Goeldia utcuyacu, Almeida-Silva & Brescovit, 2024

Almeida-Silva, Lina M. & Brescovit, Antonio D., 2024, Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae), Zootaxa 5428 (2), pp. 151-193 : 185-187

Goeldia utcuyacu sp. nov.

( Figs 21 ; 22 ; 25 )

Type material. Holotype ♂ from Utcuyacu (11°11’11.9”S 75°27’51.4”W), Junin, Peru, elev. 1600–2200 m, III.1984, F. Woytkowski coll. ( AMNH); paratype female with same data as holotype ( AMNH).

Additional Material Examined. PERU. Cuzco. Ocongate (13°37’45”S 71°23’10”W), Hacienda Capana , elev. 3670 m, 2♀, 6–11.IV.1947, J. C. Pallister coll. ( AMNH) ; Urubamba, Machu-Picchu , (13°09’47”S 72°32’44”W), elev. 2600–2800 m, 1♀, 1–5.VIII.1964, B. Malkin coll. ( AMNH) . Junin: San Ramon (11°07’29”S 75°21’25”W), Estância Naranjal , 2♀, 20– 7.VII.1965 ( AMNH) ; Huacapistana (11°13’59”S 75°28’59”W), elev. 1800 m, 1♂, 2♀ 1juv., 27–30.VII.1965 ( AMNH) all collected by P. & B. Wygodzinsky; San José de Utcuyacu (11°11’10”S 75°27’47”W), 3♀, 1♂, 4 juv., III.1948, F. Woytkowski coll. ( AMNH) ; Colonia Perene, Rio Perene , 18 mi. NE La Merced, 2♂ 2♀, 3.I.1955, E. Schlinger & E. Ross coll. ( CAS 9021317 , SEM, male palp and female leg IV) .

Etymology. The specific epithet is a noun in apposition from the type locality, San José de Utcuyacu, Peru.

Diagnosis. Goeldia utcuyacu sp. nov. differs from other species of the genus by the thick patellar apophysis with sinuous tip ( Figs 21B–D ; 22E ); and by the tegular process subrectangular and with enlarged base in the male palp ( Figs 21A ; 22A ). Females can be recognized by the epigynum with median field wide, with posterior edge almost straight ( Fig. 21E ), and the vertical spermathecae slender and longer than copulatory ducts ( Fig. 21F ).

Description. Male (Holotype, AMNH). Carapace brown with cephalic area darker; femur I brown; other leg segments yellowish-brown; chelicerae dark brown, labium and endites brown. Total length 4.8. Carapace: length 2.4; width 1.8; height 1.34. Clypeus: height 0.16. Opisthosoma: length 2.43; width 1.4. Eye diameters and interdistances: AME 0.06; ALE 0.11; PME 0.10; PLE 0.12; AME–AME 0.11; AME–ALE 0.11; ALE–PLE 0.5; PME–PLE 0.20; PME–PME 0.13; AME–PME 0.12. Chelicerae paturon: length 1.25. Leg measurements: I: femur 2.2/ patella 1.3/ tibia 2.3/ metatarsus 2.2/ tarsus 1.0/total 9.0; II: 2.0/ 0.8/ 1.9/ 1.85/ 0.95/ 7.5; III: 1.85/ 0.7/ 1.5/ 1.6/ 0.7/ 6.35; IV: 2.35/ 0.7/ 2.2/ 2.0/ 0.85/ 8.1. Leg formula: 1423. Leg spination: femur I–II p1ap; tibia II v2-0-2 or v0-0-2, p1ap, r1ap; III v2sa or v1-0-2, r 0-0-1, p0-1-1; IV: v1-0-2 or v0-1-2, p1ap, r1ap; metatarsus I v1p-2- 1m, p1ap, r1ap; II v2-1r-2(1p-1r)-2(1p-1r)-2ap, p0-2-2, r2ap. Patellar apophysis dorso-posteriorly projected ( Figs 21B–D , 22E ). MLT base enlarged and apex partially sclerotized ( Figs 21C , 22D–E ). Cymbium with macrosetae ( Figs 21B , 22D ). Tegular process with uniform width and base membranous ( Figs 21A ; 22A ). Pars pendula thickening far from embolus tip ( Figs 21D ; 22A ).

Female (AMNH). Coloration: carapace, chelicerae, and femora dark brown; other leg segments, labium and endites caramel brown. Total length: 6.8. Carapace: length 2.56; width 1.92, height 1.28. Clypeus: height 0.21. Opisthosoma: length 4.4, width 3.2. Eye diameters and interdistances: AME 0.08; ALE 0.13; PME 0.12; PLE 0.13; AME–AME 0.08; AME–ALE 0.13;ALE–PLE 0.04; PME–PLE 0.21; PME–PME 0.14; AME–PME 0.09. Chelicerae paturon: length 1.29. Leg measurements: I: femur 1.85/ patella 0.85/ tibia 1.8/ metatarsus 1.55/ tarsus 0.8/ total 6.85; II: 1.8/ 0.9/ 1.5/ 1.4/ 0.7/ 6.3; III: 1.75/ 0.75/ 1.25/ 1.2/ 0.55/ 5.5; IV: 1.8/ 0.8/ 1.65/ 1.55/ 0.55/ 6.35. Leg formula 1423. Leg spination: femur I–II p1ap; tibia III p1ap; metatarsus I–II v2-2- 1m, p 1ap, r1ap; III v1r-1p-2ap, p1-0-1ap, r0-0-1p; IV v1p-1p- 2m, p1ap, r1ap. Median field of epigynum wide but not closing copulatory openings ( Fig. 21E ). Copulatory duct wall visible by transparency on the median field and partially visible at the copulatory opening area ( Fig. 21E ). Copulatory duct base enlarged ( Fig. 21E–F ). Vertical spermathecae straight ( Fig. 21F ). Horizontal spermathecae straight shorter than vertical spermathecae ( Fig. 21F ). Fertilization ducts enlarged and separated by their length ( Fig. 21F )

Variation. Length (2 ♂): total 4.20–4.80; carapace 2.00–2.40; femur I 2.10–2.20; (10 ♀): total 5.20–7.10; carapace 2.14–2.60; femur I 1.78–1.88.

Distribution. Peru ( Fig. 25 ).

Figures

FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view.

FIGURE 22. A–E. Goeldia utcuyacu sp. nov., from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view.

FIGURE 25. Distribution of Goedia spp. in northern South America: Goeldia diva (pentagon—pink); G. obscura (square— yellow); G. guayaquilensis (circle—purple); G. mirim (star—red); G.utcuyacu (rectangle—blue). Type species are marked as black versions of each symbol when there are records to multiple localities.