Amblypsilopus andasibensis, Grichanov, 2021

Grichanov, Igor Ya., 2021, Eleven new species of Amblypsilopus Bigot (Diptera: Dolichopodidae: Sciapodinae) and a key to the species of Madagascar and adjacent islands, European Journal of Taxonomy 755 (1), pp. 47-87 : 70-73

Amblypsilopus andasibensis sp. nov.

urn:lsid:zoobank.org:act:EF68C2DC-2411-4736-B379-563FB79470CB

Fig. 8

Diagnosis

Amblypsilopus andasibensis sp. nov. is very close to A. olgae sp. nov. in habitus. The latter differs from A. andasibensis sp. nov. in the fore tarsomeres 4–5 white. The fore tarsomere 4 is entirely black in A. andasibensis sp. nov.; the long lobe of the cercus has 2 long setae at the middle, but no modified setae at the apex.

Etymology

The species is named after the Andasibe-Mantadia National Park located about 150 km east of Antananarivo, consisting principally of primary growth forest of the Alaotra-Mangoro Region in eastern Madagascar. The park’s two component parts are Mantadia National Park and Analamazoatra Reserve ( Newsome & Hassell 2014).

Material examined

Holotype MADAGASCAR • ♂; Andasibe, Analamazaotra Forest ; 18°46ʹ S, 48°24ʹ E; 950 m a.s.l.; 31 Oct.–4 Nov. 2007; A. Freidberg leg.; SMNHTAU.

Description

Male ( Fig. 8A )

MEASUREMENTS. Body length 5.2 mm; antenna length 1.1 mm; wing length 4.5 mm; wing width 1.2 mm.

HEAD ( Fig. 8B ). Frons black, white pollinose; strong short anterior vertical and long postvertical bristles; upper postocular setae black, short; lateral postocular setae white, uniserial; ventral postcranium covered with irregular white hairs; face densely silvery-white pollinose, narrow, 3.3 times as high as wide under antennae, 7 times as high as wide at clypeus, at clypeus as wide as postpedicel; clypeus small, slightly bulging; antenna ( Fig. 8C ) 1.3 times as long as height of head, with scape and pedicel dirty yellow, postpedicel brown; pedicel with short bristles, with one long dorsal bristle; postpedicel ovate, 1.2 times as long as high, with short hairs; arista-like stylus black, brown at base, dorsal, microscopically haired; length (mm) of scape, pedicel, postpedicel, stylus, 0.07/0.07/0.1/0.91; proboscis and palpus orange; palpus with brown hairs, 2 black setae.

THORAX. Mesonotum orange, with brown, shining greenish blue, medial stripe slightly expanding posteriorly; scutellum metallic blue; pleura yellow, weakly pollinose; 6 strong dorsocentral bristles gradually decreasing in size anteriorly with strongest posterior pair; acrostichals short, biserial; scutellum with 2 strong bristles and 2 hairs laterally.

LEGS. Mostly yellow; coxae yellow; segment 4 of fore tarsus black, segment 5 silvery white ( Fig. 8D ); last segments of mid and hind tarsi brown; fore and mid coxae with light and dark hairs and 2–3 black subapical setae; hind coxa with 1 black seta at middle; femora without strong or long setae; fore tibia without conspicuous setae; fore basitarsus thin and long, with 1 rather short basoventral seta; segment 4 cylindrical; segment 5 flattened dorsoventrally, slightly widened; segments 4–5 with white flattened setae laterally, slightly longer than tarsomere width; with black setulae dorsally; mid tibia with one short anterodorsal and two short posterodorsals (broken), without conspicuous ventral setae; mid and hind tarsi simple, cylindrical; hind tibia with several short dorsal and ventral setae; femur, tibia and tarsomere (from first to fifth) length ratio (mm): fore leg: 1.19/1.63/3.82/0.58/0.54/0.24/0.15, mid leg: 1.51/2.23/2.15/0.58/0.4/0.23/0.15, hind leg: 1.55/2.63/1.53/0.63/0.36/0.25/0.13.

WING ( Fig. 8E ). Widened distally, almost hyaline, veins brown; R 4+5 gently curved to M 1 in apical third; M straight in basal half, slightly convex anteriad in middle part; M 1 with strong elbow, forming right angle with M 1+2; ratio of parts of costa between R 2+3 and R 4+5 to those between R 4+5 and M 1, 5/1; crossvein dm-m straight; ratio of crossvein dm-m to apical part of M 1+2 (fork-handle) to apical part of M 4, 0.37.1.03/0.33; anal vein and lobe reduced; anal angle absent; lower calypter yellow, with fine brown-black cilia; halter yellow with brown knob.

1+2

ABDOMEN. Thin and long, mostly orange-yellow, brown along distal margin of tergites 2–5, with short black hairs and marginal setae; pregenital segments combined 2.4 times as long as mesonotum; segment 7 short, symmetrical (dorsal view); tergite 7 flat, with proximal angular emargination (dorsal view); sternite 7 distinct, embracing long and thin tergite 8; sternite 8 symmetrical, rounded, with short setae; hypopygium ( Fig. 8F ) black, with brown appendages; epandrial foramen located almost symmetrically on basoventral side of epandrium; hypandrium short, with reduced arms; cercus ( Fig. 8G ) brown, trifurcated from base; dorsal lobe flat, elongate-triangular, half as long as epandrium, with pointed apex, sparsely covered with dark setae on outer side, densely covered with microscopic hairs on inner side; middle arm of cercus ( Fig. 8H ) as long as epandrium, stick-shaped, flattened laterally, with 2 very long lateral setae at distal third; ventral arm of cercus strongly curved, finger-like, reaching to middle of middle arm, with small comma-like sclerotized process at base and 4 long apical setae; surstylus projected, bilobed, with irregularly curved broad flat arms; dorsal arm of surstylus pointed at apex, with one very long apical seta, with few short setae; ventral arm of surstylus with deep distal emargination, with 4 small modified setae on distoventral apex; epandrial lobe strongly projected distally, sclerotized, broad at base, narrow distally, with 2 fine epandrial setae at base ( Fig. 8F ).

Female

Unknown.

Figures

Fig. 8. Amblypsilopus andasibensis sp. nov. Holotype, ♂ (SMNHTAU). A. Habitus. B. Head. C. Antenna. D. Fore tarsomeres 2–5, lateral view, with inset showing segments 4–5, dorsal view. E. Wing. F. Hypopygium, left lateral view, reflected light. G. Cercal lobes, lateral view, transmitted light. H. Cercal lobes, dorsal view, transmitted light.