Onthophagus fuscus, Boucomont, 1932

Joaqui, Tatiana, Moctezuma, Victor, Sánchez-Huerta, José Luis & Escobar, Federico, 2019, The Onthophagus fuscus (Coleoptera: Scarabaeidae) species complex: an update and the description of a new species, Zootaxa 4555 (2), pp. 151-186 : 177

publication ID	https://doi.org/ 10.11646/zootaxa.4555.2.1
publication LSID	lsid:zoobank.org:pub:E8F6B60F-3E7B-4BD1-938C-F46A840AA08D
DOI	https://doi.org/10.5281/zenodo.5940898
persistent identifier	https://treatment.plazi.org/id/BA2787A6-807A-FFEB-7AEA-FB43FA171AF2
treatment provided by	Plazi
scientific name	Onthophagus fuscus
status

Treatment

Key for the identification of adult major male specimens of the Onthophagus fuscus View in CoL species complex

(modified from Zunino & Halffter 1988).

1 Clypeus not transverse, obtusely pentagonal and not concave or weakly concave ( Figs. 1, 3 View FIGURE 3 , 5 View FIGURE 5 , 7 View FIGURE 7 , 9 View FIGURE 9 , 13 View FIGURE 13 )................. 2

- Clypeus transverse, obtusely trapezoidal and strongly concave ( Fig. 11 View FIGURE 11 ).......................................... 7

2 Frontal carina complete................................................................................. 3

- Frontal carina limited to the medial region.................................................................. 5

3 Dorsal setae (pronotum and elytra) relatively long (length: 0.1 ± 0.03 mm). Pronotal disc glabrous medially ( Figs. 3–4 View FIGURE 3 View FIGURE 4 , 18 – 20 View FIGURE 18 View FIGURE 19 View FIGURE 20 ). Michoacán, Mexico...................................................... O. canescens Zunino & Halffter, 1988 View in CoL

- Pronotal setae relatively short (length: 0.07 ± 0.02 mm). Pubescence present on the entire pronotal disc................. 4

4 Protonal surface coppery without olive-green sheen and elliptical impressions. Clypeal carina lacking ( Figs. 9–10 View FIGURE 9 View FIGURE 10 , 18 – 20 View FIGURE 18 View FIGURE 19 View FIGURE 20 ). Hidalgo, Puebla, and Veracruz, Mexico...................................... O. parafuscus Zunino & Halffter, 2005 View in CoL

- Protonal surface coppery with an olive-green sheen, with two more-or-less elliptical impressions on each of midline. Clypeal carina weakly impressed ( Figs. 13 – 16 View FIGURE 13 View FIGURE 14 View FIGURE 15 View FIGURE 16 ). Veracruz, Mexico.................................................................................................... O. orizabensis Moctezuma, Joaqui View in CoL , & Sánchez-Huerta, new species

5 Frontal carina produced into an acute or flattened triangular tubercle between the genal sutures. Elytral striae fine ( Figs. 7–8 View FIGURE 7 View FIGURE 8 , 18 – 20 View FIGURE 18 View FIGURE 19 View FIGURE 20 ). Durango, Jalisco, and Sinaloa, Mexico.............................. O. pseudofuscus Zunino & Halffter, 1988 View in CoL

- Frontal carina not produced into an acute tubercle. Elytral striae strongly impressed................................. 6

6 Clypeal carina present, but weakly developed. Dorsal surface dark brown with a weak greenish-cupreous sheen ( Figs. 1, 2 View FIGURE 2 , 18 – 20 View FIGURE 18 View FIGURE 19 View FIGURE 20 ). Aguascalientes, Durango, Jalisco, and Zacatecas, Mexico............................. O. fuscus Boucomont, 1932 View in CoL

- Clypeal carina absent. Dorsal surface coppery with a strong metallic sheen ( Fig. 5–6 View FIGURE 5 View FIGURE 6 , 18 – 20 View FIGURE 18 View FIGURE 19 View FIGURE 20 ). Morelos, Mexico................................................................................. O. mycetorum Zunino & Halffter, 1988 View in CoL

7 Pubescence uniform in size and density on the pronotal disc ( Fig. 11–12 View FIGURE 11 View FIGURE 12 ). Puebla, Mexico......................................................................................... O. clavijeroi Moctezuma, Rossini, & Zunino, 2016 View in CoL

- Pubescence reduced in size and density on the postero-central region of the pronotal disc. See dorsal habitus at http:// unibio.unam.mx/irekani/bitstream/123456789/12809/1/9035.jpg. Morelos, Mexico.................................................................................................... O. navarretorum Delgado & Capistan, 1996 View in CoL