pulcher Marsh, 1872b

Murphey, Paul C. & Kelly, Thomas S., 2017, Mammals from the earliest Uintan (middle Eocene) Turtle Bluff Member, Bridger Formation, southwestern Wyoming, USA, Part 2: Apatotheria, Lipotyphla, Carnivoramorpha, Condylartha, Dinocerata, Perissodactyla and Artiodactyla, Palaeontologia Electronica (Cambridge, England: 2003) 20 (2), pp. 1-51 : 14

Centetodon pulcher Marsh, 1872b

Figure 6.5-10 , Table 2

Referred specimens. From DMNH Locality 4672: Rm2, DMNH 75327. From UCM Locality 92189: Lp4, UCM 68534; partial dentary with Lp4-m1,

UCM 95767; partial dentary with Lp1 and p4-m3, UCM 68963; Rp4, UCM 67889; Rm1 or 2, UCM 95768; Rm3s, UCM 69968, 95682.

Description. The p4-m3 of Centetodon pulcher have been well described in the literature ( McKenna, 1960b; McKenna et al., 1962; West, 1973a; Lillegraven et al., 1981) and the new specimens from the TBM do not provide any additional information, so detailed descriptions of them are not necessary. However, to the best of our knowledge, the p1 of C. pulcher has not been previously described. The p1 is moderately worn. It has two roots that are about equal in size and inclined anteriorly. The crown is anteroposteriorly elongated, transversely compressed, and procumbent with the apex of the primary cusp positioned anteriorly. A short ridge is present that extends anteriorly from the primary cusp to terminate as what appears to be an incipient cuspid, but this may just be an artifact of wear. A long, wedge-shaped, slightly convex ridge extends posteriorly from the apex of the primary cusp to terminate at a small posterior accessory cuspid. A labial cingulid is lacking.

Remarks. The eight Centetodon specimens from the TBM are significantly larger than those referred above to C. bemicophagus ( Table 2). In size and occlusal morphology, they are indistinguishable from those of C. pulcher ( Lillegraven et al., 1981) and are referred to the species.

Figures

FIGURE 6. Geolabididae from TBM. 1-4, Centetodon bembicophagus: LP4 (UCM 68681) in occlusal view (1); partial Lm1 or 2 (UCM 68899) in occlusal view (2); Lm1 or 2 (DMNH 75256) in occlusal view (3); Lm3 (DMNH 75257) in occlusal view (4). 5-10, Centetodon pulcher: partial dentary with Lp4-m1 (UCM 95767) in occlusal (5) and labial (6) views; Rm1 or 2 (UCM 95768) in occlusal view (7); partial dentary with Lp1 and p4-m3 (UCM 68963) in occlusal (8), labial (9), and lingual (10) views. Scale bar equals 1 mm.

FIGURE 7. Nyctitherium gunnelli new species from TBM, associated Lp4-m2 (SDSNH 110393, holotype): Lp4 in occlusal (1), labial (4), and lingual (7) views; Lm1 in occlusal (2), labial (5), and lingual (8) views; Lm2 in occlusal (3), labial (6), and lingual (9) views. Scale bar equals 1 mm.

FIGURE 8. Nyctitheriidae from TBM. 1-6, Nyctitherium velox: RM2 (DMNH 75292) in occlusal view (1); RM2 (UCM 95771) in occlusal view (2); RM2 (UCM 68581) in occlusal view (3); RM3 (DMNH 75291) in occlusal view (4); partial Lm2 (DMNH 75317) in occlusal view (5); partial Lm2 (SDSNH 110409) in occlusal view (6). (7), Pontifactor bestiola, RM1 (UCM 71129) in occlusal view. 8-10, Nyctitherium gunnelli new species, LP4 (UCM 95769) in occlusal (8), labial (9), and lingual (10) views. Scale bar equals 1 mm.

FIGURE 9. Chart comparing m1-2 anteroposterior length (ap) versus m1-2 talonid width (trp) in mm of specimens of Nyctitherium velox (including two referred specimens from TBM, SDSNH 110409 and UCM 68967) and holotype m1- 2 (SDSNH 110393) of Nyctitherium gunnelli new species. Black solid filled squares/circles are m1s and white filled squares/circles are m2s. Except for SDSNH 110409 and UCM 68967, measurements of other specimens of N. velox from Robinson (1968a) and Krishtalka (1976b).

FIGURE 10. Oligoryctidae and unnamed lipotyphlan from TBM. 1-4, unnamed apternodontid sp.: partial LM1 or 2 (SDSNH 110348) in occlusal (1) and anterior (2) views; partial Rm1 or 2 (SDSNH 110349) in occlusal (3) and labial (4) views. 5-6, unnamed lipotyphlan sp., partial dentary with Rp4 (DMNH 75258) in occlusal (5) and labial (6) views. Scale bars equals 1 mm.