Chelagyrtodes glacicola Seago & Leschen

Seago, Ainsley E. & Leschen, Richard A. B., 2011, Revision and phylogeny of Chelagyrtodes Szymczakowski (Coleoptera: Leiodidae: Camiarinae: Agyrtodini), Zootaxa 3090, pp. 1-20 : 10-11

Chelagyrtodes glacicola Seago & Leschen , new species

( Figs. 4 c, 5c, 8)

Type material. Holotype: Male, labeled: NEW ZEALAND, WD, 2.7 km S Franz Josef Glacier, Tk Head of Alex Knob, 150 m, 15 Jan 1998, R Leschen & C Carlton, coastal forest leaf litter ( NZAC). Paratypes: NEW ZEA- LAND: WD: 1, Monroe Beach Tk, 8 Nov 2007, R Leschen & C Carlton, rotten wood / leaf litter berlesate ( NZAC); 1, 2.7 km S Franz Josef Glacier, Tk Head of Alex Knob, 150 m, 15 Jan 1998, R Leschen & C Carlton, coastal forest leaf litter ( NZAC); 1, 11.2 km NE Franz Josef Glacier, nr L Mapourika, Ottos Cnr picnic area, 150 m, 15 Jan 1998, R Leschen & C Carlton, coastal forest leaf litter ( NZAC); 1, 9 mi S Franz Josef town, 7 Feb 1965, N A Walker ( NZAC); 3, Fox Glacier, Westland, 12 Nov 1968, Townsend, moss ( NZAC); 3, Westland NP, 8.8 km NE Fox Glacier, 315 m, 18 Jan 1985, A Newton & M Thayer, berl litter, low hardwood forest ( FMNH); 2, Mananui Bush, 3 km NNE Ruatapu, 10 m, 9 Jan 1985, 19 Jan 1985, A Newton & M Thayer, coastal hdwd. Dicksonia dune forest, window trap ( FMNH); Fox Glacier, Westland, 12 Nov 68, J. Townsend, moss, 68 / 172; FD: Tempest Spur, W. Olivine Ra. FD 1220-1463, 25 Jan 75, G. W. Ramsay, Litter 75 / 33 ( NZAC).

Diagnosis. Male protibia simple. Body form elongately oval, hind margin of pronotum same width as elytral base. Epistomal suture absent. Elytra with sides converging in apical third, apex of closed elytra bluntly rounded; transverse strigae present. Pronotum with sparse, even vestiture of fine setae. Metaventrite moderately compressed, narrowest at midline. Abdominal ventrite 6 medially emarginate, not divided. Apex of median lobe blunt, very weakly produced; endophallus with two large paired internal sclerites not fused into a single median process. Parameres long, slender, weakly apically curved.

Description. Length 1.5 mm (n = 4). Body about 1.6 X as long as wide (BL / EW = 1.63 mm) greatest depth 0.6 X as great as elytral width (GD / EW = 0.58). Body form elongate, oval, with sides of elytra converging in apical ¼. Color of body unicolorous red to red-brown; mouthparts, tarsi, and clypeus light brown to pale, with femora same color as tibiae; vestiture of pale golden setae. Dorsum with sparse, setiferous punctures.

Antennomere ratios: 13:13:10:7:10:7:11:5:12:12:17; segment 6 quadrate, segment 8 transverse. Eyes small, protuberant, with weak posterior emargination. Eyes not coarsely facetted (edges of ommatidia smooth), greatest length 5 facets, greatest height 7 facets. Epistomal suture absent, clypeus subquadrate. Mandibles with single apical tooth, pronounced prostheca and mola present. Maxillae with slender galea, slightly broad lacinia, with loose, nonarticulated apical brush of minute, curved spines; maxillary palp with penultimate segment wider than other segments, nearly as long as attenuate terminal segment. Labial palpi weakly elongate, three-segmented, each segment slightly wider than long. Pronotum about 0.6 X as long as wide (PL / PW = 0.57). Rim of procoxal fossae strongly developed, parallel to and nearly confluent with anterior margin of prosternum. Elytra about 1.1 X as long as wide (EL / EW = 1.13) and about twice as long as pronotum (EL / PL = 1.96), widest in basal third; sides parallel and converging apically from posterior ½ to ¼; transverse strigae present; setae biseriate, decumbent to suberect, with setae not extending to insertion of next seta in series; width of epipleuron at level of metacoxa subequal to width of metanepisternum. Metaventrite moderately compressed, narrowest at midline. Metanepisternal clamp weakly developed. Mesothoracic, metathoracic and abdominal ventrites glabrous, with abundant small, deep punctures. Abdominal ventrites 1–4 each with two or more transverse rows of minute setae; abdominal ventrite 5 scarcely longer than 4, twice as long as 6. Ventrite 6 complete, at most weakly emarginate medially.

Tibia with 5 large blunt spines at apex, protarsi with segments 1–3 not expanded; male protibia simple, tenent setae present on protarsomeres 1–3, absent from mesotarsomeres. Tibiae and femorae unicolorous, reddish-brown. Male genital segment short, subquadrate to transverse; tergite 9 bluntly pointed at apices with setae confined apically, anterior apophysis slender and elongate, weakly expanded apically; sternite 9 longer than wide; tergite 10 (= proctiger) lacking apical notch. Aedeagus relatively short, broad at base, median lobe triangular, about 2 X longer than wide; apex of median lobe simple, median lobe triangular in dorsal view. Phallobase (basal piece) forming a narrow hyaline band surrounding the anterior portion of the median lobe; parameres extending scarcely beyond apex of median lobe, slender, slightly flattened, weakly bent inward distally, with cluster of two long and 2–3 short setae at apices. Endophallus membranous, with abundant minute, weakly-sclerotized setae and large, paired, irregular internal sclerites fused at base into a single prominent process. Spermatheca present, unsclerotized; gonocoxite with 4 long setae, stylus approximately ¼ as long and ½ as wide as gonocoxite, bearing one minute subapical seta and one elongate, robust seta equal in length to both styli segments combined.

Remarks. This species is very similar to C. haasti , and only differs in the form of the male genitalia which have highly hooked apical parameres. Specimens were collected mainly from moss, but also from leaf litter from coastal and lowland podocarp forests.

Distribution. New Zealand: South Island, Westland (WD) region.

Etymology. The specific epithet refers to the Westland glacial region where this species has been collected.

Figures

FIGURE 4. Habitus photographs of adult Chelagyrtodes. a) C. crowsoni; b) C. davidi; c) C. glacicola; d) C. haasti; e) C. newtoni; f) C. rotundus.