Clivina excisa, Balkenohl, 2021

Balkenohl, Michael, 2021, The Clivina komareki-species group from Asia with the description of a new species from the Philippines (Coleoptera, Carabidae, Scaritinae), Linzer biologische Beiträge 53 (2), pp. 535-543 : 539-541

Clivina excisa nov.sp. ( Figs 2 , 4)

T y p e m a t e r i a l: Holotype: ♁, with labels and data: white, printed in black, "Kapatagan, Davao de Sur, Mindanao JANUARY 2020 " / "local coll. lgt." (CBB). Paratype: 1♁, with labels and data: white, printed in green, "Kidapawan, South Cotabato, Mindanao MARCH 2020 " / "local coll. lgt." (OLML).

E t y m o l o g y: The name refers to the excised anterior margin of the clypeus, the labrum, and the pronotum which is expressed by the Latin adjective excisa in the feminine genus.

D i a g n o s i s: The new species differs from C. komareki KULT mainly in the head with the gently excised clypeus and the emarginated labrum, the missing triangle-like depression at middle of the frons, and the distinctly moniliform antennomeres; in the pronotum which is distinctly wider than long, and the anterior margin is distinctly excised. In addition, the lateral channel of the pronotum is distinctly wider between the lateral setigerous punctures. The elytra are shorter oval and the humerus exhibits a humeral tooth.

D e s c r i p t i o n: Measurements: Holotype / paratype: Body length 5.97 / 5.57 mm, width 2 / 1.84 mm, ratio L/W of pronotum 0.75 / 0.76, ratio L/W of elytra 1.64 / 1.63.

Colour: Glossy. Piceous. Legs uniformly fuscous, antenna and mandible slightly paler fuscous.

Head: A third narrower than pronotum. Clypeus gently excised anteriorly; wing moderately developed, not more protruding anteriorly than clypeus, fused with clypeus; supraantennal plate wide, long-convex in dorsal and moderately convex in frontal view; clypeus and wing indistinctly margined, supraantennal plate distinctly reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by right-angled notch, separated from supraorbital carina by a transverse sulcus. Clypeus transverse, flattened, separated from frons by a moderately sharp furrow. Frons convex. Clypeus with longitudinal irregular carinae, frons with few scattered microscopic punctures. Supraantennal plate separated from clypeus and frons by deep moderately wide furrow continuing posteriorly up to mid-eye level as wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eyes. With two supraorbital setigerous punctures situated over posterior third of eye. Neck constriction not developed. Eye hemispherical, projecting laterally. Gena small, enclosing completely eye posteriorly. Antenna not reaching posterior seta of pronotum, antennomeres four to ten distinctly moniliform (ratio L/W around 1.02). Labrum moderately excised, isodiametrically to irregularly reticulated, six setose. Mandible wide. Mentum with lateral lobe long oval anteriorly, with two indistinct longitudinal carinae, median tooth much shorter than lobe, distinctly keeled.

Pronotum: Disk slightly convex in lateral view, moderately convex in frontal view. Quadrate, distinctly wider than long. Anterior margin distinctly excised. Reflexed lateral margin smooth, straight to slightly concave at middle, slightly attenuating in anterior half, widest in posterior third; anterior angle distinct; posterior angle marked by distinct tooth, laterally not projecting. Lateral channel indistinctly foveolate, distinctly wider between lateral setigerous punctures. Median line narrow, line-like, joining anterior transverse line and basal constriction; anterior transverse line as deep as median line, not reaching lateral channel. Surface with microscopic punctures, with small group of few moderately sized punctures laterally in basal half, with few transverse wrinkles laterally and at base.

Elytron: Disk slightly convex in lateral view, distinctly convex on frontal view. Elytra concavely impressed in apical quarter (best visible in anterior-frontal view at an angle of 45°). Outline long oval, distinctly less than twice as long as wide, with maximum width at middle. Humerus broadly rounded off, with small humeral tooth pointing backwards. Reflexed lateral margin smooth. Lateral channel moderately wide. Scutellar striole short, indistinct; with distinct setigerous puncture at base of first stria, with blunt tubercle at base of third interval. Striae moderately deep, with distinct punctures, one to four free at base, five to seven joining at humerus, one running up to apex, two ending free at apex, three and four, and five and six joining apically. Intervals moderately convex, third to fifth indistinctly more elevated at base, seventh more elevated apically. Third interval with two setigerous punctures (but see variation). Surface of intervals smooth, glossy.

Hind wing: Distinctly reduced, around half as long as elytron.

Lower surface: Proepisternum covered with distinct punctures. All terminal sternites of abdomen laterally with moderately sized punctures. The two setigerous punctures on each side of the abdominal sternum VII are very widely separated.

Legs: Legs nearly smooth. Protibia with three spines of moderate length and a small but distinct one basally, sulcate dorsally, movable spur regularly arcuate. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with four fine seta at the inner side.

Male genitalia (Fig. 4): Median lobe broadened apically, with longitudinal basal opening, nearly straight, apex with wide and distinct arcuate lamella. Endophallus with longitudinal group of short slender spines. Parameres slender, nearly straight, acuminated in apical half, at apex with one long and one minute seta.

Female genitalia: Unknown. Variation: On the clypeus, the two specimens differ in form and number of the longitudinal irregular carinae. In the holotype, the third interval of the elytron shows one setigerous puncture at the right side and two at the left side. In the paratype, there is no puncture at the right side.

D i s t r i b u t i o n: Known from two localities on the Island of Mindanao.

Figures

Figs 1-2: Clivina species, habitus,dorsal view; (1)C. komareki KULT, holotype; (2) C. excisa nov.sp., holotype.